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+ <head>
+ <title>
+ Lecture to Working Men, No. 2 (of 6), The Past Condition of Organic
+ Nature, by Thomas H. Huxley
+ </title>
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+
+Project Gutenberg's The Past Condition of Organic Nature, by Thomas H. Huxley
+
+This eBook is for the use of anyone anywhere at no cost and with
+almost no restrictions whatsoever. You may copy it, give it away or
+re-use it under the terms of the Project Gutenberg License included
+with this eBook or online at www.gutenberg.org
+
+
+Title: The Past Condition of Organic Nature
+ Lecture II. (of VI.), Lectures To Working Men, at the
+ Museum of Practical Geology, 1863, On Darwin's work: "Origin
+ of Species".
+
+Author: Thomas H. Huxley
+
+Release Date: January 4, 2009 [EBook #2922]
+Last Updated: January 22, 2013
+
+Language: English
+
+Character set encoding: ASCII
+
+*** START OF THIS PROJECT GUTENBERG EBOOK CONDITION OF ORGANIC NATURE ***
+
+
+
+
+Produced by Amy E. Zelmer, and David Widger
+
+
+
+
+
+
+</pre>
+ <p>
+ <br /><br />
+ </p>
+ <h1>
+ THE PAST CONDITION <br /> OF ORGANIC NATURE
+ </h1>
+ <p>
+ <br />
+ </p>
+ <h3>
+ Lecture II. (of VI.), "Lectures To Working Men", at the Museum of
+ Practical Geology, 1863, On Darwin's work: "Origin of Species".
+ </h3>
+ <p>
+ <br /><br />
+ </p>
+ <h2>
+ By Thomas H. Huxley
+ </h2>
+ <p>
+ <br /> <br />
+ </p>
+ <hr />
+ <p>
+ <br /> <br />
+ </p>
+ <h3>
+ List of Illustrations
+ </h3>
+ <table summary="" style="margin-right: auto; margin-left: auto">
+ <tr>
+ <td>
+ <p class="toc">
+ <a href="#linkimage-0001"> Fig.4. </a>
+ </p>
+ <p class="toc">
+ <a href="#linkimage-0002"> Fig. 5. </a>
+ </p>
+ </td>
+ </tr>
+ </table>
+ <p>
+ <br /> <br />
+ </p>
+ <hr />
+ <p>
+ <br /> <br />
+ </p>
+ <p>
+ IN the lecture which I delivered last Monday evening, I endeavoured to
+ sketch in a very brief manner, but as well as the time at my disposal
+ would permit, the present condition of organic nature, meaning by that
+ large title simply an indication of the great, broad, and general
+ principles which are to be discovered by those who look attentively at the
+ phenomena of organic nature as at present displayed. The general result of
+ our investigations might be summed up thus: we found that the multiplicity
+ of the forms of animal life, great as that may be, may be reduced to a
+ comparatively few primitive plans or types of construction; that a further
+ study of the development of those different forms revealed to us that they
+ were again reducible, until we at last brought the infinite diversity of
+ animal, and even vegetable life, down to the primordial form of a single
+ cell.
+ </p>
+ <p>
+ We found that our analysis of the organic world, whether animals or
+ plants, showed, in the long run, that they might both be reduced into, and
+ were, in fact, composed of, the same constituents. And we saw that the
+ plant obtained the materials constituting its substance by a peculiar
+ combination of matters belonging entirely to the inorganic world; that,
+ then, the animal was constantly appropriating the nitrogenous matters of
+ the plant to its own nourishment, and returning them back to the inorganic
+ world, in what we spoke of as its waste; and that finally, when the animal
+ ceased to exist, the constituents of its body were dissolved and
+ transmitted to that inorganic world whence they had been at first
+ abstracted. Thus we saw in both the blade of grass and the horse but the
+ same elements differently combined and arranged. We discovered a continual
+ circulation going on,&mdash;the plant drawing in the elements of inorganic
+ nature and combining them into food for the animal creation; the animal
+ borrowing from the plant the matter for its own support, giving off during
+ its life products which returned immediately to the inorganic world; and
+ that, eventually, the constituent materials of the whole structure of both
+ animals and plants were thus returned to their original source: there was
+ a constant passage from one state of existence to another, and a returning
+ back again.
+ </p>
+ <p>
+ Lastly, when we endeavoured to form some notion of the nature of the
+ forces exercised by living beings, we discovered that they&mdash;if not
+ capable of being subjected to the same minute analysis as the constituents
+ of those beings themselves&mdash;that they were correlative with&mdash;that
+ they were the equivalents of the forces of inorganic nature&mdash;that
+ they were, in the sense in which the term is now used, convertible with
+ them. That was our general result.
+ </p>
+ <p>
+ And now, leaving the Present, I must endeavour in the same manner to put
+ before you the facts that are to be discovered in the Past history of the
+ living world, in the past conditions of organic nature. We have, to-night,
+ to deal with the facts of that history&mdash;a history involving periods
+ of time before which our mere human records sink into utter insignificance&mdash;a
+ history the variety and physical magnitude of whose events cannot even be
+ foreshadowed by the history of human life and human phenomena&mdash;a
+ history of the most varied and complex character.
+ </p>
+ <p>
+ We must deal with the history, then, in the first place, as we should deal
+ with all other histories. The historical student knows that his first
+ business should be to inquire into the validity of his evidence, and the
+ nature of the record in which the evidence is contained, that he may be
+ able to form a proper estimate of the correctness of the conclusions which
+ have been drawn from that evidence. So, here, we must pass, in the first
+ place, to the consideration of a matter which may seem foreign to the
+ question under discussion. We must dwell upon the nature of the records,
+ and the credibility of the evidence they contain; we must look to the
+ completeness or incompleteness of those records themselves, before we turn
+ to that which they contain and reveal. The question of the credibility of
+ the history, happily for us, will not require much consideration, for, in
+ this history, unlike those of human origin, there can be no cavilling, no
+ differences as to the reality and truth of the facts of which it is made
+ up; the facts state themselves, and are laid out clearly before us.
+ </p>
+ <p>
+ But, although one of the greatest difficulties of the historical student
+ is cleared out of our path, there are other difficulties&mdash;difficulties
+ in rightly interpreting the facts as they are presented to us&mdash;which
+ may be compared with the greatest difficulties of any other kinds of
+ historical study.
+ </p>
+ <p>
+ What is this record of the past history of the globe, and what are the
+ questions which are involved in an inquiry into its completeness or
+ incompleteness? That record is composed of mud; and the question which we
+ have to investigate this evening resolves itself into a question of the
+ formation of mud. You may think, perhaps, that this is a vast step&mdash;of
+ almost from the sublime to the ridiculous&mdash;from the contemplation of
+ the history of the past ages of the world's existence to the consideration
+ of the history of the formation of mud! But, in nature, there is nothing
+ mean and unworthy of attention; there is nothing ridiculous or
+ contemptible in any of her works; and this inquiry, you will soon see, I
+ hope, takes us to the very root and foundations of our subject.
+ </p>
+ <p>
+ How, then, is mud formed? Always, with some trifling exception, which I
+ need not consider now&mdash;always, as the result of the action of water,
+ wearing down and disintegrating the surface of the earth and rocks with
+ which it comes in contact&mdash;pounding and grinding it down, and
+ carrying the particles away to places where they cease to be disturbed by
+ this mechanical action, and where they can subside and rest. For the
+ ocean, urged by winds, washes, as we know, a long extent of coast, and
+ every wave, loaded as it is with particles of sand and gravel as it breaks
+ upon the shore, does something towards the disintegrating process. And
+ thus, slowly but surely, the hardest rocks are gradually ground down to a
+ powdery substance; and the mud thus formed, coarser or finer, as the case
+ may be, is carried by the rush of the tides, or currents, till it reaches
+ the comparatively deeper parts of the ocean, in which it can sink to the
+ bottom, that is, to parts where there is a depth of about fourteen or
+ fifteen fathoms, a depth at which the water is, usually, nearly
+ motionless, and in which, of course, the finer particles of this detritus,
+ or mud as we call it, sinks to the bottom.
+ </p>
+ <p>
+ Or, again, if you take a river, rushing down from its mountain sources,
+ brawling over the stones and rocks that intersect its path, loosening,
+ removing, and carrying with it in its downward course the pebbles and
+ lighter matters from its banks, it crushes and pounds down the rocks and
+ earths in precisely the same way as the wearing action of the sea waves.
+ The matters forming the deposit are torn from the mountain-side and
+ whirled impetuously into the valley, more slowly over the plain, thence
+ into the estuary, and from the estuary they are swept into the sea. The
+ coarser and heavier fragments are obviously deposited first, that is, as
+ soon as the current begins to lose its force by becoming amalgamated with
+ the stiller depths of the ocean, but the finer and lighter particles are
+ carried further on, and eventually deposited in a deeper and stiller
+ portion of the ocean.
+ </p>
+ <p>
+ It clearly follows from this that mud gives us a chronology; for it is
+ evident that supposing this, which I now sketch, to be the sea bottom, and
+ supposing this to be a coast-line; from the washing action of the sea upon
+ the rock, wearing and grinding it down into a sediment of mud, the mud
+ will be carried down, and at length, deposited in the deeper parts of this
+ sea bottom, where it will form a layer; and then, while that first layer
+ is hardening, other mud which is coming from the same source will, of
+ course, be carried to the same place; and, as it is quite impossible for
+ it to get beneath the layer already there, it deposits itself above it,
+ and forms another layer, and in that way you gradually have layers of mud
+ constantly forming and hardening one above the other, and conveying a
+ record of time.
+ </p>
+ <p>
+ It is a necessary result of the operation of the law of gravitation that
+ the uppermost layer shall be the youngest and the lowest the oldest, and
+ that the different beds shall be older at any particular point or spot in
+ exactly the ratio of their depth from the surface. So that if they were
+ upheaved afterwards, and you had a series of these different layers of
+ mud, converted into sandstone, or limestone, as the case might be, you
+ might be sure that the bottom layer was deposited first, and that the
+ upper layers were formed afterwards. Here, you see, is the first step in
+ the history&mdash;these layers of mud give us an idea of time.
+ </p>
+ <p>
+ The whole surface of the earth,&mdash;I speak broadly, and leave out minor
+ qualifications,&mdash;is made up of such layers of mud, so hard, the
+ majority of them, that we call them rock whether limestone or sandstone,
+ or other varieties of rock. And, seeing that every part of the crust of
+ the earth is made up in this way, you might think that the determination
+ of the chronology, the fixing of the time which it has taken to form this
+ crust is a comparatively simple matter. Take a broad average, ascertain
+ how fast the mud is deposited upon the bottom of the sea, or in the
+ estuary of rivers; take it to be an inch, or two, or three inches a year,
+ or whatever you may roughly estimate it at; then take the total thickness
+ of the whole series of stratified rocks, which geologists estimate at
+ twelve or thirteen miles, or about seventy thousand feet, make a sum in
+ short division, divide the total thickness by that of the quantity
+ deposited in one year, and the result will, of course, give you the number
+ of years which the crust has taken to form.
+ </p>
+ <p>
+ Truly, that looks a very simple process! It would be so except for certain
+ difficulties, the very first of which is that of finding how rapidly
+ sediments are deposited; but the main difficulty&mdash;a difficulty which
+ renders any certain calculations of such a matter out of the question&mdash;is
+ this, the sea-bottom on which the deposit takes place is continually
+ shifting.
+ </p>
+ <p>
+ Instead of the surface of the earth being that stable, fixed thing that it
+ is popularly believed to be, being, in common parlance, the very emblem of
+ fixity itself, it is incessantly moving, and is, in fact, as unstable as
+ the surface of the sea, except that its undulations are infinitely slower
+ and enormously higher and deeper.
+ </p>
+ <p>
+ Now, what is the effect of this oscillation? Take the case to which I have
+ previously referred. The finer or coarser sediments that are carried down
+ by the current of the river, will only be carried out a certain distance,
+ and eventually, as we have already seen, on reaching the stiller part of
+ the ocean, will be deposited at the bottom.
+ </p>
+ <p>
+ Let C y (Fig. 4) be the sea-bottom, y D the shore, x y the sea-level, then
+ the coarser deposit will subside over the region B, the finer over A,
+ while beyond A there will be no deposit at all; and, consequently, no
+ record will be kept, simply because no deposit is going on. Now, suppose
+ that the whole land, C, D, which we have regarded as stationary, goes
+ down, as it does so, both A and B go further out from the shore, which
+ will be at y1; x1, y1, being the new sea-level. The consequence will be
+ that the layer of mud (A), being now, for the most part, further than the
+ force of the current is strong enough to convey even the finest 'debris',
+ will, of course, receive no more deposits, and having attained a certain
+ thickness will now grow no thicker.
+ </p>
+ <p>
+ We should be misled in taking the thickness of that layer, whenever it may
+ be exposed to our view, as a record of time in the manner in which we are
+ now regarding this subject, as it would give us only an imperfect and
+ partial record: it would seem to represent too short a period of time.
+ </p>
+ <p>
+ <a name="linkimage-0001" id="linkimage-0001">
+ <!-- IMG --></a>
+ </p>
+ <div class="fig" style="width:80%">
+ <img src="images/fig04.jpg" alt="Fig.4. " width="100%" /><br />
+ </div>
+ <p>
+ Suppose, on the other hand, that the land (C D) had gone on rising slowly
+ and gradually&mdash;say an inch or two inches in the course of a century,&mdash;what
+ would be the practical effect of that movement? Why, that the sediment A
+ and B which has been already deposited, would eventually be brought nearer
+ to the shore-level, and again subjected to the wear and tear of the sea;
+ and directly the sea begins to act upon it, it would of course soon cut up
+ and carry it away, to a greater or less extent, to be re-deposited further
+ out.
+ </p>
+ <p>
+ Well, as there is, in all probability, not one single spot on the whole
+ surface of the earth, which has not been up and down in this way a great
+ many times, it follows that the thickness of the deposits formed at any
+ particular spot cannot be taken (even supposing we had at first obtained
+ correct data as to the rate at which they took place) as affording
+ reliable information as to the period of time occupied in its deposit. So
+ that you see it is absolutely necessary from these facts, seeing that our
+ record entirely consists of accumulations of mud, superimposed one on the
+ other; seeing in the next place that any particular spots on which
+ accumulations have occurred, have been constantly moving up and down, and
+ sometimes out of the reach of a deposit, and at other times its own
+ deposit broken up and carried away, it follows that our record must be in
+ the highest degree imperfect, and we have hardly a trace left of thick
+ deposits, or any definite knowledge of the area that they occupied, in a
+ great many cases. And mark this! That supposing even that the whole
+ surface of the earth had been accessible to the geologist,&mdash;that man
+ had had access to every part of the earth, and had made sections of the
+ whole, and put them all together,&mdash;even then his record must of
+ necessity be imperfect.
+ </p>
+ <p>
+ But to how much has man really access? If you will look at this Map you
+ will see that it represents the proportion of the sea to the earth: this
+ coloured part indicates all the dry land, and this other portion is the
+ water. You will notice at once that the water covers three-fifths of the
+ whole surface of the globe, and has covered it in the same manner ever
+ since man has kept any record of his own observations, to say nothing of
+ the minute period during which he has cultivated geological inquiry. So
+ that three-fifths of the surface of the earth is shut out from us because
+ it is under the sea. Let us look at the other two-fifths, and see what are
+ the countries in which anything that may be termed searching geological
+ inquiry has been carried out: a good deal of France, Germany, and Great
+ Britain and Ireland, bits of Spain, of Italy, and of Russia, have been
+ examined, but of the whole great mass of Africa, except parts of the
+ southern extremity, we know next to nothing; little bits of India, but of
+ the greater part of the Asiatic continent nothing; bits of the Northern
+ American States and of Canada, but of the greater part of the continent of
+ North America, and in still larger proportion, of South America, nothing!
+ </p>
+ <p>
+ Under these circumstances, it follows that even with reference to that
+ kind of imperfect information which we can possess, it is only of about
+ the ten-thousandth part of the accessible parts of the earth that has been
+ examined properly. Therefore, it is with justice that the most thoughtful
+ of those who are concerned in these inquiries insist continually upon the
+ imperfection of the geological record; for, I repeat, it is absolutely
+ necessary, from the nature of things, that that record should be of the
+ most fragmentary and imperfect character. Unfortunately this circumstance
+ has been constantly forgotten. Men of science, like young colts in a fresh
+ pasture, are apt to be exhilarated on being turned into a new field of
+ inquiry, to go off at a hand-gallop, in total disregard of hedges and
+ ditches, losing sight of the real limitation of their inquiries, and to
+ forget the extreme imperfection of what is really known. Geologists have
+ imagined that they could tell us what was going on at all parts of the
+ earth's surface during a given epoch; they have talked of this deposit
+ being contemporaneous with that deposit, until, from our little local
+ histories of the changes at limited spots of the earth's surface, they
+ have constructed a universal history of the globe as full of wonders and
+ portents as any other story of antiquity.
+ </p>
+ <p>
+ But what does this attempt to construct a universal history of the globe
+ imply? It implies that we shall not only have a precise knowledge of the
+ events which have occurred at any particular point, but that we shall be
+ able to say what events, at any one spot, took place at the same time with
+ those at other spots.
+ </p>
+ <p>
+ Let us see how far that is in the nature of things practicable. Suppose
+ that here I make a section of the Lake of Killarney, and here the section
+ of another lake&mdash;that of Loch Lomond in Scotland for instance. The
+ rivers that flow into them are constantly carrying down deposits of mud,
+ and beds, or strata, are being as constantly formed, one above the other,
+ at the bottom of those lakes. Now, there is not a shadow of doubt that in
+ these two lakes the lower beds are all older than the upper&mdash;there is
+ no doubt about that; but what does 'this' tell us about the age of any
+ given bed in Loch Lomond, as compared with that of any given bed in the
+ Lake of Killarney? It is, indeed, obvious that if any two sets of deposits
+ are separated and discontinuous, there is absolutely no means whatever
+ given you by the nature of the deposit of saying whether one is much
+ younger or older than the other; but you may say, as many have said and
+ think, that the case is very much altered if the beds which we are
+ comparing are continuous. Suppose two beds of mud hardened into rock,&mdash;A
+ and B-are seen in section. (Fig. 5.)
+ </p>
+ <p>
+ <a name="linkimage-0002" id="linkimage-0002">
+ <!-- IMG --></a>
+ </p>
+ <div class="fig" style="width:80%">
+ <img src="images/fig05.jpg" alt="Fig. 5. " width="100%" /><br />
+ </div>
+ <p>
+ Well, you say, it is admitted that the lowermost bed is always the older.
+ Very well; B, therefore, is older than A. No doubt, 'as a whole', it is
+ so; or if any parts of the two beds which are in the same vertical line
+ are compared, it is so. But suppose you take what seems a very natural
+ step further, and say that the part 'a' of the bed A is younger than the
+ part 'b' of the bed B. Is this sound reasoning? If you find any record of
+ changes taking place at 'b', did they occur before any events which took
+ place while 'a' was being deposited? It looks all very plain sailing,
+ indeed, to say that they did; and yet there is no proof of anything of the
+ kind. As the former Director of this Institution, Sir H. De la Beche, long
+ ago showed, this reasoning may involve an entire fallacy. It is extremely
+ possible that 'a' may have been deposited ages before 'b'. It is very easy
+ to understand how that can be. To return to Fig. 4; when A and B were
+ deposited, they were 'substantially' contemporaneous; A being simply the
+ finer deposit, and B the coarser of the same detritus or waste of land.
+ Now suppose that that sea-bottom goes down (as shown in Fig. 4), so that
+ the first deposit is carried no farther than 'a', forming the bed Al, and
+ the coarse no farther than 'b', forming the bed B1, the result will be the
+ formation of two continuous beds, one of fine sediment (A A1) over-lapping
+ another of coarse sediment (B B1). Now suppose the whole sea-bottom is
+ raised up, and a section exposed about the point Al; no doubt, 'at this
+ spot', the upper bed is younger than the lower. But we should obviously
+ greatly err if we concluded that the mass of the upper bed at A was
+ younger than the lower bed at B; for we have just seen that they are
+ contemporaneous deposits. Still more should we be in error if we supposed
+ the upper bed at A to be younger than the continuation of the lower bed at
+ Bl; for A was deposited long before B1. In fine, if, instead of comparing
+ immediately adjacent parts of two beds, one of which lies upon another, we
+ compare distant parts, it is quite possible that the upper may be any
+ number of years older than the under, and the under any number of years
+ younger than the upper.
+ </p>
+ <p>
+ Now you must not suppose that I put this before you for the purpose of
+ raising a paradoxical difficulty; the fact is, that the great mass of
+ deposits have taken place in sea-bottoms which are gradually sinking, and
+ have been formed under the very conditions I am here supposing.
+ </p>
+ <p>
+ Do not run away with the notion that this subverts the principle I laid
+ down at first. The error lies in extending a principle which is perfectly
+ applicable to deposits in the same vertical line to deposits which are not
+ in that relation to one another.
+ </p>
+ <p>
+ It is in consequence of circumstances of this kind, and of others that I
+ might mention to you, that our conclusions on and interpretations of the
+ record are really and strictly only valid so long as we confine ourselves
+ to one vertical section. I do not mean to tell you that there are no
+ qualifying circumstances, so that, even in very considerable areas, we may
+ safely speak of conformably superimposed beds being older or younger than
+ others at many different points. But we can never be quite sure in coming
+ to that conclusion, and especially we cannot be sure if there is any break
+ in their continuity, or any very great distance between the points to be
+ compared.
+ </p>
+ <p>
+ Well now, so much for the record itself,&mdash;so much for its
+ imperfections,&mdash;so much for the conditions to be observed in
+ interpreting it, and its chronological indications, the moment we pass
+ beyond the limits of a vertical linear section.
+ </p>
+ <p>
+ Now let us pass from the record to that which it contains,&mdash;from the
+ book itself to the writing and the figures on its pages. This writing and
+ these figures consist of remains of animals and plants which, in the great
+ majority of cases, have lived and died in the very spot in which we now
+ find them, or at least in the immediate vicinity. You must all of you be
+ aware&mdash;and I referred to the fact in my last lecture&mdash;that there
+ are vast numbers of creatures living at the bottom of the sea. These
+ creatures, like all others, sooner or later die, and their shells and hard
+ parts lie at the bottom; and then the fine mud which is being constantly
+ brought down by rivers and the action of the wear and tear of the sea,
+ covers them over and protects them from any further change or alteration;
+ and, of course, as in process of time the mud becomes hardened and
+ solidified, the shells of these animals are preserved and firmly imbedded
+ in the limestone or sandstone which is being thus formed. You may see in
+ the galleries of the Museum up stairs specimens of limestones in which
+ such fossil remains of existing animals are imbedded. There are some
+ specimens in which turtles' eggs have been imbedded in calcareous sand,
+ and before the sun had hatched the young turtles, they became covered over
+ with calcareous mud, and thus have been preserved and fossilized.
+ </p>
+ <p>
+ Not only does this process of imbedding and fossilization occur with
+ marine and other aquatic animals and plants, but it affects those land
+ animals and plants which are drifted away to sea, or become buried in bogs
+ or morasses; and the animals which have been trodden down by their fellows
+ and crushed in the mud at the river's bank, as the herd have come to
+ drink. In any of these cases, the organisms may be crushed or be
+ mutilated, before or after putrefaction, in such a manner that perhaps
+ only a part will be left in the form in which it reaches us. It is,
+ indeed, a most remarkable fact, that it is quite an exceptional case to
+ find a skeleton of any one of all the thousands of wild land animals that
+ we know are constantly being killed, or dying in the course of nature:
+ they are preyed on and devoured by other animals or die in places where
+ their bodies are not afterwards protected by mud. There are other animals
+ existing in the sea, the shells of which form exceedingly large deposits.
+ You are probably aware that before the attempt was made to lay the
+ Atlantic telegraphic cable, the Government employed vessels in making a
+ series of very careful observations and soundings of the bottom of the
+ Atlantic; and although, as we must all regret, up to the present time that
+ project has not succeeded, we have the satisfaction of knowing that it
+ yielded some most remarkable results to science. The Atlantic Ocean had to
+ be sounded right across, to depths of several miles in some places, and
+ the nature of its bottom was carefully ascertained. Well, now, a space of
+ about 1,000 miles wide from east to west, and I do not exactly know how
+ many from north to south, but at any rate 600 or 700 miles, was carefully
+ examined, and it was found that over the whole of that immense area an
+ excessively fine chalky mud is being deposited; and this deposit is
+ entirely made up of animals whose hard parts are deposited in this part of
+ the ocean, and are doubtless gradually acquiring solidity and becoming
+ metamorphosed into a chalky limestone. Thus, you see, it is quite possible
+ in this way to preserve unmistakable records of animal and vegetable life.
+ Whenever the sea-bottom, by some of those undulations of the earth's crust
+ that I have referred to, becomes upheaved, and sections or borings are
+ made, or pits are dug, then we become able to examine the contents and
+ constituents of these ancient sea-bottoms, and find out what manner of
+ animals lived at that period.
+ </p>
+ <p>
+ Now it is a very important consideration in its bearing on the
+ completeness of the record, to inquire how far the remains contained in
+ these fossiliferous limestones are able to convey anything like an
+ accurate or complete account of the animals which were in existence at the
+ time of its formation. Upon that point we can form a very clear judgment,
+ and one in which there is no possible room for any mistake. There are of
+ course a great number of animals&mdash;such as jelly-fishes, and other
+ animals&mdash;without any hard parts, of which we cannot reasonably expect
+ to find any traces whatever: there is nothing of them to preserve. Within
+ a very short time, you will have noticed, after they are removed from the
+ water, they dry up to a mere nothing; certainly they are not of a nature
+ to leave any very visible traces of their existence on such bodies as
+ chalk or mud. Then again, look at land animals; it is, as I have said, a
+ very uncommon thing to find a land animal entire after death. Insects and
+ other carnivorous animals very speedily pull them to pieces, putrefaction
+ takes place, and so, out of the hundreds of thousands that are known to
+ die every year, it is the rarest thing in the world to see one imbedded in
+ such a way that its remains would be preserved for a lengthened period.
+ Not only is this the case, but even when animal remains have been safely
+ imbedded, certain natural agents may wholly destroy and remove them.
+ </p>
+ <p>
+ Almost all the hard parts of animals&mdash;the bones and so on&mdash;are
+ composed chiefly of phosphate of lime and carbonate of lime. Some years
+ ago, I had to make an inquiry into the nature of some very curious fossils
+ sent to me from the North of Scotland. Fossils are usually hard bony
+ structures that have become imbedded in the way I have described, and have
+ gradually acquired the nature and solidity of the body with which they are
+ associated; but in this case I had a series of 'holes' in some pieces of
+ rock, and nothing else. Those holes, however, had a certain definite shape
+ about them, and when I got a skilful workman to make castings of the
+ interior of these holes, I found that they were the impressions of the
+ joints of a backbone and of the armour of a great reptile, twelve or more
+ feet long. This great beast had died and got buried in the sand; the sand
+ had gradually hardened over the bones, but remained porous. Water had
+ trickled through it, and that water being probably charged with a
+ superfluity of carbonic acid, had dissolved all the phosphate and
+ carbonate of lime, and the bones themselves had thus decayed and entirely
+ disappeared; but as the sandstone happened to have consolidated by that
+ time, the precise shape of the bones was retained. If that sandstone had
+ remained soft a little longer, we should have known nothing whatsoever of
+ the existence of the reptile whose bones it had encased.
+ </p>
+ <p>
+ How certain it is that a vast number of animals which have existed at one
+ period on this earth have entirely perished, and left no trace whatever of
+ their forms, may be proved to you by other considerations. There are large
+ tracts of sandstone in various parts of the world, in which nobody has yet
+ found anything but footsteps. Not a bone of any description, but an
+ enormous number of traces of footsteps. There is no question about them.
+ There is a whole valley in Connecticut covered with these footsteps, and
+ not a single fragment of the animals which made them has yet been found.
+ Let me mention another case while upon that matter, which is even more
+ surprising than those to which I have yet referred. There is a limestone
+ formation near Oxford, at a place called Stonesfield, which has yielded
+ the remains of certain very interesting mammalian animals, and up to this
+ time, if I recollect rightly, there have been found seven specimens of its
+ lower jaws, and not a bit of anything else, neither limb-bones nor skull,
+ or any part whatever; not a fragment of the whole system! Of course, it
+ would be preposterous to imagine that the beasts had nothing else but a
+ lower jaw! The probability is, as Dr. Buckland showed, as the result of
+ his observations on dead dogs in the river Thames, that the lower jaw, not
+ being secured by very firm ligaments to the bones of the head, and being a
+ weighty affair, would easily be knocked off, or might drop away from the
+ body as it floated in water in a state of decomposition. The jaw would
+ thus be deposited immediately, while the rest of the body would float and
+ drift away altogether, ultimately reaching the sea, and perhaps becoming
+ destroyed. The jaw becomes covered up and preserved in the river silt, and
+ thus it comes that we have such a curious circumstance as that of the
+ lower jaws in the Stonesfield slates. So that, you see, faulty as these
+ layers of stone in the earth's crust are, defective as they necessarily
+ are as a record, the account of contemporaneous vital phenomena presented
+ by them is, by the necessity of the case, infinitely more defective and
+ fragmentary.
+ </p>
+ <p>
+ It was necessary that I should put all this very strongly before you,
+ because, otherwise, you might have been led to think differently of the
+ completeness of our knowledge by the next facts I shall state to you.
+ </p>
+ <p>
+ The researches of the last three-quarters of a century have, in truth,
+ revealed a wonderful richness of organic life in those rocks. Certainly
+ not fewer than thirty or forty thousand different species of fossils have
+ been discovered. You have no more ground for doubting that these creatures
+ really lived and died at or near the places in which we find them than you
+ have for like scepticism about a shell on the sea-shore. The evidence is
+ as good in the one case as in the other.
+ </p>
+ <p>
+ Our next business is to look at the general character of these fossil
+ remains, and it is a subject which it will be requisite to consider
+ carefully; and the first point for us is to examine how much the extinct
+ 'Flora' and 'Fauna' as a 'whole'&mdash;disregarding altogether the
+ 'succession' of their constituents, of which I shall speak afterwards&mdash;differ
+ from the 'Flora' and 'Fauna' of the present day;&mdash;how far they differ
+ in what we 'do' know about them, leaving altogether out of consideration
+ speculations based upon what we 'do not' know.
+ </p>
+ <p>
+ I strongly imagine that if it were not for the peculiar appearance that
+ fossilised animals have, any of you might readily walk through a museum
+ which contains fossil remains mixed up with those of the present forms of
+ life, and I doubt very much whether your uninstructed eyes would lead you
+ to see any vast or wonderful difference between the two. If you looked
+ closely, you would notice, in the first place, a great many things very
+ like animals with which you are acquainted now: you would see differences
+ of shape and proportion, but on the whole a close similarity.
+ </p>
+ <p>
+ I explained what I meant by ORDERS the other day, when I described the
+ animal kingdom as being divided in sub-kingdoms, classes and orders. If
+ you divide the animal kingdom into orders, you will find that there are
+ about one hundred and twenty. The number may vary on one side or the
+ other, but this is a fair estimate. That is the sum total of the orders of
+ all the animals which we know now, and which have been known in past
+ times, and left remains behind.
+ </p>
+ <p>
+ Now, how many of those are absolutely extinct? That is to say, how many of
+ these orders of animals have lived at a former period of the world's
+ history, but have at present no representatives? That is the sense in
+ which I meant to use the word "extinct." I mean that those animals did
+ live on this earth at one time, but have left no one of their kind with us
+ at the present moment. So that estimating the number of extinct animals is
+ a sort of way of comparing the past creation as a whole with the present
+ as a whole. Among the mammalia and birds there are none extinct; but when
+ we come to the reptiles there is a most wonderful thing: out of the eight
+ orders, or thereabouts, which you can make among reptiles, one-half are
+ extinct. These diagrams of the plesiosaurus, the ichthyosaurus, the
+ pterodactyle, give you a notion of some of these extinct reptiles. And
+ here is a cast of the pterodactyle and bones of the ichthyosaurus and the
+ plesiosaurus, just as fresh as if it had been recently dug up in a
+ churchyard. Thus, in the reptile class, there are no less than half of the
+ orders which are absolutely extinct. If we turn to the 'Amphibia', there
+ was one extinct order, the Labyrinthodonts, typified by the large
+ salamander-like beast shown in this diagram.
+ </p>
+ <p>
+ No order of fishes is known to be extinct. Every fish that we find in the
+ strata&mdash;to which I have been referring&mdash;can be identified and
+ placed in one of the orders which exist at the present day. There is not
+ known to be a single ordinal form of insect extinct. There are only two
+ orders extinct among the 'Crustacea'. There is not known to be an extinct
+ order of these creatures, the parasitic and other worms; but there are
+ two, not to say three, absolutely extinct orders of this class, the
+ 'Echinodermata'; out of all the orders of the 'Coelenterata' and
+ 'Protozoa' only one, the Rugose Corals.
+ </p>
+ <p>
+ So that, you see, out of somewhere about 120 orders of animals, taking
+ them altogether, you will not, at the outside estimate, find above ten or
+ a dozen extinct. Summing up all the orders of animals which have left
+ remains behind them, you will not find above ten or a dozen which cannot
+ be arranged with those of the present day; that is to say, that the
+ difference does not amount to much more than ten per cent.: and the
+ proportion of extinct orders of plants is still smaller. I think that that
+ is a very astounding, a most astonishing fact, seeing the enormous epochs
+ of time which have elapsed during the constitution of the surface of the
+ earth as it at present exists; it is, indeed, a most astounding thing that
+ the proportion of extinct ordinal types should be so exceedingly small.
+ </p>
+ <p>
+ But now, there is another point of view in which we must look at this past
+ creation. Suppose that we were to sink a vertical pit through the floor
+ beneath us, and that I could succeed in making a section right through in
+ the direction of New Zealand, I should find in each of the different beds
+ through which I passed the remains of animals which I should find in that
+ stratum and not in the others. First, I should come upon beds of gravel or
+ drift containing the bones of large animals, such as the elephant,
+ rhinoceros, and cave tiger. Rather curious things to fall across in
+ Piccadilly! If I should dig lower still, I should come upon a bed of what
+ we call the London clay, and in this, as you will see in our galleries
+ upstairs, are found remains of strange cattle, remains of turtles, palms,
+ and large tropical fruits; with shell-fish such as you see the like of now
+ only in tropical regions. If I went below that, I should come upon the
+ chalk, and there I should find something altogether different, the remains
+ of ichthyosauri and pterodactyles, and ammonites, and so forth.
+ </p>
+ <p>
+ I do not know what Mr. Godwin Austin would say comes next, but probably
+ rocks containing more ammonites, and more ichthyosauri and plesiosauri,
+ with a vast number of other things; and under that I should meet with yet
+ older rocks, containing numbers of strange shells and fishes; and in thus
+ passing from the surface to the lowest depths of the earth's crust, the
+ forms of animal life and vegetable life which I should meet with in the
+ successive beds would, looking at them broadly, be the more different the
+ further that I went down. Or, in other words, inasmuch as we started with
+ the clear principle, that in a series of naturally-disposed mud beds the
+ lowest are the oldest, we should come to this result, that the further we
+ go back in time the more difference exists between the animal and
+ vegetable life of an epoch and that which now exists. That was the
+ conclusion to which I wished to bring you at the end of this Lecture.
+ </p>
+ <p>
+ <br /><br /><br /><br />
+ </p>
+<pre xml:space="preserve">
+
+
+
+
+
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+</pre>
+ </body>
+</html>
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