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You may copy it, give it away or +re-use it under the terms of the Project Gutenberg License included +with this eBook or online at www.gutenberg.org + + +Title: The Elements of Botany + For Beginners and For Schools + +Author: Asa Gray + +Release Date: September 18, 2010 [EBook #33757] + +Language: English + +Character set encoding: ISO-8859-1 + +*** START OF THIS PROJECT GUTENBERG EBOOK THE ELEMENTS OF BOTANY *** + + + + +Produced by Curtis Weyant, Stephen H. Sentoff and the +Online Distributed Proofreading Team at https://www.pgdp.net + + + + + + +</pre> + + + + + + +<div class="likeheading2">GRAY'S LESSONS IN BOTANY<br /> + +REVISED EDITION</div> + + + +<hr class="chap" /> +<h1>THE<br /> +ELEMENTS OF BOTANY</h1> + +<div class="likeheading2">FOR BEGINNERS AND FOR SCHOOLS</div> + + +<div class="likeheading3"><span class="smcap">By ASA GRAY</span></div> + + + +<hr class="chap" /> +<p class="center">IVISON, BLAKEMAN, AND COMPANY<br /> +NEW YORK AND CHICAGO</p> + + + +<hr class="chap" /> +<p class="center"><i>Copyright</i>,<br /> +<span class="smcap">By Asa Gray</span>.<br /> +1887. +</p> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_iii" id="Page_iii">[Pg iii]</a></span></p> +<h2><a name="PREFACE" id="PREFACE"></a>PREFACE.</h2> + + +<p>This volume takes the place of the author's <span class="smcap">Lessons in Botany and +Vegetable Physiology</span>, published over a quarter of a century ago. It is +constructed on the same lines, and is a kind of new and much revised +edition of that successful work. While in some respects more extended, +it is also more concise and terse than its predecessor. This should the +better fit it for its purpose now that competent teachers are common. +They may in many cases develop paragraphs into lectures, and fully +illustrate points which are barely, but it is hoped clearly, stated. +Indeed, even for those without a teacher, it may be that a condensed is +better than a diffuse exposition.</p> + +<p>The book is adapted to the higher schools, "How Plants Grow and Behave" +being the "Botany for Young People and Common Schools." It is intended +to ground beginners in Structural Botany and the principles of vegetable +life, mainly as concerns Flowering or Phanerogamous plants, with which +botanical instruction should always begin; also to be a companion and +interpreter to the Manuals and Floras by which the student threads his +flowery way to a clear knowledge of the surrounding vegetable creation. +Such a book, like a grammar, must needs abound in technical words, which +thus arrayed may seem formidable; nevertheless, if rightly apprehended, +this treatise should teach that the study of botany is not the learning +of names and terms, but the acquisition of knowledge and ideas. No +effort should be made to commit technical terms to memory. Any term used +in describing a plant or explaining its structure can be looked up when +it is wanted, and that should suffice. On the other hand, plans of +<span class="pagenum"><a name="Page_iv" id="Page_iv">[Pg iv]</a></span>structure, types, adaptations, and modifications, once understood, are +not readily forgotten; and they give meaning and interest to the +technical terms used in explaining them.</p> + +<p>In these "Elements" naturally no mention has been made of certain terms +and names which recent cryptogamically-minded botanists, with lack of +proportion and just perspective, are endeavoring to introduce into +phanerogamous botany, and which are not needed nor appropriate, even in +more advanced works, for the adequate recognition of the ascertained +analogies and homologies.</p> + +<p>As this volume will be the grammar and dictionary to more than one or +two Manuals, Floras, etc., the particular directions for procedure which +were given in the "First Lessons" are now relegated to those works +themselves, which in their new editions will provide the requisite +explanations. On the other hand, in view of such extended use, the +Glossary at the end of this book has been considerably enlarged. It will +be found to include not merely the common terms of botanical description +but also many which are unusual or obsolete; yet any of them may now and +then be encountered. Moreover, no small number of the Latin and Greek +words which form the whole or part of the commoner specific names are +added to this Glossary, some in an Anglicized, others in their Latin +form. This may be helpful to students with small Latin and less Greek, +in catching the meaning of a botanical name or term.</p> + +<p>The illustrations in this volume are largely increased in number. They +are mostly from the hand of Isaac Sprague.</p> + +<p>It happens that the title chosen for this book is that of the author's +earliest publication, in the year 1836, of which copies are rarely seen; +so that no inconvenience is likely to arise from the present use of the +name.</p> + +<p style="margin-left: 50%;"> +ASA GRAY.<br /> +<br /> +<span class="smcap">Cambridge, Massachusetts</span>,<br /> +<i>March, 1887</i>. +</p> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_v" id="Page_v">[Pg v]</a></span></p> +<h2><a name="CONTENTS" id="CONTENTS"></a>CONTENTS.</h2> + +<div style="margin-right:15em;"> +<ul class="lsoff"> +<li> <span class="tocnum">Page</span></li> +<li><a href="#Section_I">SECTION I. INTRODUCTORY</a> <span class="tocnum"><a href="#Page_9">9</a></span></li> + + +<li><a href="#Section_II">SECTION II. FLAX AS A PATTERN PLANT</a> <span class="tocnum"><a href="#Page_11">11</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_8">Growth from the Seed</a>, <a href="#UNIT_12">Organs of Vegetation</a> <span class="tocnum"><a href="#Page_11">11</a></span></li> +<li><a href="#UNIT_15">Blossoming</a>, <a href="#UNIT_16">Flower</a>, &c. <span class="tocnum"><a href="#Page_14">14</a></span></li> +</ul></li> + + +<li><a href="#Section_III">SECTION III. MORPHOLOGY OF SEEDLINGS</a> <span class="tocnum"><a href="#Page_15">15</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_19">Germinating Maples</a> <span class="tocnum"><a href="#Page_15">15</a></span></li> +<li><a href="#UNIT_24">Cotyledons thickened</a>, <a href="#UNIT_26">hypogæous in germination</a> <span class="tocnum"><a href="#Page_18">18</a></span></li> +<li><a href="#UNIT_31">Store of Food external to the Embryo</a> <span class="tocnum"><a href="#Page_20">20</a></span></li> +<li><a href="#UNIT_36">Cotyledons as to number</a> <span class="tocnum"><a href="#Page_22">22</a></span></li> +<li><a href="#UNIT_37">Dicotyledonous</a> and <a href="#UNIT_38">Polycotyledonous</a> <span class="tocnum"><a href="#Page_23">23</a></span></li> +<li><a href="#UNIT_40">Monocotyledonous</a> <span class="tocnum"><a href="#Page_24">24</a></span></li> +<li><a href="#UNIT_44">Simple-stemmed Plants</a> <span class="tocnum"><a href="#Page_26">26</a></span></li> +</ul></li> + +<li><a href="#Section_IV">SECTION IV. GROWTH FROM BUDS; BRANCHING</a> <span class="tocnum"><a href="#Page_27">27</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_47">Buds, situation and kinds</a> <span class="tocnum"><a href="#Page_27">27</a></span></li> +<li><a href="#UNIT_53">Vigorous vegetation from strong Buds</a> <span class="tocnum"><a href="#Page_28">28</a></span></li> +<li><a href="#UNIT_54">Arrangement of Branches</a> <span class="tocnum"><a href="#Page_29">29</a></span></li> +<li><a href="#UNIT_56">Non-developed</a>, <a href="#UNIT_57">Latent</a>, and <a href="#UNIT_59">Accessory Buds</a> <span class="tocnum"><a href="#Page_30">30</a></span></li> +<li><a href="#UNIT_60">Enumeration of kinds of Buds</a> <span class="tocnum"><a href="#Page_31">31</a></span></li> +<li><a href="#UNIT_61">Definite</a> and <a href="#UNIT_62">Indefinite growth</a>; <a href="#UNIT_63">Deliquescent and Excurrent</a> <span class="tocnum"><a href="#Page_31">31</a></span></li> +</ul></li> + +<li><a href="#Section_V">SECTION V. ROOTS</a> <span class="tocnum"><a href="#Page_33">33</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_66">Primary</a> and <a href="#UNIT_67">Secondary</a>. <a href="#UNIT_68">Contrast between Stem and Root</a> <span class="tocnum"><a href="#Page_34">34</a></span></li> +<li><a href="#UNIT_71">Fibrous</a> and <a href="#UNIT_74">Fleshy Roots</a>; names of kinds <span class="tocnum"><a href="#Page_34">34</a></span></li> +<li><a href="#UNIT_78">Anomalous Roots</a>. <a href="#UNIT_79">Epiphytic</a> and <a href="#UNIT_80">Parasitic Plants</a> <span class="tocnum"><a href="#Page_36">36</a></span></li> +<li><a href="#UNIT_83">Duration</a>: <a href="#UNIT_84">Annuals</a>, <a href="#UNIT_85">Biennials</a>, <a href="#UNIT_86">Perennials</a> <span class="tocnum"><a href="#Page_37">37</a></span></li> +</ul></li> + +<li><a href="#Section_VI">SECTION VI. STEMS</a> <span class="tocnum"><a href="#Page_38">38</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_89">Those above Ground: kinds and modifications</a> <span class="tocnum"><a href="#Page_39">39</a></span></li> +<li><a href="#UNIT_103">Subterranean Stems and Branches</a> <span class="tocnum"><a href="#Page_42">42</a></span></li> +<li><a href="#UNIT_104">Rootstock</a> <span class="tocnum"><a href="#Page_42">42</a></span></li> +<li><a href="#UNIT_110">Tuber</a> <span class="tocnum"><a href="#Page_44">44</a></span></li> +<li><a href="#UNIT_111">Corm</a> <span class="tocnum"><a href="#Page_45">45</a></span></li> +<li><a href="#UNIT_113">Bulb</a> and <a href="#UNIT_116">Bulblets</a> <span class="tocnum"><a href="#Page_46">46</a></span></li> +<li><a href="#UNIT_117">Consolidated Vegetation</a> <span class="tocnum"><a href="#Page_47">47</a></span></li> +</ul></li> + +<li><a href="#Section_VII">SECTION VII. LEAVES</a> <span class="tocnum"><a href="#Page_49">49</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_VII_1">§ 1. <span class="smcap">Leaves as Foliage</span></a> <span class="tocnum"><a href="#Page_49">49</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_122">Parts</a> and <a href="#UNIT_126">Venation</a> <span class="tocnum"><a href="#Page_50">50</a></span></li> +<li><a href="#UNIT_134">Forms as to general outline</a> <span class="tocnum"><a href="#Page_52">52</a></span></li> +<li><a href="#UNIT_137">As to apex and particular outline</a> <span class="tocnum"><a href="#Page_53">53</a></span></li> +<li><span class="pagenum"><a name="Page_vi" id="Page_vi">[Pg vi]</a></span><a href="#UNIT_141">As to lobing or division</a> <span class="tocnum"><a href="#Page_56">56</a></span></li> +<li><a href="#UNIT_146">Compound</a>, <a href="#UNIT_158">Perfoliate</a>, and <a href="#UNIT_160">Equitant Leaves</a> <span class="tocnum"><a href="#Page_57">57</a></span></li> +<li><a href="#UNIT_161">With no distinction of Petiole and Blade</a>, <a href="#UNIT_162">Phyllodia</a>, &c. <span class="tocnum"><a href="#Page_61">61</a></span></li> +</ul></li> +<li><a href="#Subsect_VII_2">§ 2. <span class="smcap">Leaves of Special Conformation and Use</span></a> <span class="tocnum"><a href="#Page_62">62</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_165">Leaves for storage</a> <span class="tocnum"><a href="#Page_62">62</a></span></li> +<li><a href="#UNIT_166">Leaves as bud-scales</a> <span class="tocnum"><a href="#Page_63">63</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_167">Spines</a> <span class="tocnum"><a href="#Page_64">64</a></span></li> +<li>and <a href="#UNIT_168">for Climbing</a> <span class="tocnum"><a href="#Page_64">64</a></span></li> +</ul></li> +<li><a href="#UNIT_171">Pitchers</a> <span class="tocnum"><a href="#Page_64">64</a></span> +<ul class="lsoff"> +<li>and <a href="#UNIT_173">Fly-traps</a> <span class="tocnum"><a href="#Page_65">65</a></span></li> +</ul></li> +</ul></li> +<li><a href="#Subsect_VII_3">§ 3. <span class="smcap">Stipules</span></a> <span class="tocnum"><a href="#Page_66">66</a></span></li> + +<li><a href="#Subsect_VII_4">§ 4. <span class="smcap">The Arrangement of Leaves</span></a> <span class="tocnum"><a href="#Page_67">67</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_181">Phyllotaxy</a> <span class="tocnum"><a href="#Page_67">67</a></span></li> +<li><a href="#UNIT_185">Of Alternate Leaves</a> <span class="tocnum"><a href="#Page_69">69</a></span></li> +<li><a href="#UNIT_193">Of Opposite and Whorled Leaves</a> <span class="tocnum"><a href="#Page_71">71</a></span></li> +<li><a href="#UNIT_194">Vernation or Præfoliation</a> <span class="tocnum"><a href="#Page_71">71</a></span></li> +</ul></li> +</ul></li> +<li><a href="#Section_VIII">SECTION VIII. FLOWERS</a> <span class="tocnum"><a href="#Page_72">72</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_VIII_1">§ 1. <span class="smcap">Position and Arrangement, Inflorescence</span></a> <span class="tocnum"><a href="#Page_73">73</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_205">Raceme</a> <span class="tocnum"><a href="#Page_73">73</a></span></li> +<li><a href="#UNIT_206">Corymb</a>, <a href="#UNIT_207">Umbel</a>, <a href="#UNIT_210">Spike</a>, <a href="#UNIT_211">Head</a> <span class="tocnum"><a href="#Page_74">74</a></span></li> +<li><a href="#UNIT_212">Spadix</a>, <a href="#UNIT_213">Catkin, or Ament</a> <span class="tocnum"><a href="#Page_75">75</a></span></li> +<li><a href="#UNIT_217">Panicle</a>: <a href="#UNIT_218">Determinate Inflorescence</a> <span class="tocnum"><a href="#Page_76">76</a></span></li> +<li><a href="#UNIT_219">Cyme</a>, <a href="#UNIT_221">Fascicle</a>, <a href="#UNIT_222">Glomerule</a>, <a href="#UNIT_225">Scorpioid or Helicoid Cymes</a> <span class="tocnum"><a href="#Page_77">77</a></span></li> +<li><a href="#UNIT_227">Mixed Inflorescence</a> <span class="tocnum"><a href="#Page_78">78</a></span></li> +</ul></li> +<li><a href="#Subsect_VIII_2">§ 2. <span class="smcap">Parts or Organs of the Flower</span></a> <span class="tocnum"><a href="#Page_79">79</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_229">Floral Envelopes: Perianth</a>, <a href="#UNIT_230">Calyx</a>, <a href="#UNIT_231">Corolla</a> <span class="tocnum"><a href="#Page_79">79</a></span></li> +<li><a href="#UNIT_234">Essential Organs</a>: <a href="#UNIT_235">Stamen</a>, <a href="#UNIT_236">Pistil</a> <span class="tocnum"><a href="#Page_80">80</a></span></li> +<li><a href="#UNIT_237">Torus or Receptacle</a> <span class="tocnum"><a href="#Page_81">81</a></span></li> +</ul></li> +<li><a href="#Subsect_VIII_3">§ 3. <span class="smcap">Plan of the Flower</span></a> <span class="tocnum"><a href="#Page_81">81</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_239">When perfect, complete, regular, or symmetrical</a> <span class="tocnum"><a href="#Page_81">81</a></span></li> +<li><a href="#UNIT_241">Numerical Plan</a> and <a href="#UNIT_242">Alternation of Organs</a> <span class="tocnum"><a href="#Page_82">82</a></span></li> +<li><a href="#UNIT_243">Flowers are altered branches</a> <span class="tocnum"><a href="#Page_83">83</a></span></li> +</ul></li> +<li><a href="#Subsect_VIII_4">§ 4. <span class="smcap">Modifications of the Type</span></a> <span class="tocnum"><a href="#Page_85">85</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_249">Unisexual or diclinous</a> <span class="tocnum"><a href="#Page_85">85</a></span></li> +<li><a href="#UNIT_251">Incomplete</a>, <a href="#UNIT_253">Irregular, and Unsymmetrical</a> <span class="tocnum"><a href="#Page_86">86</a></span></li> +<li><a href="#UNIT_255">Flowers with Multiplication of Parts</a> <span class="tocnum"><a href="#Page_88">88</a></span></li> +<li><a href="#UNIT_256">Flowers with Union of Parts</a>: <a href="#UNIT_257">Coalescence</a> <span class="tocnum"><a href="#Page_88">88</a></span></li> +<li><a href="#UNIT_258">Regular Forms</a> <span class="tocnum"><a href="#Page_89">89</a></span></li> +<li><a href="#UNIT_263">Irregular Forms</a> <span class="tocnum"><a href="#Page_90">90</a></span></li> +<li><a href="#UNIT_264">Papilionaceous</a> <span class="tocnum"><a href="#Page_91">91</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_265">Labiate</a> <span class="tocnum"><a href="#Page_92">92</a></span></li> +<li>and <a href="#UNIT_267">Ligulate Corollas</a> <span class="tocnum"><a href="#Page_93">93</a></span></li> +</ul></li> +<li><a href="#UNIT_269">Adnation or Consolidation</a> <span class="tocnum"><a href="#Page_94">94</a></span></li> +<li><a href="#UNIT_273">Position of Flower or of its Parts</a> <span class="tocnum"><a href="#Page_96">96</a></span></li> +</ul></li> +<li><a href="#Subsect_VIII_5">§ 5. <span class="smcap">Arrangement of Parts in the Bud</span></a> <span class="tocnum"><a href="#Page_97">97</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_276">Æstivation or Præfloration, its kinds</a> <span class="tocnum"><a href="#Page_97">97</a></span></li> +</ul></li> +</ul></li> + +<li><a href="#Section_IX">SECTION IX. STAMENS IN PARTICULAR</a> <span class="tocnum"><a href="#Page_98">98</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_281">Andrœcium</a> <span class="tocnum"><a href="#Page_98">98</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_283">Insertion, Relation, &c.</a> <span class="tocnum"><a href="#Page_99">99</a></span></li> +</ul></li> +<li><a href="#UNIT_287">Anther</a> and <a href="#UNIT_286">Filament</a>. <a href="#UNIT_296">Pollen</a> <span class="tocnum"><a href="#Page_101">101</a></span></li> +</ul></li> + +<li><a href="#Section_X">SECTION X. PISTILS IN PARTICULAR</a> <span class="tocnum"><a href="#Page_105">105</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_X_1">§ 1. <span class="smcap">Angiospermous or Ordinary Gynœcium</span></a> <span class="tocnum"><a href="#Page_105">105</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_302">Parts of a complete Pistil</a> <span class="tocnum"><a href="#Page_105">105</a></span></li> +<li><a href="#UNIT_305">Carpels</a>, <a href="#UNIT_306">Simple Pistil</a> <span class="tocnum"><a href="#Page_106">106</a></span></li> +<li><span class="pagenum"><a name="Page_vii" id="Page_vii">[Pg vii]</a></span><a href="#UNIT_309">Compound Pistil</a> <a href="#UNIT_310">with Cells and Axile Placentæ</a> <span class="tocnum"><a href="#Page_107">107</a></span></li> +<li><a href="#UNIT_311">One-celled with Free Central Placenta</a> <span class="tocnum"><a href="#Page_108">108</a></span></li> +<li><a href="#UNIT_312">One-celled with Parietal Placentæ</a> <span class="tocnum"><a href="#Page_108">108</a></span></li> +</ul></li> +<li><a href="#Subsect_X_2">§ 2. <span class="smcap">Gymnospermous Gynœcium</span></a> <span class="tocnum"><a href="#Page_109">109</a></span></li> +</ul></li> + +<li><a href="#Section_XI">SECTION XI. OVULES</a> <span class="tocnum"><a href="#Page_110">110</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_317">Their Parts, Insertion</a>, and <a href="#UNIT_321">Kinds</a> <span class="tocnum"><a href="#Page_111">111</a></span></li> +</ul></li> + +<li><a href="#Section_XII">SECTION XII. MODIFICATIONS OF THE RECEPTACLE</a> <span class="tocnum"><a href="#Page_112">112</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_323">Torus, Stipe</a>, <a href="#UNIT_324">Carpophore</a>, <a href="#UNIT_327">Disk</a> <span class="tocnum"><a href="#Page_113">113</a></span></li> +</ul></li> + +<li><a href="#Section_XIII">SECTION XIII. FERTILIZATION</a> <span class="tocnum"><a href="#Page_114">114</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_XIII_1">§ 1. <span class="smcap">Adaptations for Pollination of the Stigma</span></a> <span class="tocnum"><a href="#Page_114">114</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_332">Close</a> and <a href="#UNIT_333">Cross Fertilization</a>, <a href="#UNIT_334">Anemophilous</a> and <a href="#UNIT_335">Entomophilous</a> <span class="tocnum"><a href="#Page_115">115</a></span></li> +<li><a href="#UNIT_339">Dichogamy</a> and <a href="#UNIT_341">Heterogony</a> <span class="tocnum"><a href="#Page_116">116</a></span></li> +</ul></li> +<li><a href="#Subsect_XIII_2">§ 2. <span class="smcap">Action of the Pollen and Formation of the Embryo</span></a> <span class="tocnum"><a href="#Page_117">117</a></span></li> +</ul></li> + +<li><a href="#Section_XIV">SECTION XIV. THE FRUIT</a> <span class="tocnum"><a href="#Page_117">117</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_345">Nature</a> and <a href="#UNIT_347">kinds</a> <span class="tocnum"><a href="#Page_118">118</a></span></li> +<li><a href="#UNIT_352">Berry</a>, <a href="#UNIT_353">Pepo</a>, <a href="#UNIT_354">Pome</a> <span class="tocnum"><a href="#Page_119">119</a></span></li> +<li><a href="#UNIT_356">Drupe</a> and <a href="#UNIT_359">Akene</a> <span class="tocnum"><a href="#Page_120">120</a></span></li> +<li><a href="#UNIT_361">Cremocarp</a>, <a href="#UNIT_363">Caryopsis</a>, <a href="#UNIT_364">Nut</a> <span class="tocnum"><a href="#Page_121">121</a></span></li> +<li><a href="#UNIT_367">Follicle</a>, <a href="#UNIT_368">Legume</a>, <a href="#UNIT_370">Capsule</a> <span class="tocnum"><a href="#Page_122">122</a></span></li> +<li><a href="#UNIT_371">Capsular Dehiscence</a>, <a href="#UNIT_374">Silique</a> and <a href="#UNIT_375">Silicle</a> <span class="tocnum"><a href="#Page_123">123</a></span></li> +<li><a href="#UNIT_376">Pyxis</a>, <a href="#UNIT_379">Strobile or Cone</a> <span class="tocnum"><a href="#Page_124">124</a></span></li> +</ul></li> + +<li><a href="#Section_XV">SECTION XV. THE SEED</a> <span class="tocnum"><a href="#Page_125">125</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_381">Seed-coats</a> and <a href="#UNIT_383">their appendages</a> <span class="tocnum"><a href="#Page_125">125</a></span></li> +<li><a href="#UNIT_386">The Kernel or Nucleus, Embryo and its parts, Albumen</a> <span class="tocnum"><a href="#Page_127">127</a></span></li> +</ul></li> + +<li><a href="#Section_XVI">SECTION XVI. VEGETABLE LIFE AND WORK</a> <span class="tocnum"><a href="#Page_128">128</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_XVI_1">§ 1. <span class="smcap">Anatomical Structure and Growth</span></a> <span class="tocnum"><a href="#Page_129">129</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_394">Nature of Growth</a>, <a href="#UNIT_396">Protoplasm</a> <span class="tocnum"><a href="#Page_129">129</a></span></li> +<li><a href="#UNIT_399">Cells and Cell-walls</a>. <a href="#UNIT_401">Cellular Structure or Tissue</a> <span class="tocnum"><a href="#Page_130">130</a></span></li> +<li><a href="#UNIT_407">Strengthening Cells</a>. <a href="#UNIT_408">Wood</a>, <a href="#UNIT_410">Wood-cells</a>, <a href="#UNIT_413">Vessels or Ducts</a> <span class="tocnum"><a href="#Page_132">132</a></span></li> +</ul></li> +<li><a href="#Subsect_XVI_2">§ 2. <span class="smcap">Cell-contents</span></a> <span class="tocnum"><a href="#Page_136">136</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_415">Sap</a>, <a href="#UNIT_417">Chlorophyll</a>, <a href="#UNIT_419">Starch</a> <span class="tocnum"><a href="#Page_136">136</a></span></li> +<li><a href="#UNIT_422">Crystals, Rhaphides</a> <span class="tocnum"><a href="#Page_137">137</a></span></li> +</ul></li> +<li><a href="#Subsect_XVI_3">§ 3. <span class="smcap">Anatomy of Roots and Stems</span></a> <span class="tocnum"><a href="#Page_138">138</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_427">Endogenous</a> and <a href="#UNIT_428">Exogenous Stems</a> <span class="tocnum"><a href="#Page_139">139</a></span></li> +<li><a href="#UNIT_431">Particular structure of the latter</a> <span class="tocnum"><a href="#Page_140">140</a></span></li> +<li><a href="#UNIT_434">Wood</a>, <a href="#UNIT_436">Sapwood</a> and <a href="#UNIT_437">Heart-wood</a>. <a href="#UNIT_438">The living parts of a Tree</a> <span class="tocnum"><a href="#Page_141">141</a></span></li> +</ul></li> +<li><a href="#Subsect_XVI_4">§ 4. <span class="smcap">Anatomy of Leaves</span></a> <span class="tocnum"><a href="#Page_142">142</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_442">Epidermis</a>, <a href="#UNIT_443">Stomata or Breathing pores</a> <span class="tocnum"><a href="#Page_143">143</a></span></li> +</ul></li> +<li><a href="#Subsect_XVI_5">§ 5. <span class="smcap">Plant Food and Assimilation</span></a> <span class="tocnum"><a href="#Page_144">144</a></span></li> + +<li><a href="#Subsect_XVI_6">§ 6. <span class="smcap">Plant Work and Movement</span></a> <span class="tocnum"><a href="#Page_149">149</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_461">Movements in Cells or Cyclosis</a> <span class="tocnum"><a href="#Page_149">149</a></span></li> +<li><a href="#UNIT_462">Transference from Cell to Cell</a> <span class="tocnum"><a href="#Page_150">150</a></span></li> +<li><span class="pagenum"><a name="Page_viii" id="Page_viii">[Pg viii]</a></span><a href="#UNIT_465">Movements of Organs</a>, <a href="#UNIT_468">Twining Stems</a>, <a href="#UNIT_469">Leaf-movements</a> <span class="tocnum"><a href="#Page_150">150</a></span></li> +<li><a href="#UNIT_471">Movements of Tendrils</a>, <a href="#UNIT_472">Sensitiveness</a> <span class="tocnum"><a href="#Page_152">152</a></span></li> +<li><a href="#UNIT_475">Movements in Flowers</a> <span class="tocnum"><a href="#Page_153">153</a></span></li> +<li><a href="#UNIT_476">Movements for capture of Insects</a> <span class="tocnum"><a href="#Page_154">154</a></span></li> +<li><a href="#UNIT_480">Work costs, using up Material and Energy</a> <span class="tocnum"><a href="#Page_155">155</a></span></li> +</ul></li> +</ul></li> + +<li><a href="#Section_XVII">SECTION XVII. CRYPTOGAMOUS OR FLOWERLESS PLANTS</a> <span class="tocnum"><a href="#Page_156">156</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_483">Vascular Cryptogams</a>, <a href="#UNIT_484">Pteridophytes</a> <span class="tocnum"><a href="#Page_156">156</a></span></li> +<li><a href="#UNIT_485">Horsetails (Equisetaceæ)</a>, <a href="#UNIT_486">Ferns</a> <span class="tocnum"><a href="#Page_157">157</a></span></li> +<li><a href="#UNIT_492">Club-Mosses (Lycopodium), &c.</a> <span class="tocnum"><a href="#Page_161">161</a></span></li> +<li><a href="#UNIT_494">Quillworts (Isoetes)</a>, <a href="#UNIT_495">Pillworts (Marsilia)</a> <span class="tocnum"><a href="#Page_161">161</a></span></li> +<li><a href="#UNIT_496">Azolla</a>. <a href="#UNIT_497">Cellular Cryptogams</a> <span class="tocnum"><a href="#Page_162">162</a></span></li> +<li><a href="#UNIT_498">Bryophytes</a>. <a href="#UNIT_499">Mosses (Musci)</a> <span class="tocnum"><a href="#Page_163">163</a></span></li> +<li><a href="#UNIT_501">Liverworts (Hepaticæ)</a> <span class="tocnum"><a href="#Page_164">164</a></span></li> +<li><a href="#UNIT_503">Thallophytes</a> <span class="tocnum"><a href="#Page_165">165</a></span></li> +<li><a href="#UNIT_506">Characeæ</a> <span class="tocnum"><a href="#Page_167">167</a></span></li> +<li><a href="#UNIT_507">Algæ, Seaweeds, &c.</a> <span class="tocnum"><a href="#Page_168">168</a></span></li> +<li><a href="#UNIT_515">Lichenes or Lichens</a> <span class="tocnum"><a href="#Page_171">171</a></span></li> +<li><a href="#UNIT_516">Fungi</a> <span class="tocnum"><a href="#Page_172">172</a></span></li> +</ul></li> + +<li><a href="#Section_XVIII">SECTION XVIII. CLASSIFICATION AND NOMENCLATURE</a> <span class="tocnum"><a href="#Page_175">175</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_XVIII_1">§ 1. <span class="smcap">Kinds and Relationship</span></a> <span class="tocnum"><a href="#Page_175">175</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_522">Species</a>, <a href="#UNIT_524">Varieties</a>, <a href="#UNIT_521">Individuals</a> <span class="tocnum"><a href="#Page_176">176</a></span></li> +<li><a href="#UNIT_528">Genera</a>, <a href="#UNIT_529">Orders</a>, <a href="#UNIT_531">Classes</a>, &c. <span class="tocnum"><a href="#Page_177">177</a></span></li> +</ul></li> +<li><a href="#Subsect_XVIII_2">§ 2. <span class="smcap">Names, Terms and Characters</span></a> <span class="tocnum"><a href="#Page_178">178</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_536">Nomenclature of Genera</a>, <a href="#UNIT_537">Species</a>, and <a href="#UNIT_538">Varieties</a> <span class="tocnum"><a href="#Page_179">179</a></span></li> +<li><a href="#UNIT_540">Nomenclature of Orders</a>, <a href="#UNIT_542">Classes</a>, &c. <a href="#UNIT_545">Terminology</a> <span class="tocnum"><a href="#Page_180">180</a></span></li> +</ul></li> +<li><a href="#Subsect_XVIII_3">§ 3. <span class="smcap">System</span></a> <span class="tocnum"><a href="#Page_181">181</a></span> +<ul class="lsoff"> +<li><a href="#UNIT_549">Artificial</a> and <a href="#UNIT_550">Natural</a> <span class="tocnum"><a href="#Page_182">182</a></span></li> +<li><a href="#UNIT_554">Synopsis of Series, Classes, &c.</a> <span class="tocnum"><a href="#Page_183">183</a></span></li> +</ul></li> +</ul></li> + +<li><a href="#Section_XIX">SECTION XIX. BOTANICAL WORK</a> <span class="tocnum"><a href="#Page_184">184</a></span> +<ul class="lsoff"> +<li><a href="#Subsect_XIX_1">§ 1. <span class="smcap">Collection or Herborization</span></a> <span class="tocnum"><a href="#Page_184">184</a></span></li> + +<li><a href="#Subsect_XIX_2">§ 2. <span class="smcap">Herbarium</span></a> <span class="tocnum"><a href="#Page_186">186</a></span></li> + +<li><a href="#Subsect_XIX_3">§ 3. <span class="smcap">Investigation and Determination of Plants</span></a> <span class="tocnum"><a href="#Page_187">187</a></span></li> + +<li><a href="#Subsect_XIX_4">§ 4. <span class="smcap">Signs and Abbreviations</span></a> <span class="tocnum"><a href="#Page_188">188</a></span></li> +</ul></li> + +<li><a href="#ABBREVIATIONS"><span class="smcap">Abbreviations of the Names of Botanists</span></a> <span class="tocnum"><a href="#Page_190">190</a></span></li> + + +<li><a href="#INDEX"><span class="smcap">Glossary combined with Index</span></a> <span class="tocnum"><a href="#Page_193">193</a></span></li> +</ul></div> + + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_9" id="Page_9">[Pg 9]</a></span></p> +<div class="likeheading1"><a name="ELEMENTS_OF_BOTANY" id="ELEMENTS_OF_BOTANY"></a>ELEMENTS OF BOTANY.</div> + + + +<hr class="chap" /> +<h2><a name="Section_I" id="Section_I"></a><span class="smcap">Section I.</span> INTRODUCTORY.</h2> + + +<p>1. <span class="smcap">Botany</span> is the name of the science of the vegetable kingdom in +general; that is, of plants.</p> + +<p>2. Plants may be studied as to their kinds and relationships. This study +is <span class="smcap">Systematic Botany</span>. An enumeration of the kinds of vegetables, as far +as known, classified according to their various degrees of resemblance +or difference, constitutes a general <i>System of plants</i>. A similar +account of the vegetables of any particular country or district is +called a <i>Flora</i>.</p> + +<p><a name="UNIT_3" id="UNIT_3">3.</a> Plants may be studied as to their structure and parts. This is +<span class="smcap">Structural Botany</span>, or <span class="smcap">Organography</span>. The study of the organs or parts of +plants in regard to the different forms and different uses which the +same kind of organ may assume,—the comparison, for instance, of a +flower-leaf or a bud-scale with a common leaf,—is <span class="smcap">Vegetable Morphology</span>, +or <span class="smcap">Morphological Botany</span>. The study of the minute structure of the parts, +to learn by the microscope what they themselves are formed of, is +<span class="smcap">Vegetable Anatomy</span>, or <span class="smcap">Histology</span>; in other words, it is Microscopical +Structural Botany. The study of the actions of plants or of their parts, +of the ways in which a plant lives, grows, and acts, is the province of +<span class="smcap">Physiological Botany</span>, or <span class="smcap">Vegetable Physiology</span>.</p> + +<p>4. This book is to teach the outlines of Structural Botany and of the +simpler parts of the physiology of plants, that it may be known how +plants are constructed and adapted to their surroundings, and how they +live, move, propagate, and have their being in an existence no less +real, although more simple, than that of the animal creation which they +support. Particularly, this book is to teach the principles of the +structure and relationships of plants, the nature and names of their +parts and their modifications, and so to prepare for the study of +Systematic Botany; in which the learner may ascertain the name and the +place in the system of any or all of the ordinary plants within reach, +whether wild or cultivated. And in ascertaining the name of any plant, +the student, if rightly taught, will come to know all about its general +or particular structure, rank, and relationship to other plants.</p> + +<p><span class="pagenum"><a name="Page_10" id="Page_10">[Pg 10]</a></span></p><p>5. The vegetable kingdom is so vast and various, and the difference is +so wide between ordinary trees, shrubs, and herbs on the one hand, and +mosses, moulds, and such like on the other, that it is hardly possible +to frame an intelligible account of plants as a whole without +contradictions or misstatements, or endless and troublesome +qualifications. If we say that plants come from seeds, bear flowers, and +have roots, stems, and leaves, this is not true of the lower orders. It +is best for the beginner, therefore, to treat of the higher orders of +plants by themselves, without particular reference to the lower.</p> + +<p><a name="UNIT_6" id="UNIT_6">6.</a> Let it be understood, accordingly, that there is a higher and a lower +series of plants; namely:—</p> + +<p><span class="smcap">Phanerogamous Plants</span>, which come from seed and bear <i>flowers</i>, +essentially stamens and pistils, through the co-operation of which seed +is produced. For shortness, these are commonly called <span class="smcap">Phanerogams</span>, or +<i>Phænogams</i>, or by the equivalent English name of <span class="smcap">Flowering Plants</span>.<a name="FNanchor_1_1" id="FNanchor_1_1"></a><a href="#Footnote_1_1" class="fnanchor">[1]</a></p> + +<p><span class="smcap">Cryptogamous Plants</span>, or <span class="smcap">Cryptogams</span>, come from minute bodies, which +answer to seeds, but are of much simpler structure, and such plants have +not stamens and pistils. Therefore they are called in English <span class="smcap">Flowerless +Plants</span>. Such are Ferns, Mosses, Algæ or Seaweeds, Fungi, etc. These +sorts have each to be studied separately, for each class or order has a +plan of its own.</p> + +<p>7. But Phanerogamous, or Flowering, Plants are all constructed on one +plan, or <i>type</i>. That is, taking almost any ordinary herb, shrub, or +tree for a pattern, it will exemplify the whole series: the parts of one +plant answer to the parts of any other, with only certain differences in +particulars. And the occupation and the delight of the scientific +botanist is in tracing out this common plan, in detecting the likenesses +under all the diversities, and in noting the meaning of these manifold +diversities. So the attentive study of any one plant, from its growth +out of the seed to the flowering and fruiting state and the production +of seed like to that from which the plant grew, would not only give a +correct general idea of the structure, growth, and characteristics of +Flowering Plants in general, but also serve as a pattern or standard of +comparison. Some plants will serve this purpose of a pattern much better +than others. A proper pattern will be one that is perfect in the sense +of having all the principal parts of a phanerogamous plant, and simple +and regular in having these parts free from complications or disguises. +The common Flax-plant may very well serve this purpose. Being an annual, +it has the advantage of being easily raised and carried in a short time +through its circle of existence, from seedling to fruit and seed.</p> + + +<div class="footnotes"><h3>FOOTNOTES:</h3> + +<div class="footnote"><p><a name="Footnote_1_1" id="Footnote_1_1"></a><a href="#FNanchor_1_1"><span class="label">[1]</span></a> The name is sometimes <i>Phanerogamous</i>, sometimes +<i>Phænogamous</i> (<i>Phanerogams</i>, or <i>Phænogams</i>), terms of the same meaning +etymologically; the former of preferable form, but the latter shorter. +The meaning of such terms is explained in the Glossary.</p></div> +</div> + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_11" id="Page_11">[Pg 11]</a></span></p> +<h2><a name="Section_II" id="Section_II"></a><span class="smcap">Section II.</span> FLAX AS A PATTERN PLANT.</h2> + + +<p><a name="UNIT_8" id="UNIT_8">8.</a> <b>Growth from the Seed.</b> Phanerogamous plants grow from seed, and their +flowers are destined to the production of seeds. A seed has a +rudimentary plant ready formed in it,—sometimes with the two most +essential parts, i. e. stem and leaf, plainly discernible; sometimes +with no obvious distinction of organs until germination begins. This +incipient plant is called an <span class="smcap">Embryo</span>.</p> + +<p>9. In this section the Flax-plant is taken as a specimen, or type, and +the development and history of common plants in general is illustrated +by it. In flax-seed the embryo nearly fills the coats, but not quite. +There is a small deposit of nourishment between the seed-coat and the +embryo: this may for the present be left out of the account. This embryo +consists of a pair of leaves, pressed together face to face, and +attached to an extremely short stem. (Fig. <a href="#Fig1">2-4</a>.) In this rudimentary +condition the real nature of the parts is not at once apparent; but when +the seed grows they promptly reveal their character,—as the +accompanying figures (Fig. <a href="#Fig5">5-7</a>) show.</p> + +<div class="figcenter"> +<a name="Fig1" id="Fig1"></a> +<img src="images/fig001_004.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 1. Pod of Flax. 2. Section lengthwise, showing two +of the seeds; one whole, the other cut half away, bringing contained +embryo into view. 3. Similar section of a flax-seed more magnified and +divided flatwise; turned round, so that the stem-end (caulicle) of the +embryo is below: the whole broad upper part is the inner face of one of +the cotyledons; the minute nick at its base is the plumule. 4. Similar +section through a seed turned edgewise, showing the thickness of the +cotyledons, and the minute plumule between them, i. e. the minute bud on +the upper end of the caulicle.</p> +</div> + +<p><a name="UNIT_10" id="UNIT_10">10.</a> Before the nature of these parts in the seed was altogether +understood, technical names were given to them, which are still in use. +These initial leaves were named <span class="smcap">Cotyledons</span>. The initial stem on which +they stand was called the <span class="smcap">Radicle</span>. That was because it gives rise to the +first root; but, as it is really the beginning of the stem, and because +it is the stem that produces the root and not the root that produces the +stem, it is better to name it the <span class="smcap">Caulicle</span>. Recently it has been named +<i>Hypocotyle</i>; which signifies something below the cotyledons, without +pronouncing what its nature is.</p> + +<p><span class="pagenum"><a name="Page_12" id="Page_12">[Pg 12]</a></span></p> +<div class="figcenter"> +<a name="Fig5" id="Fig5"></a> +<img src="images/fig005_007.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 5. Early Flax seedling; stem (caulicle), root at +lower end, expanded seed-leaves (cotyledons) at the other: minute bud +(plumule) between these. 6. Same later; the bud developed into second +pair of leaves, with hardly any stem-part below them; then into a third +pair of leaves, raised on a short joint of stem; and a fifth leaf also +showing. 7. Same still older, with more leaves developed, but these +singly (one after another), and with joints of stem between them.</p> +</div> + +<p><a name="UNIT_11" id="UNIT_11">11.</a> On committing these seeds to moist and warm soil they soon sprout, +i. e. <i>germinate</i>. The very short stem-part of the embryo is the first +to grow. It lengthens, protrudes its root-end; this turns downward, if +not already pointing in that direction, and while it is lengthening a +root forms at its point and grows downward into the ground. This root +continues to grow on from its lower end, and thus insinuates itself and +penetrates into the soil. The stem meanwhile is adding to its length +throughout; it erects itself, and, seeking the light, brings the seed up +out of the ground. The materials for this growth have been supplied by +the cotyledons or seed-leaves, still in the seed: it was the store of +nourishing material they held which gave them their thickish shape, so +unlike that of ordinary leaves. Now, relieved of a part of this store of +food, which has formed the growth by which they have been raised into +the air and light, they appropriate the remainder to their own growth. +In enlarging they open and throw off the seed-husk; they expand, diverge +into a horizontal position, turn green, and thus become a pair of +evident leaves, the first foliage of a tiny plant. This seedling, +although diminutive and most simple, possesses and puts into use, all +the <span class="smcap">Organs</span> of <span class="smcap">Vegetation</span>, namely, root, stem, and leaves, each in its +proper element,—the root in the soil, the stem rising out of it, the +leaves in the light and open air. It now draws in moisture and some +food-materials from the soil by its root, conveys this through the stem +into the leaves, where these materials, along with other crude food +which these imbibe from the air, are assimilated into vegetable matter, +i. e. into the material for further growth.</p> + +<p><a name="UNIT_12" id="UNIT_12">12.</a> <b>Further Growth</b> soon proceeds to the formation of new +parts,—downward in the production of more root, or of branches of the +main root, upward in the development of more stem and leaves. That from +which a stem with its leaves is continued, or a new stem (i. e. branch) +originated, is a <span class="smcap">Bud</span>. The most conspicuous and familiar buds are those +of most shrubs and trees, bearing buds formed in summer or autumn, to +grow the following <span class="pagenum"><a name="Page_13" id="Page_13">[Pg 13]</a></span>spring. But every such point for new growth may +equally bear the name. When there is such a bud between the cotyledons +in the seed or seedling it is called the <span class="smcap">Plumule</span>. This is conspicuous +enough in a bean (Fig. <a href="#Fig26">29</a>.), where the young leaf of the new growth +looks like a little plume, whence the name, <i>plumule</i>. In flax-seed this +is very minute indeed, but is discernible with a magnifier, and in the +seedling it shows itself distinctly (Fig. <a href="#Fig5">5, 6, 7</a>).</p> + +<p><a name="UNIT_13" id="UNIT_13">13.</a> As it grows it shapes itself into a second pair of leaves, which of +course rests on a second joint of stem, although in this instance that +remains too short to be well seen. Upon its summit appears the third +pair of leaves, soon to be raised upon its proper joint of stem; the +next leaf is single, and is carried up still further upon its supporting +joint of stem; and so on. The root, meanwhile, continues to grow +underground, not joint after joint, but continuously, from its lower +end; and commonly it before long multiplies itself by branches, which +lengthen by the same continuous growth. But stems are built up by a +succession of leaf-bearing growths, such as are strongly marked in a +reed or corn-stalk, and less so in such an herb as Flax. The word +"joint" is ambiguous: it may mean either the portion between successive +leaves, or their junction, where the leaves are attached. For precision, +therefore, the place where the leaf or leaves are borne is called a +<span class="smcap">Node</span>, and the naked interval between two nodes, an <span class="smcap">Internode</span>.</p> + +<div class="figcenter"> +<a name="Fig8" id="Fig8"></a> +<img src="images/fig008.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 8. Upper part of Flax-plant in blossom. +</div> + +<p>14. In this way a simple stem with its garniture of leaves is developed +from the seed. But besides this direct continuation, buds may form and +develop into lateral stems, that is, <i>into branches</i>, from any node. The +proper origin of branches is from the <span class="smcap">Axil</span> of a leaf, i. e. the angle +between leaf and stem on the upper side; and branches may again branch, +so building up the herb, shrub, or tree. But sooner or later, and +without long delay in an annual like Flax, instead of this continuance +of mere vegetation, reproduction is prepared for by</p> + +<p><span class="pagenum"><a name="Page_14" id="Page_14">[Pg 14]</a></span></p><p><a name="UNIT_15" id="UNIT_15">15.</a> <b>Blossoming.</b> In Flax the flowers make their appearance at the end of +the stem and branches. The growth, which otherwise might continue them +farther or indefinitely, now takes the form of blossom, and is +subservient to the production of seed.</p> + +<div class="figcenter"> +<a name="Fig9" id="Fig9"></a> +<img src="images/fig009_010.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 9. Flax-flowers about natural size. 10. Section of a +flower moderately enlarged, showing a part of the petals and stamens, +all five styles, and a section of ovary with two ovules or rudimentary +seeds.</p> +</div> + +<p><a name="UNIT_16" id="UNIT_16">16.</a> <b>The Flower</b> of Flax consists, first, of five small green leaves, +crowded into a circle: this is the <span class="smcap">Calyx</span>, or flower-cup. When its +separate leaves are referred to they are called <span class="smcap">Sepals</span>, a name which +distinguishes them from foliage-leaves on the one hand, and from petals +on the other. Then come five delicate and <i>colored</i> leaves (in the Flax, +blue), which form the <span class="smcap">Corolla</span>, and its leaves are <span class="smcap">Petals</span>; then a circle +of organs, in which all likeness to leaves is lost, consisting of +slender stalks with a knob at summit, the <span class="smcap">Stamens</span>; and lastly, in the +centre, the rounded body, which becomes a pod, surmounted by five +slender or stalk-like bodies. This, all together, is the <span class="smcap">Pistil</span>. The +lower part of it, which is to contain the seeds, is the <span class="smcap">Ovary</span>; the +slender organs surmounting this are <span class="smcap">Styles</span>; the knob borne on the apex +of each style is a <span class="smcap">Stigma</span>. Going back to the stamens, these are of two +parts, viz. the stalk, called <span class="smcap">Filament</span>, and the body it bears, the +<span class="smcap">Anther</span>. Anthers are filled with <span class="smcap">Pollen</span>, a powdery substance made up of +minute grains.</p> + +<p>17. The pollen shed from the anthers when they open falls upon or is +conveyed to the stigmas; then the pollen-grains set up a kind of growth +(to be discerned only by aid of a good microscope), which penetrates the +style: this growth takes the form of a thread more delicate than the +finest spider's web, and reaches the bodies which are to become seeds +(<span class="smcap">Ovules</span> they are called until this change occurs); these, touched by +this influence, are incited to a new growth within, which becomes an +embryo. So, as the ovary ripens into the seed-pod or capsule (Fig. <a href="#Fig1">1</a>, +etc.) containing seeds, each seed enclosing a rudimentary new plantlet, +the round of this vegetable existence is completed.</p> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_15" id="Page_15">[Pg 15]</a></span></p> +<h2><a name="Section_III" id="Section_III"></a><span class="smcap">Section III.</span> MORPHOLOGY OF SEEDLINGS.</h2> + + +<p>18. Having obtained a general idea of the growth and parts of a +phanerogamous plant from the common Flax of the field, the seeds and +seedlings of other familiar plants may be taken up, and their variations +from the assumed pattern examined.</p> + +<p><a name="UNIT_19" id="UNIT_19">19.</a> <b>Germinating Maples</b> are excellent to begin with, the parts being so +much larger than in Flax that a common magnifying glass, although +convenient, is hardly necessary. The only disadvantage is that fresh +seeds are not readily to be had at all seasons.</p> + +<div class="figcenter"> +<a name="Fig11" id="Fig11"></a> +<img src="images/fig011_013.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 11. Embryo of Sugar Maple, cut through lengthwise +and taken out of the seed. 12, 13. Whole embryo of same just beginning +to grow; <i>a</i>, the stemlet or caulicle, which in 13 has considerably +lengthened.</p> +</div> + +<p>20. The seeds of Sugar Maple ripen at the end of summer, and germinate +in early spring. The embryo fills the whole seed, in which it is nicely +packed; and the nature of the parts is obvious even before growth +begins. There is a stemlet (caulicle) and a pair of long and narrow +seed-leaves (cotyledons), doubled up and coiled, green even in the seed, +and in germination at once unfolding into the first pair of +foliage-leaves, though of shape quite unlike those that follow.</p> + +<p>21. Red Maple seeds are ripe and ready to germinate at the beginning of +summer, and are therefore more convenient for study. The cotyledons are +crumpled in the seed, and not easy to straighten out until they unfold +themselves in germination. The story of their development into the +seedling is told by the accompanying Fig. <a href="#Fig14">14-20</a>; and that of Sugar Maple +is closely similar. No plumule or bud appears in the embryo of these two +Maples until the seed-leaves have nearly attained their full growth and +are acting as foliage-leaves, and until a root is formed below. There is +no great store of nourishment in these thin cotyledons; so further +growth has to wait until the root and seed-leaves have collected and +elaborated sufficient material for the formation of the second internode +and its pair of leaves, which lending their help the third pair is more +promptly produced, and so on.</p> + +<p><a name="UNIT_22" id="UNIT_22">22.</a> Some change in the plan comes with the Silver or Soft White Maple. +(Fig. <a href="#Fig21">21-25</a>). This blossoms in earliest spring, and it drops its large +and ripened keys only a few weeks later. Its cotyledons have not at all +the appearance of leaves; they are short and broad, and (as there is no +room to be saved by folding) they are straight, except a small fold at +the top,—a vestige of the habit of Maples in general. Their unusual +thickness is due <span class="pagenum"><a name="Page_16" id="Page_16">[Pg 16]</a></span>to the large store of nutritive matter they contain, +and this prevents their developing into actual leaves. Correspondingly, +their caulicle does not lengthen to elevate them above the surface of +the soil; the growth below the cotyledons is nearly all of root. It is +the little plumule or bud between them which makes the upward growth, +and which, being well fed by the cotyledons, rapidly develops the next +pair of leaves and raises them upon a long internode, and so on. The +cotyledons all the while remain below, in the husk of the fruit and +seed, and perish when they have yielded up the store of food which they +contained.</p> + +<div class="figcenter"> +<a name="Fig14" id="Fig14"></a> +<img src="images/fig014_020.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 14. One of the pair of keys or winged fruits of Red +Maple; the seed-bearing portion cut open to show the seed. 15. Seed +enlarged, and divided to show the crumpled embryo which fills it. 16. +Embryo taken out and partly opened. 17. Embryo which has unfolded in +early stage of germination and begun to grow. 18. Seedling with next +joint of stem and leaves apparent; and 19 with these parts full-grown, +and bud at apex for further growth. 20. Seedling with another joint of +stem and pair of leaves.</p> +</div> + +<p>23. So, even in plants so much alike as Maples, there is considerable +difference in the amount of food stored up in the cotyledons by which +the growth is to be made; and there are corresponding differences in the +germination. <span class="pagenum"><a name="Page_17" id="Page_17">[Pg 17]</a></span>The larger the supply to draw upon, the stronger the +growth, and the quicker the formation of root below and of stem and +leaves above. This deposit of food thickens the cotyledons, and renders +them less and less leaf-like in proportion to its amount.</p> + +<div class="figcenter"> +<a name="Fig21" id="Fig21"></a> +<img src="images/fig021_025.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 21. Fruit (one key) of Silver Maple, Acer +dasycarpum, of natural size, the seed-bearing portion divided to show +the seed. 22. Embryo of the seed taken out. 23. Same opened out, to show +the thick cotyledons and the little plumule or bud between them. 24. +Germination of Silver Maple, natural size; merely the base of the fruit, +containing the seed, is shown. 25. Embryo of same, taken out of the +husk; upper part of growing stem cut off, for want of room.</p> +</div> + +<p><a name="UNIT_24" id="UNIT_24">24.</a> <b>Examples of Embryos with thickened Cotyledons.</b> In the Pumpkin and +Squash (Fig. <a href="#Fig26">26, 27</a>), the cotyledons are well supplied with nourishing +matter, as their sweet taste demonstrates. Still, they are flat and not +very thick. In germination this store is promptly utilized in the +development of the caulicle to twenty or thirty times its length in the +seed, and to corresponding thickness, in the formation of a cluster of +roots at its lower end, and the early production of the incipient +plumule; also in their own growth into efficient green leaves. The case +of our common Bean (Phaseolus vulgaris, Fig. <a href="#Fig26">28-30</a>) is nearly the same, +except that the cotyledons are much more gorged; so that, although +carried up into the air and light upon the lengthening caulicle, and +there acquiring a green color, they never expand into useful leaves. +Instead of this, they nourish into rapid growth the plumule, which is +plainly visible in the seed, as a pair of incipient leaves; and these +form the first actual foliage.</p> + +<p>25. Very similar is the germination of the Beech (Fig. <a href="#Fig31">31-33</a>), except +that the caulicle lengthens less, hardly raising the cotyledons out of +the ground. Nothing would be gained by elevating them, as they never +grow out into efficient leaves; but the joint of stem belonging to the +plumule lengthens well, carrying up its pair of real foliage-leaves.</p> + +<p><a name="UNIT_26" id="UNIT_26">26.</a> It is nearly the same in the Bean of the Old World (Vicia Faba, here +called Horse Bean and Windsor Bean): the caulicle lengthens very little, +does not undertake to elevate the heavy seed, which is left below or +<span class="pagenum"><a name="Page_18" id="Page_18">[Pg 18]</a></span>upon the surface of the soil, the flat but thick cotyledons remaining +in it, and supplying food for the growth of the root below and the +plumule above. In its near relative, the Pea (Fig. <a href="#Fig31">34, 35</a>), this use of +cotyledons for storage only is most completely carried out. For they are +thickened to the utmost, even into hemispheres; the caulicle does not +lengthen at all; merely sends out roots from the lower end, and develops +its strong plumule from the upper, the seed remaining unmoved +underground. That is, in technical language, the germination is +<i>hypogæous</i>.</p> + +<div class="figcenter"> +<a name="Fig26" id="Fig26"></a> +<img src="images/fig026_030.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 26. Embryo of Pumpkin-seed, partly opened. 27. Young +seedling of same.</p> +<p><span class="smcap">Fig.</span> 28. Embryo of Common Bean (Phaseolus vulgaris): +caulicle bent down over edge of cotyledons. 29. Same germinating: +caulicle well lengthened and root beginning; thick cotyledons partly +spreading; and plumule (pair of leaves) growing between them. 30. Same, +older, with plumule developed into internode and pair of leaves.</p> +</div> + +<p>27. There is sufficient nourishment in the cotyledons of a pea to make a +very considerable growth before any actual foliage is required. So it is +the stem-portion of the plumule which is at first conspicuous and +strong-growing. Here, as seen in Fig. <a href="#Fig31">35</a>, its lower nodes bear each a +useless leaf-scale instead of an efficient leaf, and only the later ones +bear leaves fitted for foliage.</p> + +<p><span class="pagenum"><a name="Page_19" id="Page_19">[Pg 19]</a></span></p><div class="figcenter"> +<a name="Fig31" id="Fig31"></a> +<img src="images/fig031_035.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 31. A Beech-nut, cut across. 32. Beginning +germination of the Beech, showing the plumule growing before the +cotyledons have opened or the root has scarcely formed. 33. The same, a +little later, with the plumule-leaves developing, and elevated on a long +internode.</p> +<p><span class="smcap">Fig.</span> 34. Embryo of Pea, i. e. a pea with the coats +removed; the short and thick caulicle presented to view. 35. Same in +advanced germination: the plumule has developed four or five internodes, +bearing single leaves; but the first and second leaves are mere scales, +the third begins to serve as foliage; the next more so.</p> +</div> + +<p>28. This <i>hypogæous</i> germination is exemplified on a larger scale by the +Oak (Fig. <a href="#Fig36">36, 37</a>) and Horse-chestnut (Fig. <a href="#Fig36">38, 39</a>); but in these the +downward growth is wholly a stout tap-root. It is not the caulicle; for +this lengthens hardly any. Indeed, the earliest growth which carries the +very short caulicle out of the shell comes from the formation of +foot-stalks to the cotyledons; above these develops the strong plumule, +below grows the stout root. The growth is at first entirely, for a long +time <span class="pagenum"><a name="Page_20" id="Page_20">[Pg 20]</a></span>mainly, at the expense of the great store of food in the +cotyledons. These, after serving their purpose, decay and fall away.</p> + +<div class="figcenter"> +<a name="Fig36" id="Fig36"></a> +<img src="images/fig036_039.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 36. Half of an acorn, cut lengthwise, filled by the +very thick cotyledons, the base of which encloses the minute caulicle. +37. Oak-seedling.</p> +<p><span class="smcap">Fig.</span> 38. Half of a horse-chestnut, similarly cut; the +caulicle is curved down on the side of one of the thick cotyledons. 39. +Horse-chestnut in germination; foot-stalks are formed to the cotyledons, +pushing out in their lengthening the growing parts.</p> +</div> + +<p>29. Such thick cotyledons never separate; indeed, they sometimes grow +together by some part of their contiguous faces; so that the germination +seems to proceed from a solid bulb-like mass. This is the case in a +horse-chestnut.</p> + +<p><a name="UNIT_30" id="UNIT_30">30.</a> <b>Germinating Embryo supplied by its own Store of Nourishment</b>, i. e. +the store in the cotyledons. This is so in all the illustrations thus +far, essentially so even in the Flax. This nourishment was supplied by +the mother plant to the ovule and seed, and thence taken into the embryo +during its growth. Such embryos, filling the whole seed, are +comparatively large and strong, and vigorous in germination in +proportion to the amount of their growth while connected with the parent +plant.</p> + +<p><a name="UNIT_31" id="UNIT_31">31.</a> <b>Germinating Embryo supplied from a Deposit outside of Itself.</b> This +is as common as the other mode; and it occurs in all degrees. <span class="pagenum"><a name="Page_21" id="Page_21">[Pg 21]</a></span>Some +seeds have very little of this deposit, but a comparatively large +embryo, with its parts more or less developed and recognizable. In +others this deposit forms the main bulk of the seed, and the embryo is +small or minute, and comparatively rudimentary. The following +illustrations exemplify these various grades. When an embryo in a seed +is thus surrounded by a white substance, it was natural to liken the +latter to the white of an egg, and the embryo or germ to the yolk. So +the matter around or by the side of the embryo was called the <i>Albumen</i>, +i. e. the white of the seed. The analogy is not very good; and to avoid +ambiguity some botanists call it the <span class="smcap">Endosperm</span>. As that means in English +merely the inwards of a seed, the new name is little better than the old +one; and, since we do not change names in botany except when it cannot +be avoided, this name of <i>albumen</i> is generally kept up. A seed with +such a deposit is <i>albuminous</i>, one with none is <i>exalbuminous</i>.</p> + +<p>32. The <span class="smcap">Albumen</span> forms the main bulk of the seed in wheat, maize, rice, +buckwheat, and the like. It is the floury part of the seed. Also of the +cocoa-nut, of coffee (where it is dense and hard), etc.; while in peas, +beans, almonds, and in most edible nuts, the store of food, although +essentially the same in nature and in use, is in the embryo itself, and +therefore is not counted as anything to be separately named. In both +forms this concentrated food for the germinating plant is food also for +man and for animals.</p> + +<div class="figcenter"> +<a name="Fig40" id="Fig40"></a> +<img src="images/fig040_043.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 40. Seed of Morning Glory divided, moderately +magnified; shows a longitudinal section through the centre of the embryo +as it lies crumpled in the albumen. 41. Embryo taken out whole and +unfolded; the broad and very thin cotyledons notched at summit; the +caulicle below. 42. Early state of germination. 43. Same, more advanced; +caulicle or primary stem, cotyledons or seed-leaves, and below, the +root, well developed.</p> +</div> + +<p>33. For an albuminous seed with a well-developed embryo, the common +Morning Glory (Ipomœa purpurea, Fig. <a href="#Fig40">40-43</a>) is a convenient example, +being easy and prompt to grow, and having all the parts well apparent. +The seeds (duly soaked for examination) and the germination should be +compared with those of Sugar and Red Maple (<a href="#UNIT_19">19-21</a>). The only essential +difference is that here the embryo is surrounded by and crumpled up in +the albumen. This substance, which is pulpy or mucilaginous in fresh and +young seeds, hardens as the seed ripens, but becomes again pulpy in +germination; and, as it liquefies, the thin cotyledons absorb it by +their <span class="pagenum"><a name="Page_22" id="Page_22">[Pg 22]</a></span>whole surface. It supplements the nutritive matter contained in +the embryo. Both together form no large store, but sufficient for +establishing the seedling, with tiny root, stem, and pair of leaves for +initiating its independent growth; which in due time proceeds as in Fig. +<a href="#Fig44">44, 45</a>.</p> + +<div class="figcenter"> +<a name="Fig44" id="Fig44"></a> +<img src="images/fig044_045.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 44. Seedling of Morning Glory more advanced (root +cut away); cotyledons well developed into foliage-leaves: succeeding +internode and leaf well developed, and the next forming. 45. Seedling +more advanced; reduced to much below natural size.</p> +</div> + +<p>34. Smaller embryos, less developed in the seed, are more dependent upon +the extraneous supply of food. The figures <a href="#Fig46">46-53</a> illustrate four grades +in this respect. The smallest, that of the Peony, is still large enough +to be seen with a hand magnifying glass, and even its cotyledons may be +discerned by the aid of a simple stage microscope.</p> + +<p>35. The broad cotyledons of Mirabilis, or Four-o'clock (Fig. <a href="#Fig46">52, 53</a>), +with the slender caulicle almost encircle and enclose the floury +albumen, instead of being enclosed in it, as in the other illustrations. +Evidently here the germinating embryo is principally fed by one of the +leaf-like cotyledons, the other being out of contact with the supply. In +the embryo of Abronia (Fig. <a href="#Fig46">54, 55</a>), a near relative of Mirabilis, there +is a singular modification; one cotyledon is almost wanting, being +reduced to a rudiment, leaving it for the other to do the work. This +leads to the question of the</p> + +<p><a name="UNIT_36" id="UNIT_36">36.</a> <b>Number of Cotyledons.</b> In all the preceding illustrations, the +embryo, however different in shape and degree of development, is +evidently <span class="pagenum"><a name="Page_23" id="Page_23">[Pg 23]</a></span>constructed upon one and the same plan, namely, that of two +leaves on a caulicle or initial stem,—a plan which is obvious even when +one cotyledon becomes very much smaller than the other, as in the rare +instance of Abronia (Fig. <a href="#Fig46">54, 55</a>). In other words, the embryos so far +examined are all</p> + +<p><a name="UNIT_37" id="UNIT_37">37.</a> <b>Dicotyledonous</b>, that is, two-cotyledoned. Plants which are thus +similar in the plan of the embryo agree likewise in the general +structure of their stems, leaves, and blossoms; and thus form a class, +named from their embryo <span class="smcap">Dicotyledones</span>, or in English, <span class="smcap">Dicotyledonous +Plants</span>. So long a name being inconvenient, it may be shortened into +<span class="smcap">Dicotyls</span>.</p> + +<div class="figcenter"> +<a name="Fig46" id="Fig46"></a> +<img src="images/fig046_055.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 46. Section of a seed of a Peony, showing a very +small embryo in the albumen, near one end. 47. This embryo detached, and +more magnified.</p> +<p><span class="smcap">Fig.</span> 48. Section of a seed of Barberry, showing the +straight embryo in the middle of the albumen. 49. Its embryo detached.</p> +<p><span class="smcap">Fig.</span> 50. Section of a Potato seed, showing the embryo +coiled in the albumen. 51. Its embryo detached.</p> +<p><span class="smcap">Fig.</span> 52. Section of the seed of Mirabilis or Four-o'clock, +showing the embryo coiled round the outside of the albumen. 53. +Embryo detached; showing the very broad and leaf-like cotyledons, +applied face to face, and the pair incurved.</p> +<p><span class="smcap">Fig.</span> 54. Embryo of Abronia umbellata; one of the +cotyledons very small. 55. Same straightened out.</p> +</div> + +<p><a name="UNIT_38" id="UNIT_38">38.</a> <b>Polycotyledonous</b> is a name employed for the less usual case in which +there are more than two cotyledons. The Pine is the most familiar case. +This occurs in all Pines, the number of cotyledons varying from three to +twelve; in Fig. <a href="#Fig56">56, 57</a> they are six. Note that they are all on the same +level, that is, belong to the same node, so as to form a circle or +<i>whorl</i> at the summit of the caulicle. When there are only three +cotyledons, they divide the space equally, are one third of the circle +apart. When only two they are 180° apart, that is, are <i>opposite</i>.</p> + +<p>39. The case of three or more cotyledons, which is constant in Pines and +in some of their relatives (but not in all of them), is occasional among +Dicotyls. And the polycotyledonous is only a variation of the +dicotyledonous type,—a difference in the number of leaves in the whorl; +for a pair is a whorl reduced to two members. Some suppose that there +are really only <span class="pagenum"><a name="Page_24" id="Page_24">[Pg 24]</a></span>two cotyledons even in a Pine embryo, but these divided +or split up congenitally so as to imitate a greater number. But as +leaves are often in whorls on ordinary stems, they may be so at the very +beginning.</p> + +<div class="figcenter"> +<a name="Fig56" id="Fig56"></a> +<img src="images/fig056_057.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 56. Section of a Pine-seed, showing its +polycotyledonous embryo in the centre of the albumen, moderately +magnified. 57. Seedling of same, showing the freshly expanded six +cotyledons in a whorl, and the plumule just appearing.</p> +</div> + +<p><a name="UNIT_40" id="UNIT_40">40.</a> <b>Monocotyledonous</b> (meaning with single cotyledon) is the name of the +one-cotyledoned sort of embryo. This goes along with peculiarities in +stem, leaves, and flowers, which all together associate such plants into +a great class, called <span class="smcap">Monocotyledonous Plants</span>, or, for shortness, +<span class="smcap">Monocotyls</span>. It means merely that the leaves are alternate from the very +first.</p> + +<p>41. In Iris (Fig. <a href="#Fig58">58, 59</a>) the embryo in the seed is a small cylinder at +one end of the mass of the albumen, with no apparent distinction of +parts. The end which almost touches the seed coat is caulicle, the other +end belongs to the solitary cotyledon. In germination the whole +lengthens (but mainly the cotyledon) only enough to push the proximate +end fairly out of the seed; from this end the root is formed, and from a +little higher the plumule later emerges. It would appear therefore that +the cotyledon answers to a minute leaf rolled up, and that a chink +through which the plumule grows out is a part of the inrolled edges. The +embryo of Indian Corn shows these parts on a larger scale and in a more +open state (Fig. <a href="#Fig62">66-68</a>). There, in the seed, the cotyledon remains, +imbibing nourishment from the softened albumen, and transmitting it to +the growing root below and new-forming leaves above.</p> + +<div class="figcenter"> +<a name="Fig58" id="Fig58"></a> +<img src="images/fig058_061.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 58. Section of a seed of the Iris, or +Flower-de-Luce, enlarged, showing its small embryo in the albumen, near +the bottom. 59. A germinating seedling of the same, its plumule +developed into the first four leaves (alternate), the first one +rudimentary, the cotyledon remains in the seed.</p> +<p><span class="smcap">Fig.</span> 60. Section of an Onion seed showing the slender and +coiled embryo in the albumen, moderately magnified. 61. Seed of same in +early germination.</p> +</div> + +<div class="figcenter"> +<a name="Fig62" id="Fig62"></a> +<img src="images/fig062_070.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 62. Germinating Onion, more advanced, the chink at +base of cotyledon opening for the protrusion of the plumule, consisting +of a thread-shaped leaf. 63. Section of base of Fig. 62, showing plumule +enclosed. 64. Section of same later, plumule emerging. 65. Later stage +of 62, upper part cut off. 66. A grain of Indian Corn, flatwise, cut +away a little, so as to show the embryo, lying on the albumen which +makes the principal bulk of the seed. 67. A grain cut through the middle +in the opposite direction, dividing the embryo through its thick +cotyledon and its plumule, the latter consisting of two leaves, one +enclosing the other. 68. The embryo taken out whole; the thick mass is +the cotyledon, the narrow body partly enclosed by it is the plumule, the +little projection at its base is the very short radicle enclosed in the +sheathing base of the first leaf of the plumule.</p> +<p><span class="smcap">Fig.</span> 69. Grain of Indian Corn in germination, the +ascending sprout is the first leaf of the plumule, enclosing the younger +leaves within, at its base the primary root has broken through. 70. The +same, advanced; the second and third leaves developing, while the +sheathing first leaf does not further develop.</p> +</div> + +<p>42. The general plan is the same in the Onion (Fig. <a href="#Fig58">60</a>-<a href="#Fig62">65</a>), but with a +striking difference. The embryo is long, and coiled in the albumen of +the seed. To ordinary examination it shows no distinction of parts. But +germination plainly shows that all except the lower end of it is +cotyledon. For after it has lengthened into a long thread, the chink +from which the <span class="pagenum"><a name="Page_25" id="Page_25">[Pg 25]</a></span>plumule in time emerges is seen at the base, or near it, +so the caulicle is extremely short, and does not elongate, but sends out +from its base a simple root, and afterwards others in a cluster. Not +only does the cotyledon lengthen enormously in the seedling, but (unlike +that of Iris, Indian Corn, and all <span class="pagenum"><a name="Page_26" id="Page_26">[Pg 26]</a></span>the cereal grains) it raises the +comparatively light seed into the air, the tip still remaining in the +seed and feeding upon the albumen. When this food is exhausted and the +seedling is well established in the soil, the upper end decays and the +emptied husk of the seed falls away.</p> + +<p>43. In Maize or Indian Corn (Fig. <a href="#Fig62">66-70</a>), the embryo is more developed +in the seed, and its parts can be made out. It lies against the starchy +albumen, but is not enclosed therein. The larger part of it is the +cotyledon, thickish, its edges involute, and its back in contact with +the albumen; partly enclosed by it is the well-developed plumule or bud +which is to grow. For the cotyledon remains in the seed to fulfil its +office of imbibing nourishment from the softened albumen, which it +conveys to the growing sprout; the part of this sprout which is visible +is the first leaf of the plumule rolled up into a sheath and enclosing +the rudiments of the succeeding leaves, at the base enclosing even the +minute caulicle. In germination the first leaf of the plumule develops +only as a sort of sheath, protecting the tender parts within; the second +and the third form the first foliage. The caulicle never lengthens: the +first root, which is formed at its lower end, or from any part of it, +has to break through the enclosing sheath; and succeeding roots soon +spring from all or any of the nodes of the plumule.</p> + +<p><a name="UNIT_44" id="UNIT_44">44.</a> <b>Simple-stemmed Plants</b> are thus built up, by the continuous +production of one leaf-bearing portion of stem from the summit of the +preceding one, beginning with the initial stem (or caulicle) in the +embryo. Some Dicotyls and many Monocotyls develop only in this single +line of growth (as to parts above ground) until the flowering state is +approached. For some examples, see Cycas (Fig. <a href="#Fig71">71</a>, front, at the left); +a tall Yucca or Spanish Bayonet, and two Cocoa-nut Palms behind; at the +right, a group of Sugar-canes, and a Banana behind.</p> + +<div class="figcenter"> +<a name="Fig71" id="Fig71"></a> +<img src="images/fig071.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 71. Simple-stemmed vegetation. +</div> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_27" id="Page_27">[Pg 27]</a></span></p> +<h2><a name="Section_IV" id="Section_IV"></a><span class="smcap">Section IV.</span> GROWTH FROM BUDS: BRANCHING.</h2> + + +<p>45. Most plants increase the amount of their vegetation by branching, +that is, by producing lateral shoots.</p> + +<p>46. Roots branch from any part and usually without definite order. Stems +normally give rise to branches only at definite points, namely, at the +nodes, and there only from the axils of leaves.</p> + +<p><a name="UNIT_47" id="UNIT_47">47.</a> <b>Buds</b> (Fig. <a href="#Fig72">72, 73</a>). Every incipient shoot is a <i>Bud</i> (<a href="#UNIT_12">12</a>). A stem +continues its growth by its <i>terminal bud</i>; it branches by the formation +and development of <i>lateral buds</i>. As normal lateral buds occupy the +axils of leaves, they are called <i>axillary buds</i>. As leaves are +symmetrically arranged on the stem, the buds in their axils and the +branches into which axillary buds grow partake of this symmetry. The +most conspicuous buds are the scaly winter-buds of most shrubs and trees +of temperate and cold climates; but the name belongs as well to the +forming shoot or branch of any herb.</p> + +<div class="figcenter"> +<a name="Fig72" id="Fig72"></a> +<img src="images/fig072_073.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 72. Shoot of Horse-chestnut, of one year's growth, +taken in autumn after the leaves have fallen; showing the large terminal +bud and smaller axillary buds.</p> +<p><span class="smcap">Fig.</span> 73. Similar shoot of Shagbark Hickory, Carya alba.</p> +</div> + +<p><a name="UNIT_48" id="UNIT_48">48.</a> <b>The Terminal Bud</b>, in the most general sense, may be said to exist in +the embryo,—as cotyledons, or the cotyledons and plumule,—and to crown +each successive growth of the simple stem so long as the summit is +capable of growth. The whole ascending growth of the Palm, Cycas, and +the like (such as in Fig. <a href="#Fig71">71</a>) is from a terminal bud. Branches, being +repetitions of the main stem and growing in the same way, are also +lengthened by terminal buds. Those of Horse-chestnut, Hickory, Maples, +and such trees, being the resting buds of winter, are conspicuous by +their protective covering of scales. These bud-scales, as will hereafter +be shown, are themselves a kind of leaves.</p> + +<p><a name="UNIT_49" id="UNIT_49">49.</a> <b>Axillary Buds</b> were formed on these annual shoots early in the +summer. Occasionally they grow the same season into branches; at least, +some of them are pretty sure to do so whenever the growing terminal bud +at the end of the shoot is injured or destroyed. Otherwise they may lie +dormant until the following spring. In many trees or shrubs these +axillary buds do not show themselves until spring; but if searched for, +they may be detected, though of small size, hidden under the bark. +Sometimes, although early <span class="pagenum"><a name="Page_28" id="Page_28">[Pg 28]</a></span>formed, they are concealed all summer long +under the base of the leaf-stalk, which is then hollowed out into a sort +of inverted cup, like a candle-extinguisher, to cover them; as in the +Locust, the Yellow-wood, or more strikingly in the Button-wood or +Plane-tree (Fig. <a href="#Fig74">74</a>).</p> + +<div class="figcenter"> +<a name="Fig74" id="Fig74"></a> +<img src="images/fig074.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 74. An axillary bud, concealed under the hollowed +base of the leaf-stalk, in Buttonwood or Plane-tree.</p> +</div> + +<p>50. The <i>leaf-scars</i>, so conspicuous in Fig. <a href="#Fig72">72, 73</a>, under each axillary +bud, mark the place where the stalk of the subtending leaf was attached +until it fell in autumn.</p> + +<p><a name="UNIT_51" id="UNIT_51">51.</a> <b>Scaly Buds</b>, which are well represented in Fig. <a href="#Fig72">72, 73</a>, commonly +belong to trees and shrubs of countries in which growth is suspended +during winter. The scaly coverings protect the tender young parts +beneath, not so much by keeping out the cold, which of course would +penetrate the bud in time, as by shielding the interior from the effects +of sudden changes. There are all gradations between these and</p> + +<p><a name="UNIT_52" id="UNIT_52">52.</a> <b>Naked Buds</b>, in which these scales are inconspicuous or wanting, as +in most herbs, at least above ground, and most tropical trees and +shrubs. But nearly related plants of the same climate may differ widely +in this respect. Rhododendrons have strong and scaly winter-buds; while +in Kalmia they are naked. One species of Viburnum, the Hobble-bush, has +completely naked buds, what would be a pair of scales developing into +the first leaves in spring; while another (the Snowball) has conspicuous +scaly buds.</p> + +<p><a name="UNIT_53" id="UNIT_53">53.</a> <b>Vigor of Vegetation from strong buds.</b> Large and strong buds, like +those of the Horse-chestnut, Hickory, and the like, contain several +leaves, or pairs of leaves, ready formed, folded and packed away in +small compass, just as the seed-leaves of a strong embryo are packed +away in the seed: they may even contain all the blossoms of the ensuing +season, plainly visible as small buds. And the stems upon which these +buds rest are filled with abundant nourishment, which was deposited the +summer before in the <span class="pagenum"><a name="Page_29" id="Page_29">[Pg 29]</a></span>wood or in the bark. Under the surface of the +soil, or on it covered with the fallen leaves of autumn, similar strong +buds of our perennial herbs may be found; while beneath are thick roots, +rootstocks, or tubers, charged with a great store of nourishment for +their use. This explains how it is that vegetation from such buds shoots +forth so vigorously in the spring of the year, and clothes the bare and +lately frozen surface of the soil, as well as the naked boughs of trees, +very promptly with a covering of fresh green, and often with brilliant +blossoms. Everything was prepared, and even formed, beforehand: the +short joints of stem in the bud have only to lengthen, and to separate +the leaves from each other so that they may unfold and grow. Only a +small part of the vegetation of the season comes directly from the seed, +and none of the earliest vernal vegetation. This is all from buds which +have lived through the winter.</p> + +<p><a name="UNIT_54" id="UNIT_54">54.</a> <b>The Arrangement of Branches</b>, being that of axillary buds, answers to +that of the leaves. Now leaves principally are either <i>opposite</i> or +<i>alternate</i>. Leaves are <i>opposite</i> when there are two from the same +joint of stem, as in Maples (Fig. <a href="#Fig14">20</a>), the two being on opposite sides +of the stem; and so the axillary buds and branches are opposite, as in +Fig. <a href="#Fig75">75</a>. Leaves are <i>alternate</i> when there is only one from each joint +of stem, as in the Oak, Lime-tree, Poplar, Button-wood (Fig. <a href="#Fig74">74</a>), +Morning-Glory (Fig. <a href="#Fig44">45</a>,—not counting the seed-leaves, which of course +are opposite, there being a pair of them); also in Indian Corn (Fig. +<a href="#Fig62">70</a>), and Iris (Fig. <a href="#Fig58">59</a>). Consequently the axillary buds are also +alternate, as in Hickory (Fig. <a href="#Fig72">73</a>); and the branches they form +alternate,—making a different kind of spray from the other mode, one +branch shooting on one side of the stem and the next on some other. For +in the alternate arrangement no leaf is on the same side of the stem as +the one next above or next below it.</p> + +<p>55. But the symmetry of branches (unlike that of the leaves) is rarely +complete. This is due to several causes, and most commonly to the</p> + +<p><a name="UNIT_56" id="UNIT_56">56.</a> <b>Non-development of buds.</b> It never happens that all the buds grow. If +they did, there might be as many branches in any year as there were +leaves the year before. And of those which do begin to grow, a large +portion perish, sooner or later, for want of nourishment, or for want of +light, or because those which first begin to grow have an advantage, +which they are apt to keep, taking to themselves the nourishment of the +stem, and starving the weaker buds. In the Horse-chestnut (Fig. <a href="#Fig72">72</a>), +Hickory (Fig. <a href="#Fig72">73</a>), Magnolia, and most other trees with large scaly buds, +the terminal bud is the strongest, and has the advantage in growth; and +next in strength are the upper axillary buds: while the former continues +the shoot of the last year, some of the latter give rise to branches, +and the rest fail to grow. In the Lilac also (Fig. <a href="#Fig75">75</a>), the uppermost +axillary buds are stronger than the lower; but the terminal bud rarely +appears at all; in its place the uppermost pair of axillary buds grow, +and so each stem branches every year into two,—making a repeatedly +two-forked ramification, as in Fig. <a href="#Fig75">76</a>.</p> + +<p><span class="pagenum"><a name="Page_30" id="Page_30">[Pg 30]</a></span></p><div class="figcenter"> +<a name="Fig75" id="Fig75"></a> +<img src="images/fig075_076.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 75. Shoot of Lilac, with winter buds; the two +uppermost axillary ones strong; the terminal not developed. 76. Forking +ramification of Lilac; reduced in size.</p> +</div> + +<p><a name="UNIT_57" id="UNIT_57">57.</a> <b>Latent Buds.</b> Axillary buds that do not grow at the proper season, +and especially those which make no appearance externally, may long +remain latent, and at length upon a favorable occasion start into +growth, so forming branches apparently out of place as they are out of +time. The new shoots seen springing directly out of large stems may +sometimes originate from such latent buds, which have preserved their +life for years. But commonly these arise from</p> + +<p><a name="UNIT_58" id="UNIT_58">58.</a> <b>Adventitious Buds.</b> These are buds which certain shrubs and trees +produce anywhere on the surface of the wood, especially where it has +been injured. They give rise to the slender twigs which often feather +the sides of great branches of our American Elms. They sometimes form on +the root, which naturally is destitute of buds; they are even found upon +some leaves; and they are sure to appear on the trunks and roots of +Willows, Poplars, and Chestnuts, when these are wounded or mutilated. +Indeed Osier-Willows are <i>pollarded</i>, or cut off, from time to time, by +the cultivator, for the purpose of producing a crop of slender +adventitious twigs, suitable for basket-work. Such branches, being +altogether irregular, of course interfere with the natural symmetry of +the tree. Another cause of irregularity, in certain trees and shrubs, is +the formation of what are called</p> + +<div class="figcenter"> +<a name="Fig77" id="Fig77"></a> +<img src="images/fig077.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 77. Tartarean Honeysuckle, with three accessory buds +in each axil. +</div> + +<p><a name="UNIT_59" id="UNIT_59">59.</a> <b>Accessory or Supernumerary Buds.</b> There are cases where two, three, +or more buds spring from the axil of a leaf, instead of the single one +which is ordinarily found there. Sometimes they are placed one over the +other, as in the Aristolochia or Pipe-Vine, and in the Tartarean +Honeysuckle (Fig. <a href="#Fig77">77</a>); also in the Honey-Locust, and in the Walnut and +Butternut (Fig. <a href="#Fig78">78</a>), where <span class="pagenum"><a name="Page_31" id="Page_31">[Pg 31]</a></span>the upper supernumerary bud is a good way +out of the axil and above the others. And this is here stronger than the +others, and grows into a branch which is considerably out of the axil, +while the lower and smaller ones commonly do not grow at all. In other +cases three buds stand side by side in the axil, as in the Hawthorn, and +the Red Maple (Fig. <a href="#Fig78">79</a>.) If these were all to grow into branches, they +would stifle each other. But some of them are commonly flower-buds: in +the Red Maple, only the middle one is a leaf-bud, and it does not grow +until after those on each side of it have expanded the blossoms they +contain.</p> + +<div class="figcenter"> +<a name="Fig78" id="Fig78"></a> +<img src="images/fig078_079.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 78. Butternut branch, with accessory buds, the +uppermost above the axil.</p> +<p><span class="smcap">Fig.</span> 79. Red-Maple branch, with accessory buds placed +side by side. The annular lines toward the base in this and in Fig. <a href="#Fig72">72</a> +are scars of the bud-scales, and indicate the place of the winter-bud of +the preceding year.</p> +</div> + +<p><a name="UNIT_60" id="UNIT_60">60.</a> <b>Sorts of Buds.</b> It may be useful to enumerate the kinds of buds which +have been described or mentioned. They are</p> + +<p><i>Terminal</i>, when they occupy the summit of (or terminate) a stem,</p> + +<p><i>Lateral</i>, when they are borne on the side of a stem; of which the +regular kind is the</p> + +<p><i>Axillary</i>, situated in the axil of a leaf. These are</p> + +<p><i>Accessory</i> or <i>Supernumerary</i>, when they are in addition to the normal +solitary bud; and these are <i>Collateral</i>, when side by side; +<i>Superposed</i>, when one above another;</p> + +<p><i>Extra-axillary</i>, when they appear above the axil, as some do when +superposed, and as occasionally is the case when single.</p> + +<p><i>Naked buds</i>; those which have no protecting scales.</p> + +<p><i>Scaly buds</i>; those which have protecting scales, which are altered +leaves or bases of leaves.</p> + +<p><i>Leaf-buds</i>, contain or give rise to leaves, and develop into a leafy +shoot.</p> + +<p><i>Flower-buds</i>, contain or consist of blossoms, and no leaves.</p> + +<p><i>Mixed buds</i>, contain both leaves and blossoms.</p> + +<p><a name="UNIT_61" id="UNIT_61">61.</a> <b>Definite annual Growth</b> from winter buds is marked in most of the +shoots from strong buds, such as those of the Horse-chestnut and Hickory +(Fig. <a href="#Fig72">72, 73</a>). Such a bud generally contains, already formed in +miniature, all or a great part of the leaves and joints of stem it is to +produce, makes its whole growth in length in the course of a few weeks, +or sometimes even in a few days, and then forms and ripens its buds for +the next year's similar growth.</p> + +<p><a name="UNIT_62" id="UNIT_62">62.</a> <b>Indefinite annual Growth</b>, on the other hand, is well marked in such +trees or shrubs as the Honey-Locust, Sumac, and in sterile shoots of +<span class="pagenum"><a name="Page_32" id="Page_32">[Pg 32]</a></span>the Rose, Blackberry, and Raspberry. That is, these shoots are apt to +grow all summer long, until stopped by the frosts of autumn or some +other cause. Consequently they form and ripen no terminal bud protected +by scales, and the upper axillary buds are produced so late in the +season that they have no time to mature, nor has their wood time to +solidify and ripen. Such stems therefore commonly die back from the top +in winter, or at least all their upper buds are small and feeble; so the +growth of the succeeding year takes place mainly from the lower axillary +buds, which are more mature.</p> + +<p><a name="UNIT_63" id="UNIT_63">63.</a> <b>Deliquescent and Excurrent Growth.</b> In the former case, and wherever +axillary buds take the lead, there is, of course, no single main stem, +continued year after year in a direct line, but the trunk is soon lost +in the branches. Trees so formed commonly have rounded or spreading +tops. Of such trees with <i>deliquescent</i> stems,—that is, with the trunk +dissolved, as it were, into the successively divided branches,—the +common American Elm (Fig. <a href="#Fig80">80</a>) is a good illustration.</p> + +<div class="figcenter"> +<a name="Fig80" id="Fig80"></a> +<img src="images/fig080.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 80. An American Elm, with Spruce-trees, and on the +left Arbor Vitæ.</p> +</div> + +<p>64. On the other hand, the main stem of Firs and Spruces, unless +destroyed by some injury, is carried on in a direct line throughout the +whole growth of the tree, by the development year after year of a +terminal bud: this forms a single, uninterrupted shaft,—an <i>excurrent</i> +trunk, which cannot be confounded with the branches that proceed from +it. Of such <i>spiry</i> or <i>spire-shaped</i> trees, the Firs or Spruces are +characteristic and familiar examples. There are all gradations between +the two modes.</p> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_33" id="Page_33">[Pg 33]</a></span></p> +<h2><a name="Section_V" id="Section_V"></a><span class="smcap">Section V.</span> ROOTS.</h2> + + +<p>65. It is a property of stems to produce roots. Stems do not spring from +roots in ordinary cases, as is generally thought, but roots from stems. +When perennial herbs arise from the ground, as they do at spring-time, +they rise from subterranean stems.</p> + +<p><a name="UNIT_66" id="UNIT_66">66.</a> <b>The Primary Root</b> is a downward growth from the root-end of the +caulicle, that is, of the initial stem of the embryo (Fig. <a href="#Fig5">5-7</a>, <a href="#Fig81">81</a>). If +it goes on to grow it makes a <i>main</i> or <i>tap-root</i>, as in Fig. <a href="#Fig36">37</a>, etc. +Some plants keep this main root throughout their whole life, and send +off only small side branches; as in the Carrot and Radish: and in +various trees, like the Oak, it takes the lead of the side-branches for +several years, unless accidentally injured, as a strong tap-root. But +commonly the main root divides off very soon, and is lost in the +branches. <i>Multiple primary roots</i> now and then occur, as in the +seedling of Pumpkin (Fig. <a href="#Fig26">27</a>), where a cluster is formed even at the +first, from the root-end of the caulicle.</p> + +<div class="figcenter"> +<a name="Fig81" id="Fig81"></a> +<img src="images/fig081_082.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 81. Seedling Maple, of the natural size; the root +well supplied with root hairs, here large enough to be seen by the naked +eye. 82. Lower end of this root, magnified, the root seen just as +root-hairs are beginning to form a little behind the tip.</p> +</div> + +<p><a name="UNIT_67" id="UNIT_67">67.</a> <b>Secondary Roots</b> are those which arise from other parts of the stem. +Any part of the stem may produce them, but they most readily come from +the nodes. As a general rule they naturally spring, or may be made to +spring, from almost any young stem, when placed in favorable +circumstances,—that is, when placed in the soil, or otherwise supplied +with moisture and screened from the light. For the special tendency of +the root is to avoid the light, seek moisture, and therefore to bury +itself in the soil. <i>Propagation by division</i>, which is so common and so +very important in cultivation, depends upon the proclivity of stems to +strike root. Stems or branches which remain under ground give out roots +as freely as roots themselves give off branches. Stems which creep on +the ground most commonly root at the joints; so will most branches when +bent to the ground, as in propagation by <i>layering</i>; and propagation by +<i>cuttings</i> equally depends upon the tendency of the cut end of a shoot +to produce roots. Thus, a piece of a plant which has stem and leaves, +either developed or in the bud, may be made to produce roots, and so +become an independent plant.</p> + +<p><span class="pagenum"><a name="Page_34" id="Page_34">[Pg 34]</a></span></p><p><a name="UNIT_68" id="UNIT_68">68.</a> <b>Contrast between Stem and Root.</b> Stems are ascending axes; roots are +descending axes. Stems grow by the successive development of internodes +(<a href="#UNIT_13">13</a>), one after another, each leaf-bearing at its summit (or node); so +that it is of the essential nature of a stem to bear leaves. Roots bear +no leaves, are not distinguishable into nodes and internodes, but grow +on continuously from the lower end. They commonly branch freely, but not +from any fixed points nor in definite order.</p> + +<p>69. Although roots generally do not give rise to stems, and therefore do +not propagate the plant, exceptions are not uncommon. For as stems may +produce adventitious buds, so also may roots. The roots of the Sweet +Potato among herbs, and of the Osage Orange among trees freely produce +adventitious buds, developing into leafy shoots; and so these plants are +propagated by <i>root-cuttings</i>. But most growths of subterranean origin +which pass for roots are forms of stems, the common Potato for example.</p> + +<p>70. Roots of ordinary kinds and uses may be roughly classed into +<i>fibrous</i> and <i>fleshy</i>.</p> + +<p><a name="UNIT_71" id="UNIT_71">71.</a> <b>Fibrous Roots</b>, such as those of Indian Corn (Fig. <a href="#Fig62">70</a>), of most +annuals, and of many perennials, serve only for absorption: these are +slender or thread-like. Fine roots of this kind, and the fine branches +which most roots send out are called <span class="smcap">Rootlets</span>.</p> + +<p><a name="UNIT_72" id="UNIT_72">72.</a> The whole surface of a root absorbs moisture from the soil while +fresh and new; and the newer roots and rootlets are, the more freely do +they imbibe. Accordingly, as long as the plant grows above ground, and +expands fresh foliage, from which moisture largely escapes into the air, +so long it continues to extend and multiply its roots in the soil +beneath, renewing and increasing the fresh surface for absorbing +moisture, in proportion to the demand from above. And when growth ceases +above ground, and the leaves die and fall, or no longer act, then the +roots generally stop growing, <span class="pagenum"><a name="Page_35" id="Page_35">[Pg 35]</a></span>and their soft and tender tips harden. +From this period, therefore, until growth begins anew the next spring, +is the best time for transplanting; especially for trees and shrubs.</p> + +<p>73. The absorbing surface of young roots is much increased by the +formation, near their tips, of <span class="smcap">Root-hairs</span> (Fig. <a href="#Fig81">81, 82</a>), which are +delicate tubular outgrowths from the surface, through the delicate walls +of which moisture is promptly imbibed.</p> + +<div class="figcenter"> +<a name="Fig83" id="Fig83"></a> +<img src="images/fig083_085.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 83-85. Forms of tap-root. +</div> + +<p><a name="UNIT_74" id="UNIT_74">74.</a> <b>Fleshy Roots</b> are those in which the root becomes a storehouse of +nourishment. Typical roots of this kind are those of such biennials as +the turnip and carrot; in which the food created in the first season's +vegetation is accumulated, to be expended the next season in a vigorous +growth and a rapid development of flowers, fruit, and seed. By the time +the seed is matured the exhausted root dies, and with it the whole +plant.</p> + +<p>75. Fleshy roots may be single or multiple. The single root of the +commoner biennials is the primary root, or tap-root, which begins to +thicken in the seedling. Names are given to its shapes, such as</p> + +<p><i>Conical</i>, when it thickens most at the crown, or where it joins the +stem, and tapers regularly downwards to a point, as in the Parsnip and +Carrot (Fig. <a href="#Fig83">84</a>);</p> + +<p><i>Turnip-shaped</i> or <i>napiform</i>, when greatly thickened above, but +abruptly becoming slender below; as the Turnip (Fig. <a href="#Fig83">83</a>); and</p> + +<p><span class="pagenum"><a name="Page_36" id="Page_36">[Pg 36]</a></span></p><p><i>Spindle-shaped</i>, or <i>Fusiform</i>, when thickest in the middle and +tapering to both ends; as the common Radish (Fig. <a href="#Fig83">85</a>).</p> + +<p>76. These examples are of primary roots. It will be seen that turnips, +carrots, and the like, are not pure root throughout; for the caulicle, +from the lower end of which the root grew, partakes of the thickening, +perhaps also some joints of stem above: so the bud-bearing and growing +top is stem.</p> + +<div class="figcenter"> +<a name="Fig86" id="Fig86"></a> +<img src="images/fig086_087.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 86. Sweet-Potato plant forming thickened roots. Some +in the middle are just beginning to thicken; one at the left has grown +more; one at the right is still larger.</p> +<p><span class="smcap">Fig.</span> 87. Fascicled fusiform roots of a Dahlia: <i>a</i>, <i>a</i>, +buds on base of stem.</p> +</div> + +<p>77. A fine example of secondary roots (<a href="#UNIT_67">67</a>), some of which remain fibrous +for absorption, while a few thicken and store up food for the next +season's growth, is furnished by the Sweet Potato (Fig. <a href="#Fig86">86</a>). As stated +above, these are used for propagation by cuttings; for any part will +produce adventitious buds and shoots. The Dahlia produces <i>fascicled</i> +(i. e. clustered) fusiform roots of the same kind, at the base of the +stem (Fig. <a href="#Fig86">87</a>): but these, like most roots, do not produce adventitious +buds. The buds by which Dahlias are propagated belong to the surviving +base of the stem above.</p> + +<p><a name="UNIT_78" id="UNIT_78">78.</a> <b>Anomalous Roots</b>, as they may be called, are those which subserve +other uses than absorption, food-storing, and fixing the plant to the +soil.</p> + +<p><i>Aerial Roots</i>, i. e. those that strike from stems in the open air, are +common in moist and warm climates, as in the Mangrove which reaches the +coast of Florida, the Banyan, and, less strikingly, in some herbaceous +plants, such as Sugar Cane, and even in Indian Corn. Such roots reach +the ground at length, or tend to do so.</p> + +<p><i>Aerial Rootlets</i> are abundantly produced by many climbing plants, such +as the Ivy, Poison Ivy, Trumpet Creeper, etc., springing from the side +of stems, which they fasten to trunks of trees, walls, or other +supports. These are used by the plant for climbing.</p> + +<div class="figcenter"> +<a name="Fig88" id="Fig88"></a> +<img src="images/fig088.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 88. Epiphytes of Florida and Georgia, viz., +Epidendrum conopseum, a small Orchid, and Tillandsia usneoides, the +so-called Long Moss or Black Moss, which is no moss, but a flowering +plant, also <i>T. recurvata</i>; on a bough of Live Oak.</p> +</div> + +<p><a name="UNIT_79" id="UNIT_79">79.</a> <b>Epiphytes, or Air-Plants</b> (Fig. <a href="#Fig88">88</a>), are called by the former name +because commonly growing <span class="pagenum"><a name="Page_37" id="Page_37">[Pg 37]</a></span>upon the trunks or limbs of other plants; by +the latter because, having no connection with the soil, they must derive +their sustenance from the air only. They have aerial roots, which do not +reach the ground, but are used to fix the plant to the surface upon +which the plant grows: they also take a part in absorbing moisture from +the air.</p> + +<p><a name="UNIT_80" id="UNIT_80">80.</a> <b>Parasitic Plants</b>, of which there are various kinds, strike their +roots, or what answer to roots, into the tissue of foster plants, or +form attachments with their surface, so as to prey upon their juices. Of +this sort is the Mistletoe, the seed of which germinates on the bough +where it falls or is left by birds; and the forming root penetrates the +bark and engrafts itself into the wood, to which it becomes united as +firmly as a natural branch to its parent stem; and indeed the parasite +lives just as if it were a branch of the tree it grows and feeds on. A +most common parasitic herb is the Dodder; which abounds in low grounds +in summer, and coils its long and slender, leafless, yellowish +stems—resembling tangled threads of yarn—round and round the stalks of +other plants; wherever they touch piercing the bark with minute and very +short rootlets in the form of suckers, which draw out the nourishing +juices of the plants laid hold of. Other parasitic plants, like the +Beech-drops and Pine-sap, fasten their roots under ground upon the roots +of neighboring plants, and rob them of their juices.</p> + +<p>81. Some plants are partly parasitic; while most of their roots act in +the ordinary way, others make suckers at their tips which grow fast to +the roots of other plants and rob them of nourishment. Some of our +species of Gerardia do this (Fig. <a href="#Fig89">89</a>).</p> + +<div class="figcenter"> +<a name="Fig89" id="Fig89"></a> +<img src="images/fig089.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 89. Roots of Yellow Gerardia, some attached to and +feeding on the root of a Blueberry-bush.</p> +</div> + +<p>82. There are phanerogamous plants, like Monotropa or Indian Pipe, the +roots of which feed mainly on decaying vegetable matter in the soil. +These are <span class="smcap">Saprophytes</span>, and they imitate Mushrooms and other Fungi in +their mode of life.</p> + +<p><a name="UNIT_83" id="UNIT_83">83.</a> <b>Duration of Roots, etc.</b> Roots are said to be either <i>annual</i>, +<i>biennial</i>, or <i>perennial</i>. As respects the first and second, these +terms may be applied either to the root or to the plant.</p> + +<p><a name="UNIT_84" id="UNIT_84">84.</a> <b>Annuals</b>, as the name denotes, live for only one year, generally for +<span class="pagenum"><a name="Page_38" id="Page_38">[Pg 38]</a></span>only a part of the year. They are of course herbs; they spring from the +seed, blossom, mature their fruit and seed, and then die, root and all. +Annuals of our temperate climates with severe winters start from the +seed in spring, and perish at or before autumn. Where the winter is a +moist and growing season and the summer is dry, <i>winter annuals</i> +prevail; their seeds germinate under autumn or winter rains, grow more +or less during winter, blossom, fructify, and perish in the following +spring or summer. Annuals are fibrous-rooted.</p> + +<p><a name="UNIT_85" id="UNIT_85">85.</a> <b>Biennials</b>, of which the Turnip, Beet, and Carrot are familiar +examples, grow the first season without blossoming, usually thicken +their roots, laying up in them a stock of nourishment, are quiescent +during the winter, but shoot vigorously, blossom, and seed the next +spring or summer, mainly at the expense of the food stored up, and then +die completely. Annuals and biennials flower only once; hence they have +been called <i>Monocarpic</i> (that is, once-fruiting) plants.</p> + +<p><a name="UNIT_86" id="UNIT_86">86.</a> <b>Perennials</b> live and blossom year after year. A perennial herb, in a +temperate or cooler climate, usually dies down to the ground at the end +of the season's growth. But subterranean portions of stem, charged with +buds, survive to renew the development. Shrubs and trees are of course +perennial; even the stems and branches above ground live on and grow +year after year.</p> + +<p>87. There are all gradations between annuals and biennials, and between +these and perennials, as also between herbs and shrubs; and the +distinction between shrubs and trees is quite arbitrary. There are +perennial herbs and even shrubs of warm climates which are annuals when +raised in a climate which has a winter,—being destroyed by frost. The +Castor-oil plant is an example. There are perennial herbs of which only +small portions survive, as off-shoots, or, in the Potato, as tubers, +etc.</p> + + + +<hr class="chap" /> +<h2><a name="Section_VI" id="Section_VI"></a><span class="smcap">Section VI.</span> STEMS.</h2> + + +<p><a name="UNIT_88" id="UNIT_88">88.</a> <b>The Stem</b> is the axis of the plant, the part which bears all the +other organs. Branches are secondary stems, that is, stems growing out +of stems. The stem at the very beginning produces roots, in most plants +a single root from the base of the embryo-stem, or caulicle. As this +root becomes a <i>descending axis</i>, so the stem, which grows in the +opposite direction is called the <i>ascending axis</i>. Rising out of the +soil, the stem bears leaves; and leaf-bearing is the particular +characteristic of the stem. But there are forms of stems that remain +underground, or make a part of their growth there. These do not bear +leaves, in the common sense; yet they bear rudiments of leaves, or what +answers to leaves, although not in the form of foliage. The so-called +stemless or <i>acaulescent</i> plants are those which bear no obvious stem +(<i>caulis</i>) above ground, but only flower-stalks, and the like.</p> + +<p><span class="pagenum"><a name="Page_39" id="Page_39">[Pg 39]</a></span></p><p><a name="UNIT_89" id="UNIT_89">89.</a> <b>Stems above ground</b>, through differences in duration, texture, and +size, form herbs, shrubs, trees, etc., or in other terms are</p> + +<p><i>Herbaceous</i>, dying down to the ground every year, or after blossoming.</p> + +<p><i>Suffrutescent</i>, slightly woody below, there surviving from year to +year.</p> + +<p><i>Suffruticose</i> or <i>Frutescent</i>, when low stems are decidedly woody +below, but herbaceous above.</p> + +<p><i>Fruticose</i> or <i>Shrubby</i>, woody, living from year to year, and of +considerable size,—not, however, more than three or four times the +height of a man.</p> + +<p><i>Arborescent</i>, when tree-like in appearance or mode of growth, or +approaching a tree in size.</p> + +<p><i>Arboreous</i>, when forming a proper tree-trunk.</p> + +<p>90. As to direction taken in growing, stems may, instead of growing +upright or erect, be</p> + +<p><i>Diffuse</i>, that is, loosely spreading in all directions.</p> + +<p><i>Declined</i>, when turned or bending over to one side.</p> + +<p><i>Decumbent</i>, reclining on the ground, as if too weak to stand.</p> + +<p><i>Assurgent</i> or <i>Ascending</i>, rising obliquely upwards.</p> + +<p><i>Procumbent</i> or <i>Prostrate</i>, lying flat on the ground from the first.</p> + +<p><i>Creeping</i> or <i>Repent</i>, prostrate on or just beneath the ground, and +striking root, as does the White Clover, the Partridge-berry, etc.</p> + +<p><i>Climbing</i> or <i>Scandent</i>, ascending by clinging to other objects for +support, whether by <i>tendrils</i>, as do the Pea, Grape-Vine, and +Passion-flower and Virginia Creeper (Fig. <a href="#Fig92">92</a>, <a href="#Fig93">93</a>); by their twisting +leaf-stalks, as the Virgin's Bower; or by rootlets, like the Ivy, Poison +Ivy, and Trumpet Creeper.</p> + +<p><i>Twining</i> or <i>Voluble</i>, when coiling spirally around other stems or +supports; like the Morning-Glory (Fig. <a href="#Fig90">90</a>) and the Hop.</p> + +<div class="figcenter"> +<a name="Fig90" id="Fig90"></a> +<img src="images/fig090.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 90. Twining or voluble stem of Morning-Glory. +</div> + +<p>91. Certain kinds of stems or branches, appropriated to special uses, +have received distinct substantive names; such as the following:</p> + +<p><a name="UNIT_92" id="UNIT_92">92.</a> <b>A Culm</b>, or straw-stem, such as that of Grasses and Sedges.</p> + +<p><a name="UNIT_93" id="UNIT_93">93.</a> <b>A Caudex</b> is the old name for such a peculiar trunk as a Palm-stem; +it is also used for an upright and thick rootstock.</p> + +<p><a name="UNIT_94" id="UNIT_94">94.</a> <b>A Sucker</b> is a branch rising from stems under ground. Such are +produced abundantly by the Rose, Raspberry, and other plants said to +multiply "by the root." If we uncover them, we see at once the great +difference between these subterranean branches and real roots. They are +only creeping branches under ground. Remarking how the upright shoots +from these branches become separate <span class="pagenum"><a name="Page_40" id="Page_40">[Pg 40]</a></span>plants, simply by the dying off of +the connecting under-ground stems, the gardener expedites the result by +cutting them through with his spade. That is, he propagates the plant +"by division."</p> + +<p><a name="UNIT_95" id="UNIT_95">95.</a> <b>A Stolon</b> is a branch from above ground, which reclines or becomes +prostrate and strikes root (usually from the nodes) wherever it rests on +the soil. Thence it may send up a vigorous shoot, which has roots of its +own, and becomes an independent plant when the connecting part dies, as +it does after a while. The Currant and the Gooseberry naturally multiply +in this way, as well as by suckers (which are the same thing, only the +connecting part is concealed under ground). Stolons must have suggested +the operation of <i>layering</i> by bending down and covering with soil +branches which do not naturally make stolons; and after they have taken +root, as they almost always will, the gardener cuts through the +connecting stem, and so converts a rooting branch into a separate plant.</p> + +<p><a name="UNIT_96" id="UNIT_96">96.</a> <b>An Offset</b> is a short stolon, or sucker, with a crown of leaves at +the end, as in the Houseleek (Fig. <a href="#Fig91">91</a>), which propagates abundantly in +this way.</p> + +<div class="figcenter"> +<a name="Fig91" id="Fig91"></a> +<img src="images/fig091.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 91. Houseleek (Sempervivum), with offsets. +</div> + +<p><a name="UNIT_97" id="UNIT_97">97.</a> <b>A Runner</b>, of which the Strawberry presents the most familiar and +characteristic example, is a long and slender, tendril-like stolon, or +branch from next the ground, destitute of conspicuous leaves. Each +runner of the Strawberry, after having grown to its full length, strikes +root from the tip, which fixes it to the ground, then forms a bud there, +which develops into a tuft of leaves, and so gives rise to a new plant, +which sends out new runners to act in the same way. In this manner a +single Strawberry plant will spread over a large space, or produce a +great number of plants, in the course of the summer, all connected at +first by the slender runners; but these die in the following winter, if +not before, and leave the plants as so many separate individuals.</p> + +<p><a name="UNIT_98" id="UNIT_98">98.</a> <b>Tendrils</b> are branches of a very slender sort, like runners, not +destined like them for propagation, and therefore always destitute of +buds or leaves, being intended only for climbing. Simple tendrils are +such as those of Passion-flowers (Fig. <a href="#Fig92">92</a>). Compound or branching +tendrils are borne by the Cucumber and Pumpkin, by the Grape-Vine, +Virginia Creeper, etc.</p> + +<div class="figcenter"> +<a name="Fig92" id="Fig92"></a> +<img src="images/fig092.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 92. A small Passion-flower (<i>Passiflora sicyoides</i>), +showing the tendrils.</p> +</div> + +<p>99. A tendril commonly grows straight and outstretched until it reaches +some neighboring support, such as a stem, when its apex hooks around it +to secure a hold; then the whole tendril shortens itself by coiling up +spirally, and so draws the shoot of the growing plant nearer to the +supporting object. But the tendrils of the Virginia Creeper (Ampelopsis, +Fig. <span class="pagenum"><a name="Page_41" id="Page_41">[Pg 41]</a></span><a href="#Fig93">93</a>), as also the shorter ones of the Japanese species, effect the +object differently, namely, by expanding the tips of the tendrils into a +flat disk, with an adhesive face. This is applied to the supporting +object, and it adheres firmly; then a shortening of the tendril and its +branches by coiling brings up the growing shoot close to the support. +This is an adaptation for climbing mural rocks or walls, or the trunks +of trees, to which ordinary tendrils are unable to cling. The Ivy and +Poison Ivy attain the same result by means of aerial rootlets (<a href="#UNIT_78">78</a>).</p> + +<div class="figcenter"> +<a name="Fig93" id="Fig93"></a> +<img src="images/fig093_094.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 93. Piece of the stem of Virginia Creeper, bearing a +leaf and a tendril. 94. Tips of a tendril, about the natural size, +showing the disks by which they hold fast to walls, etc.</p> +</div> + +<p>100. Some tendrils are leaves or parts of leaves, as those of the Pea +(Fig. <a href="#Fig31">35</a>). The nature of the tendril is known by its position. A tendril +from the axil of a leaf, like that of Passion-flowers (Fig. <a href="#Fig92">92</a>) is of +course a stem, i. e. a branch. So is one which terminates a stem, as in +the Grape-Vine.</p> + +<p><a name="UNIT_101" id="UNIT_101">101.</a> <b>Spines</b> or <b>Thorns</b> (Fig. <a href="#Fig95">95, 96</a>) are commonly stunted and hardened +branches or tips of stems or branches, as are those of Hawthorn, +Honey-Locust, etc. In the Pear and Sloe all gradations occur between +spines and spine-like (spinescent) branches. Spines may be reduced and +indurated leaves; as in the Barberry, where their nature is revealed by +their situation, underneath an axillary bud. But <span class="pagenum"><a name="Page_42" id="Page_42">[Pg 42]</a></span>prickles, such as +those of Blackberry and Roses, are only excrescences of the bark, and +not branches.</p> + +<div class="figcenter"> +<a name="Fig95" id="Fig95"></a> +<img src="images/fig095_096.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 95. A branching thorn of Honey-Locust, being an +indurated leafless branch developed from an accessory bud far above the +axil: at the cut portion below, three other buds (<i>a</i>) are concealed +under the petiole.</p> +<p><span class="smcap">Fig.</span> 96. Spine of Cockspur Thorn, developed from an +axillary bud, as the leaf-scar below witnesses: an accessory leaf-bud is +seen at its base.</p> +</div> + +<p>102. Equally strange forms of stems are characteristic of the Cactus +family (Fig. <a href="#Fig111">111</a>). These may be better understood by comparison with</p> + +<p><a name="UNIT_103" id="UNIT_103">103.</a> <b>Subterranean Stems and Branches.</b> These are very numerous and +various; but they are commonly overlooked, or else are confounded with +roots. From their situation they are out of ordinary sight; but they +will well repay examination. For the vegetation that is carried on under +ground is hardly less varied or important than that above ground. All +their forms may be referred to four principal kinds: namely, the +<i>Rhizoma</i> (<i>Rhizome</i>) or <i>Rootstock</i>, the <i>Tuber</i>, the <i>Corm</i> or solid +bulb, and the true <i>Bulb</i>.</p> + +<div class="figcenter"> +<a name="Fig97" id="Fig97"></a> +<img src="images/fig097.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 97. Rootstocks, or creeping subterranean branches, +of the Peppermint.</p> +</div> + +<p><a name="UNIT_104" id="UNIT_104">104.</a> <b>The Rootstock, or Rhizoma</b>, in its simplest form, is merely a +creeping stem or branch growing beneath the surface of the soil, or +partly covered by it. Of this kind are the so-called <i>creeping</i>, +<i>running</i>, or <i>scaly roots</i>, such as those by which the Mint (Fig. <a href="#Fig97">97</a>), +the Couch-grass, or Quick-grass, and many other plants, spread so +rapidly and widely,—"by the root," as it is said. That these are really +<i>stems</i>, and not roots, is evident from the way in which <span class="pagenum"><a name="Page_43" id="Page_43">[Pg 43]</a></span>they grow; +from their consisting of a succession of joints; and from the leaves +which they bear on each <i>node</i>, in the form of small scales, just like +the lowest ones on the upright stem next the ground. They also produce +buds in the axils of these scales, showing the scales to be leaves; +whereas real roots bear neither leaves nor axillary buds. Placed as they +are in the damp and dark soil, such stems naturally produce roots, just +as the creeping stem does where it lies on the surface of the ground.</p> + +<p>105. It is easy to see why plants with these running rootstocks take +such rapid and wide possession of the soil, and why they are so hard to +get rid of. They are always perennials; the subterranean shoots live +over the first winter, if not longer, and are provided with vigorous +buds at every joint. Some of these buds grow in spring into upright +stems, bearing foliage, to elaborate nourishment, and at length produce +blossoms for reproduction by seed; while many others, fed by nourishment +supplied from above, form a new generation of subterranean shoots; and +this is repeated over and over in the course of the season or in +succeeding years. Meanwhile, as the subterranean shoots increase in +number, the older ones, connecting the successive growths, die off year +by year, liberating the already rooted side-branches as so many separate +plants; and so on indefinitely. Cutting these running rootstocks into +pieces, therefore, by the hoe or the plough, far from destroying the +plant, only accelerates the propagation; it converts one many-branched +plant into a great number of separate individuals. Cutting into pieces +only multiplies the pest; for each piece (Fig. <a href="#Fig98">98</a>) is already a +plantlet, with its roots and with a bud in the axil of its scale-like +leaf (either latent or apparent), and with prepared nourishment enough +to develop this bud into a leafy stem; and so a single plant is all the +more speedily converted into a multitude. Whereas, when the subterranean +parts are only roots, cutting away the stem completely destroys the +plant, except in the rather rare cases where the root freely produces +adventitious buds.</p> + +<div class="figcenter"> +<a name="Fig98" id="Fig98"></a> +<img src="images/fig098.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 98. A piece of the running rootstock of the +Peppermint, with its node or joint, and an axillary bud ready to grow.</p> +</div> + +<p>106. Rootstocks are more commonly thickened by the storing up of +considerable nourishing matter in their tissue. The common species of +Iris (Fig. <a href="#Fig164">164</a>) in the gardens have stout rootstocks, which are only +partly covered by the soil, and which bear foliage-leaves instead of +mere scales, closely covering the upper part, while the lower produces +roots. As the leaves die, year by year, and decay, a scar left in the +form of a ring marks the place where each leaf was attached, that is, +marks so many nodes, separated by very short internodes.</p> + +<p>107. Some rootstocks are marked with large round scars of a different +<span class="pagenum"><a name="Page_44" id="Page_44">[Pg 44]</a></span>sort, like those of the Solomon's Seal (Fig. <a href="#Fig99">99</a>), which gave this name +to the plant, from their looking somewhat like the impression of a seal +upon wax. Here the rootstock sends up every spring an herbaceous stalk +or stem, which bears the foliage and flowers, and dies in autumn. The +<i>seal</i> is the circular scar left by the death and separation of the base +of the stout stalk from the living rootstock. As but one of these is +formed each year, they mark the limits of a year's growth. The bud at +the end of the rootstock in the figure (which was taken in summer) will +grow the next spring into the stalk of the season, which, dying in +autumn, will leave a similar scar, while another bud will be formed +farther on, crowning the ever-advancing summit or growing end of the +stem.</p> + +<div class="figcenter"> +<a name="Fig99" id="Fig99"></a> +<img src="images/fig099.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 99. Rootstock of Solomon's Seal, with the bottom of +the stalk of the season, and the bud for the next year's growth.</p> +</div> + +<p>108. As each year's growth of stem makes its own roots, it soon becomes +independent of the older parts. And after a certain age, a portion +annually dies off behind, about as fast as it increases at the growing +end, death following life with equal and certain step, with only a +narrow interval. In vigorous plants of Solomon's Seal or Iris, the +living rootstock is several inches or a foot in length; while in the +short rootstock of Trillium or Birthroot (Fig. <a href="#Fig100">100</a>) life is reduced to a +narrower span.</p> + +<div class="figcenter"> +<a name="Fig100" id="Fig100"></a> +<img src="images/fig100.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 100. The very short rootstock and strong terminal +bud of a Trillium or Birthroot.</p> +</div> + +<p>109. An upright or short rootstock, like this of Trillium, is commonly +called a <span class="smcap">Caudex</span> (<a href="#UNIT_93">93</a>); or when more shortened and thickened it would +become a corm.</p> + +<p><a name="UNIT_110" id="UNIT_110">110.</a> <b>A Tuber</b> may be understood to be a portion of a rootstock thickened, +and with buds (eyes) on the sides. Of course, there are all gradations +between a tuber and a rootstock. Helianthus tuberosus, the so-called +Jerusalem Artichoke (Fig. <a href="#Fig101">101</a>), and the common Potato, are typical and +familiar examples of the tuber. The stalks by which the tubers are +attached to the parent stem are at once seen to be different from the +roots, both in appearance and manner of growth. The scales on the tubers +are the rudiments of leaves; the eyes are the buds in their axils. The +Potato-plant <span class="pagenum"><a name="Page_45" id="Page_45">[Pg 45]</a></span>has three forms of branches: 1. Those that bear ordinary +leaves expanded in the air, to digest what they gather from it and what +the roots gather from the soil, and convert it into nourishment. 2. +After a while a second set of branches at the summit of the plant bear +flowers, which form fruit and seed out of a portion of the nourishment +which the leaves have prepared. 3. But a larger part of this +nourishment, while in a liquid state, is carried down the stem, into a +third sort of branches under ground, and accumulated in the form of +starch at their extremities, which become tubers, or depositories of +prepared solid food,—just as in the Turnip, Carrot, and Dahlia (Fig. +<a href="#Fig83">83</a>-<a href="#Fig86">87</a>), it is deposited in the root. The use of the store of food is +obvious enough. In the autumn the whole plant dies, except the seeds (if +it formed them) and the tubers; and the latter are left disconnected in +the ground. Just as that small portion of nourishing matter which is +deposited in the seed feeds the embryo when it germinates, so the much +larger portion deposited in the tuber nourishes its buds, or eyes, when +they likewise grow, the next spring, into new plants. And the great +supply enables them to shoot with a greater vigor at the beginning, and +to produce a greater amount of vegetation than the seedling plant could +do in the same space of time; which vegetation in turn may prepare and +store up, in the course of a few weeks or months, the largest quantity +of solid nourishing material, in a form most available for food. Taking +advantage of this, man has transported the Potato from the cool Andes of +Chili to other cool climates, and makes it yield him a copious supply of +food, especially important in countries where the season is too short, +or the summer's heat too little, for profitably cultivating the +principal grain-plants.</p> + +<div class="figcenter"> +<a name="Fig101" id="Fig101"></a> +<img src="images/fig101_102.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 101. Tubers of Helianthus tuberosus, called +"artichokes."</p> +<p><span class="smcap">Fig.</span> 102. Bulblet-like tubers, such as are occasionally +formed on the stem of a Potato-plant above ground.</p> +</div> + +<p><a name="UNIT_111" id="UNIT_111">111.</a> <b>The Corm or Solid Bulb</b>, like that of Cyclamen (Fig. <a href="#Fig103">103</a>), and of +Indian Turnip (Fig. <a href="#Fig103">104</a>), is a very short and thick fleshy subterranean +stem, often broader than high. It sends off roots from its lower end, or +rather face, leaves and stalks from its upper. The corm of Cyclamen goes +on to enlarge and to produce a succession of flowers and leaves year +after year. <span class="pagenum"><a name="Page_46" id="Page_46">[Pg 46]</a></span>That of Indian Turnip is formed one year and is consumed +the next. Fig. <a href="#Fig103">104</a> represents it in early summer, having below the corm +of last year, from which the roots have fallen. It is partly consumed by +the growth of the stem for the season, and the corm of the year is +forming at base of the stem above the line of roots.</p> + +<div class="figcenter"> +<a name="Fig103" id="Fig103"></a> +<img src="images/fig103_104.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 103. Corm of Cyclamen, much reduced in size: roots +from lower face, leaf-stalks and flower-stalks from the upper.</p> +<p><span class="smcap">Fig.</span> 104. Corm of Indian Turnip (Arisæma).</p> +</div> + +<p>112. The corm of Crocus (Fig. <a href="#Fig105">105, 106</a>), like that of its relative +Gladiolus, is also reproduced annually, the new ones forming upon the +summit and sides of the old. Such a corm is like a tuber in budding from +the sides, i. e. from the axils of leaves; but these leaves, instead of +being small scales, are the sheathing bases of foliage-leaves which +covered the surface. It resembles a true bulb in having these sheaths or +broad scales; but in the corm or solid bulb, this solid part or stem +makes up the principal bulk.</p> + +<div class="figcenter"> +<a name="Fig105" id="Fig105"></a> +<img src="images/fig105_106.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 105. Corm of a Crocus, the investing sheaths or dead +leaf-bases stripped off. The faint cross-lines represent the scars, +where the leaves were attached, i. e. the nodes: the spaces between are +the internodes. The exhausted corm of the previous year is underneath; +forming ones for next year on the summit and sides.</p> +<p><span class="smcap">Fig.</span> 106. Section of the same.</p> +</div> + +<p><a name="UNIT_113" id="UNIT_113">113.</a> <b>The Bulb</b>, strictly so-called, is a stem like a reduced corm as to +its solid part (or plate); while the main body consists of thickened +scales, which are leaves or leaf-bases. These are like bud-scales; so +that in fact a bulb is a bud with fleshy scales on an exceedingly short +stem. Compare a White Lily bulb (Fig. <a href="#Fig107">107</a>) with the strong scaly buds of +the Hickory and Horse-chestnut (Fig. <a href="#Fig72">72 and 73</a>), and the resemblance +will appear. In corms, as in tubers and rootstocks, the store of food +for future growth is deposited in the stem; while in the bulb, the +greater part is deposited in the bases of the leaves, changing them into +thick scales, which closely overlap or enclose one another.</p> + +<p><a name="UNIT_114" id="UNIT_114">114.</a> <b>A Scaly Bulb</b> (like that of the Lily, Fig. <a href="#Fig107">107, 108</a>) is one in which +the scales are thick but comparatively narrow.</p> + +<div class="figcenter"> +<a name="Fig107" id="Fig107"></a> +<img src="images/fig107_108.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 107. Bulb of a wild Lily. 108. The same divided +lengthwise, showing two forming buds of the next generation.</p> +</div> + +<div class="figcenter"> +<a name="Fig109" id="Fig109"></a> +<img src="images/fig109.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 109. A ground leaf of White Lily, its base (cut +across) thickened into a bulb-scale. This plainly shows that bulb-scales +are leaves.</p> +</div> + +<p><a name="UNIT_115" id="UNIT_115">115.</a> <b>A Tunicated or Coated Bulb</b> is one in which the scales enwrap each +other, forming concentric coats or layers, as in Hyacinth and Onion.</p> + +<p><span class="pagenum"><a name="Page_47" id="Page_47">[Pg 47]</a></span></p><p><a name="UNIT_116" id="UNIT_116">116.</a> <b>Bulblets</b> are very small bulbs growing out of larger ones; or small +bulbs produced above ground on some plants, as in the axils of the +leaves of the bulbiferous Lilies of the gardens (Fig. <a href="#Fig110">110</a>), and often in +the flower-clusters of the Leek and Onion. They are plainly buds with +thickened scales. They never grow into branches, but detach themselves +when full grown, fall to the ground, and take root there to form new +plants.</p> + +<div class="figcenter"> +<a name="Fig110" id="Fig110"></a> +<img src="images/fig110.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 110. Bulblets in the axils of leaves of a Tiger +Lily.</p> +</div> + +<p><a name="UNIT_117" id="UNIT_117">117.</a> <b>Consolidated Vegetation.</b> An ordinary herb, shrub, or tree is +evidently constructed on the plan developing an extensive surface. In +fleshy rootstocks, tubers, corms, and bulbs, the more enduring portion +of the plant is concentrated, and reduced for the time of struggle (as +against drought, heat, or cold) to a small amount of exposed surface, +and this mostly sheltered in the soil. There are many similar +consolidated forms which are not subterranean. Thus plants like the +Houseleek (Fig. <a href="#Fig91">91</a>) imitate a bulb. Among Cactuses the columnar species +of Cereus (Fig. <a href="#Fig111">111</a>, <i>b</i>), may be likened to rootstocks. A green rind +serves the purpose of foliage; but the surface is as nothing compared +with an ordinary leafy plant of the same bulk. Compare, for instance, +the largest Cactus known, the Giant Cereus of the Gila River (Fig. <a href="#Fig111">111</a>, +in the background), which rises to the height of fifty or sixty feet, +with a common leafy tree of the same height, such as that in Fig. <a href="#Fig89">89</a>, +and estimate how vastly greater, even without the foliage, the surface +of the latter is than that of the former. Compare, in the <span class="pagenum"><a name="Page_48" id="Page_48">[Pg 48]</a></span>same view, an +Opuntia or Prickly-Pear Cactus, its stem and branches formed of a +succession of thick and flattened joints (Fig. <a href="#Fig111">111</a>, <i>a</i>), which may be +likened to tubers, or an Epiphyllum (<i>d</i>), having short and flat joints, +with an ordinary leafy shrub or herb of equal size. And finally, in +Melon-Cactuses, Echinocactus (<i>c</i>), or other globose forms (which may be +likened to permanent corms), with their globular or bulb-like shapes, we +have plants in the compactest shape; their spherical figure being such +as to expose the least possible amount of substance to the air. These +are adaptations to climates which are very dry, either throughout or for +a part of the year. Similarly, bulbous and corm-bearing plants, and the +like, are examples of a form of vegetation which in the growing season +may expand a large surface to the air and light, while during the period +of rest the living vegetable is reduced to a globe, or solid form of the +least possible surface; and this protected by its outer coats of dead +and dry scales, as well as by its situation under ground. Such are also +adapted to a season of drought. They largely belong to countries which +have a long hot season of little or no rain, when, their stalks and +foliage above and their roots beneath early perishing, the plants rest +securely in their compact bulbs, filled with nourishment and retaining +their moisture with great tenacity, until the rainy season comes round. +Then they shoot forth leaves and flowers with wonderful rapidity, and +what was perhaps a desert of arid sand becomes green with foliage and +gay with blossoms, almost in a day.</p> + +<div class="figcenter"> +<a name="Fig111" id="Fig111"></a> +<img src="images/fig111.jpg" alt="" /><br /> +</div> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_49" id="Page_49">[Pg 49]</a></span></p> +<h2><a name="Section_VII" id="Section_VII"></a><span class="smcap">Section VII.</span> LEAVES.</h2> + + +<p>118. <span class="smcap">Stems</span> bear leaves, at definite points (nodes, <a href="#UNIT_13">13</a>); and these are +produced in a great variety of forms, and subserve various uses. The +commonest kind of leaf, which therefore may be taken as the type or +pattern, is an expanded green body, by means of which the plant exposes +to the air and light the matters which it imbibes, exhales certain +portions, and assimilates the residue into vegetable matter for its +nourishment and growth.</p> + +<p>119. But the fact is already familiar (<a href="#UNIT_10">10-30</a>) that leaves occur under +other forms and serve for other uses,—for the storage of food already +assimilated, as in thickened seed-leaves and bulb-scales; for covering, +as in bud-scales; and still other uses are to be pointed out. Indeed, +sometimes they are of no service to the plant, being reduced to mere +scales or rudiments, such as those on the rootstocks of Peppermint (Fig. +<a href="#Fig97">97</a>) or the tubers of Jerusalem Artichoke (Fig. <a href="#Fig101">101</a>). These may be said +to be of service only to the botanist, in explaining to him the plan +upon which a plant is constructed.</p> + +<p>120. Accordingly, just as a rootstock, or a tuber, or a tendril is a +kind of stem, so a bud-scale, or a bulb-scale, or a cotyledon, or a +petal of a flower, is a kind of leaf. Even in respect to ordinary +leaves, it is natural to use the word either in a wider or in a narrower +sense; as when in one sense we say that a leaf consists of blade and +petiole or leaf-stalk, and in another sense say that a leaf is petioled, +or that the leaf of Hepatica is three-lobed. The connection should make +it plain whether by leaf we mean leaf-blade only, or the blade with any +other parts it may have. And the student will readily understand that by +leaf in its largest or <i>morphological</i> sense, the botanist means the +organ which occupies the place of a leaf, whatever be its form or its +function.</p> + + +<h3><a name="Subsect_VII_1" id="Subsect_VII_1">§ 1.</a> LEAVES AS FOLIAGE.</h3> + +<p>121. This is tautological; for foliage is simply leaves: but it is very +convenient to speak of typical leaves, or those which serve the plant +for assimilation, as foliage-leaves, or ordinary leaves. These may first +be considered.</p> + +<p><a name="UNIT_122" id="UNIT_122">122.</a> <b>The Parts of a Leaf.</b> The ordinary leaf, complete in its parts, +consists of <i>blade</i>, <i>foot-stalk</i>, or <i>petiole</i>, and a pair of +<i>stipules</i>.</p> + +<p><a name="UNIT_123" id="UNIT_123">123.</a> First the <span class="smcap">Blade</span> or <span class="smcap">Lamina</span>, which is the essential part of ordinary +leaves, that is, of such as serve the purpose of foliage. In structure +it consists of a softer part, the <i>green pulp</i>, called <i>parenchyma</i>, +which is traversed and supported by a fibrous frame, the parts of which +are called <i>ribs</i> or <i>veins</i>, on account of a certain likeness in +arrangement to the veins of animals. <span class="pagenum"><a name="Page_50" id="Page_50">[Pg 50]</a></span>The whole surface is covered by a +transparent skin, the <i>Epidermis</i>, not unlike that which covers the +surface of all fresh shoots.</p> + +<p>124. Note that the leaf-blade expands horizontally,—that is, normally +presents its faces one to the sky, the other to the ground, or when the +leaf is erect the upper face looks toward the stem that bears it, the +lower face away from it. Whenever this is not the case there is +something to be explained.</p> + +<p><a name="UNIT_125" id="UNIT_125">125.</a> The framework consists of <i>wood</i>,—a fibrous and tough material +which runs from the stem through the leaf-stalk, when there is one, in +the form of parallel threads or bundles of fibres; and in the blade +these spread out in a horizontal direction, to form the <i>ribs</i> and +<i>veins</i> of the leaf. The stout main branches of the framework are called +the <i>Ribs</i>. When there is only one, as in Fig. <a href="#Fig112">112</a>, <a href="#Fig113">114</a>, or a middle one +decidedly larger than the rest, it is called the <i>Midrib</i>. The smaller +divisions are termed <i>Veins</i>; and their still smaller subdivisions, +<i>Veinlets</i>. The latter subdivide again and again, until they become so +fine that they are invisible to the naked eye. The fibres of which they +are composed are hollow; forming tubes by which the sap is brought into +the leaves and carried to every part.</p> + +<div class="figcenter"> +<a name="Fig112" id="Fig112"></a> +<img src="images/fig112.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 112. Leaf of the Quince: <i>b</i>, blade; <i>p</i>, petiole; +<i>st</i>, stipules. +</div> + +<p><a name="UNIT_126" id="UNIT_126">126.</a> <b>Venation</b> is the name of the mode of veining, that is, of the way in +which the veins are distributed in the blade. This is of two principal +kinds; namely, the <i>parallel-veined</i>, and the <i>netted-veined</i>.</p> + +<p>127. In <i>Netted-veined</i> (also called <i>Reticulated</i>) leaves, the veins +branch off from the main rib or ribs, divide into finer and finer +veinlets, and the branches unite with each other to form meshes of +network. That is, they <i>anastomose</i>, as anatomists say of the veins and +arteries of the body. The Quince-leaf, in Fig. <a href="#Fig112">112</a>, shows this kind of +veining in a leaf with a single rib. The Maple, Basswood, Plane or +Buttonwood (Fig. <a href="#Fig74">74</a>) show it in leaves of several ribs.</p> + +<p>128. In <i>parallel-veined</i> leaves, the whole framework consists of +slender ribs or veins, which run parallel with each other, or nearly so, +from the base to the point of the leaf,—not dividing and subdividing, +nor forming meshes, except by minute cross-veinlets. The leaf of any +grass, or that of the Lily of the Valley (Fig. <a href="#Fig113">113</a>) will furnish a good +illustration. Such parallel veins Linnæus called <i>Nerves</i>, and +parallel-veined leaves are still commonly called <i>nerved</i> leaves, while +those of the other kind are said to be <span class="pagenum"><a name="Page_51" id="Page_51">[Pg 51]</a></span><i>veined</i>,—terms which it is +convenient to use, although these "nerves" and "veins" are all the same +thing, and have no likeness to the <i>nerves</i> and little to the veins of +animals.</p> + +<p>129. <i>Netted-veined</i> leaves belong to plants which have a pair of +seed-leaves or cotyledons, such as the Maple (Fig. <a href="#Fig14">20</a>, <a href="#Fig21">24</a>), Beech (Fig. +<a href="#Fig31">33</a>), and the like; while <i>parallel-veined</i> or <i>nerved</i> leaves belong to +plants with one cotyledon or true seed-leaf; such as the Iris (Fig. <a href="#Fig58">59</a>), +and Indian Corn (Fig. <a href="#Fig62">70</a>). So that a mere glance at the leaves generally +tells what the structure of the embryo is, and refers the plant to one +or the other of these two grand classes,—which is a great convenience. +For when plants differ from each other in some one important respect, +they usually differ correspondingly in other respects also.</p> + +<div class="figcenter"> +<a name="Fig113" id="Fig113"></a> +<img src="images/fig113_114.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 113. A (parallel-veined) leaf of the Lily of the +Valley. 114. One of the Calla Lily.</p> +</div> + +<p>130. Parallel-veined leaves are of two sorts,—one kind, and the +commonest, having the ribs or nerves all running from the base to the +point of the leaf, as in the examples already given; while in another +kind they run from a midrib to the margin, as in the common +Pickerel-weed of our ponds, in the Banana, in Calla (Fig. <a href="#Fig113">114</a>), and many +similar plants of warm climates.</p> + +<p>131. Netted-veined leaves are also of two sorts, as in the examples +already referred to. In one case the veins all rise from a single rib +(the midrib), as in Fig. <a href="#Fig112">112</a>, <a href="#Fig115">116</a>-<a href="#Fig125">127</a>. Such leaves are called +<i>Feather-veined</i> or <i>Penni-veined</i>, i. e. <i>Pinnately-veined</i>; both terms +meaning the same thing, namely, that the veins are arranged on the sides +of the rib like the plume of a feather on each side of the shaft.</p> + +<p><span class="pagenum"><a name="Page_52" id="Page_52">[Pg 52]</a></span></p><p>132. In the other case (as in Fig. <a href="#Fig74">74</a>, <a href="#Fig128">129-132</a>), the veins branch off +from three, five, seven, or nine ribs, which spread from the top of the +leaf-stalk, and run through the blade like the toes of a web-footed +bird. Hence these are said to be <i>Palmately</i> or <i>Digitately</i> veined, or +(since the ribs diverge like rays from a centre) <i>Radiate-veined</i>.</p> + +<p>133. Since the general outline of leaves accords with the framework or +skeleton, it is plain that <i>feather-veined</i> (or <i>penni-veined</i>) leaves +will incline to elongated shapes, or at least to be longer than broad; +while in <i>radiate-veined</i> leaves more rounded forms are to be expected. +A glance at the following figures shows this.</p> + +<div class="figcenter"> +<a name="Fig115" id="Fig115"></a> +<img src="images/fig115_120.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 115-120. A series of shapes of feather-veined +leaves.</p> +</div> + +<p><a name="UNIT_134" id="UNIT_134">134.</a> <b>Forms of Leaves as to General Outline.</b> It is necessary to give +names to the principal shapes, and to define them rather precisely, +since they afford easy marks for distinguishing species. The same terms +are used for all other flattened parts as well, such as petals; so that +they make up a great part of the descriptive language of Botany. It will +be a good exercise for young students to look up leaves answering to +these names and definitions. Beginning with the narrower and proceeding +to the broadest forms, a leaf is said to be</p> + +<p><i>Linear</i> (Fig. <a href="#Fig115">115</a>), when narrow, several times longer than wide, and of +the same breadth throughout.</p> + +<p><i>Lanceolate</i>, or <i>Lance-shaped</i>, when conspicuously longer than wide, +and tapering upwards (Fig. <a href="#Fig115">116</a>), or both upwards and downwards.</p> + +<p><i>Oblong</i> (Fig. <a href="#Fig115">117</a>), when nearly twice or thrice as long as broad.</p> + +<p><i>Elliptical</i> (Fig. <a href="#Fig115">118</a>) is oblong with a flowing outline, the two ends +alike in width.</p> + +<p><i>Oval</i> is the same as broadly elliptical, or elliptical with the breadth +considerably more than half the length.</p> + +<p><i>Ovate</i> (Fig. <a href="#Fig115">119</a>), when the outline is like a section of a hen's egg +lengthwise, the broader end downward.</p> + +<p><i>Orbicular</i>, or <i>Rotund</i> (Fig. <a href="#Fig128">132</a>), circular in outline, or nearly so.</p> + +<div class="figcenter"> +<a name="Fig121" id="Fig121"></a> +<img src="images/fig121_124.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 121, oblanceolate; 122, spatulate; 123, obovate; and +124, wedge-shaped, feather-veined, leaves.</p> +</div> + +<p>135. A leaf which tapers toward the base instead of toward the apex may +be</p> + +<p><i>Oblanceolate</i> (Fig. <a href="#Fig121">121</a>) when of the lance-shaped form, only more +tapering toward the base than in the opposite direction.</p> + +<p><i>Spatulate</i> (Fig. <a href="#Fig121">122</a>) when more rounded above, but tapering thence to a +narrow base, like an old-fashioned spatula.</p> + +<p><span class="pagenum"><a name="Page_53" id="Page_53">[Pg 53]</a></span></p><p><i>Obovate</i> (Fig. <a href="#Fig121">123</a>) or inversely ovate, that is, ovate with the +narrower end down.</p> + +<p><i>Cuneate</i> or <i>Cuneiform</i>, that is, <i>Wedge-shaped</i> (Fig. <a href="#Fig121">124</a>), broad +above and tapering by nearly straight lines to an acute angle at the +base.</p> + +<div class="figcenter"> +<a name="Fig125" id="Fig125"></a> +<img src="images/fig125_127.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 125, sagittate; 126, auriculate; and 127, +halberd-shaped or hastate leaves.</p> +</div> + +<p><a name="UNIT_136" id="UNIT_136">136.</a> <b>As to the Base</b>, its shape characterizes several forms, such as</p> + +<p><i>Cordate</i> or <i>Heart-shaped</i> (Fig. <a href="#Fig115">120</a>, <a href="#Fig128">129</a>), when a leaf of an ovate +form, or something like it, has the outline of its rounded base turned +in (forming a notch or <i>sinus</i>) where the stalk is attached.</p> + +<p><i>Reniform</i>, or <i>Kidney-shaped</i> (Fig. <a href="#Fig128">131</a>), like the last, only rounder +and broader than long.</p> + +<p><i>Auriculate</i>, or <i>Eared</i>, having a pair of small and blunt projections, +or <i>ears</i>, at the base, as in one species of Magnolia (Fig. <a href="#Fig125">126</a>).</p> + +<p><i>Sagittate</i>, or <i>arrow-shaped</i>, where such ears are acute and turned +downwards, while the main body of the blade tapers upwards to a point, +as in the common Sagittaria or Arrow-head, and in the Arrow-leaved +Polygonum (Fig. <a href="#Fig125">125</a>).</p> + +<p><i>Hastate</i>, or <i>Halberd-shaped</i>, when such lobes at the base point +outwards, giving the shape of the halberd of the olden time, as in +another Polygonum (Fig. <a href="#Fig125">127</a>).</p> + +<div class="figcenter"> +<a name="Fig128" id="Fig128"></a> +<img src="images/fig128_132.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 128-132. Various forms of radiate-veined leaves. +</div> + +<p><i>Peltate</i>, or <i>Shield-shaped</i> (Fig. <a href="#Fig128">132</a>), is the name applied to a +curious modification of the leaf, commonly of a rounded form, where the +footstalk is attached to the lower surface, instead of the base, and +therefore is naturally <span class="pagenum"><a name="Page_54" id="Page_54">[Pg 54]</a></span>likened to a shield borne by the outstretched +arm. The common Watershield, the Nelumbium, and the White Water-lily, +and also the Mandrake, exhibit this sort of leaf. On comparing the +shield-shaped leaf of the common Marsh Pennywort (Fig. <a href="#Fig128">132</a>) with that of +another common species (Fig. <a href="#Fig128">130</a>), it is at once seen that a +shield-shaped leaf is like a kidney-shaped (Fig. <a href="#Fig128">130, 131</a>) or other +rounded leaf, with the margins at the base brought together and united.</p> + +<p><a name="UNIT_137" id="UNIT_137">137.</a> <b>As to the Apex</b>, the following terms express the principal +variations:—</p> + +<p><i>Acuminate</i>, <i>Pointed</i>, or <i>Taper-pointed</i>, when the summit is more or +less prolonged into a narrowed or tapering point; as in Fig. <a href="#Fig133">133</a>.</p> + +<p><i>Acute</i>, ending in an acute angle or not prolonged point; Fig. <a href="#Fig133">134</a>.</p> + +<p><i>Obtuse</i>, with a blunt or rounded apex; as in Fig. <a href="#Fig133">135</a>, etc.</p> + +<p><i>Truncate</i>, with the end as if cut off square; as in Fig. <a href="#Fig133">136</a>.</p> + +<p><i>Retuse</i>, with rounded summit slightly indented, forming a very shallow +notch, as in Fig. <a href="#Fig133">137</a>.</p> + +<p><i>Emarginate</i>, or <i>Notched</i>, indented at the end more decidedly; as in +Fig. <a href="#Fig133">138</a>.</p> + +<p><i>Obcordate</i>, that is, inversely heart-shaped, where an obovate leaf is +more deeply notched at the end (Fig. <a href="#Fig133">139</a>), as in White Clover and +Wood-sorrel; so as to resemble a cordate leaf inverted.</p> + +<p><i>Cuspidate</i>, tipped with a sharp and rigid point; as in Fig. <a href="#Fig133">140</a>.</p> + +<p><i>Mucronate</i>, abruptly tipped with a small and short point, like a mere +projection of the midrib; as in Fig. <a href="#Fig133">141</a>.</p> + +<p><i>Aristate</i>, <i>Awn-pointed</i>, and <i>Bristle-pointed</i>, are terms used when +this mucronate point is extended into a longer bristle-form or slender +appendage.</p> + +<p>The first six of these terms can be applied to the lower as well as to +the upper end of a leaf or other organ. The others belong to the apex +only.</p> + +<div class="figcenter"> +<a name="Fig133" id="Fig133"></a> +<img src="images/fig133_141.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 133-141. Forms of the apex of leaves. +</div> + +<p><a name="UNIT_138" id="UNIT_138">138.</a> <b>As to degree and nature of Division</b>, there is first of all the +difference between</p> + +<p><i>Simple Leaves</i>, those in which the blade is of one piece, however much +it may be cut up, and</p> + +<p><i>Compound Leaves</i>, those in which the blade consists of two or more +separate pieces, upon a common leaf-stalk or support. Yet between these +two kinds every intermediate gradation is to be met with.</p> + +<div class="figcenter"> +<a name="Fig142" id="Fig142"></a> +<img src="images/fig142_147.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 142-147. Kinds of margin of leaves. +</div> + +<p><a name="UNIT_139" id="UNIT_139">139.</a> <b>As to Particular Outlines of Simple Leaves</b> (and the same applies to +their separate parts), they are</p> + +<p><span class="pagenum"><a name="Page_55" id="Page_55">[Pg 55]</a></span></p><p><i>Entire</i>, when their general outline is completely filled out, so that +the margin is an even line, without teeth or notches.</p> + +<p><i>Serrate</i>, or <i>Saw-toothed</i>, when the margin only is cut into sharp +teeth, like those of a saw, and pointing forwards; as in Fig. <a href="#Fig142">142</a>.</p> + +<p><i>Dentate</i>, or <i>Toothed</i>, when such teeth point outwards, instead of +forwards; as in Fig. <a href="#Fig142">143</a>.</p> + +<p><i>Crenate</i>, or <i>Scalloped</i>, when the teeth are broad and rounded; as in +Fig. <a href="#Fig142">144</a>.</p> + +<p><i>Repand</i>, <i>Undulate</i>, or <i>Wavy</i>, when the margin of the leaf forms a +wavy line, bending slightly inwards and outwards in succession; as in +Fig. <a href="#Fig142">145</a>.</p> + +<p><i>Sinuate</i>, when the margin is more strongly sinuous or turned inwards +and outwards; as in Fig. <a href="#Fig142">146</a>.</p> + +<p><i>Incised</i>, <i>Cut</i>, or <i>Jagged</i>, when the margin is cut into sharp, deep, +and irregular teeth or incisions; as in Fig. <a href="#Fig142">147</a>.</p> + +<p><i>Lobed</i>, when deeply cut. Then the pieces are in a general way called +<span class="smcap">Lobes</span>. The number of the lobes is briefly expressed by the phrase +<i>two-lobed</i>, <i>three-lobed</i>, <i>five-lobed</i>, <i>many-lobed</i>, etc., as the +case may be.</p> + +<p>140. When the depth and character of the lobing needs to be more +particularly specified, the following terms are employed, viz.:—</p> + +<p><i>Lobed</i>, in a special sense, when the incisions do not extend deeper +than about half-way between the margin and the centre of the blade, if +so far, and are more or less rounded; as in the leaves of the Post-Oak, +Fig. <a href="#Fig148">148</a>, and the Hepatica, Fig. <a href="#Fig148">152</a>.</p> + +<p><i>Cleft</i>, when the incisions extend half way down or more, and especially +when they are sharp; as in Fig. <a href="#Fig148">149, 153</a>. And the phrases <i>two-cleft</i>, +or, in the Latin form, <i>bifid</i>, <i>three-cleft</i> or <i>trifid</i>, <i>four-cleft</i> +or <i>quadrifid</i>, <i>five-cleft</i> or <i>quinquefid</i>, etc., or <i>many-cleft</i>, in +the Latin form, <i>multifid</i>,—express the number of the <i>Segments</i>, or +portions.</p> + +<p><i>Parted</i>, when the incisions are still deeper, but yet do not quite +reach to the midrib or the base of the blade; as in Fig. <a href="#Fig148">150, 154</a>. And +the terms <i>two-parted</i>, <i>three-parted</i>, etc., express the number of such +divisions.</p> + +<p><i>Divided</i>, when the incisions extend quite to the midrib, as in the +lower part of Fig. <a href="#Fig148">151</a>, or to the leaf-stalk, as in Fig. <a href="#Fig148">155</a>; which +really makes the <span class="pagenum"><a name="Page_56" id="Page_56">[Pg 56]</a></span>leaf compound. Here, using the Latin form, the leaf is +said to be <i>bisected</i>, <i>trisected</i> (Fig. <a href="#Fig148">155</a>), etc., according to the +number of the divisions.</p> + +<div class="figcenter"> +<a name="Fig148" id="Fig148"></a> +<img src="images/fig148_155.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 148, pinnately lobed; 149, pinnately cleft; 150, +pinnately parted; 151, pinnately divided, leaves.</p> +<p><span class="smcap">Fig.</span> 152, palmately three-lobed; 153, palmately +three-cleft; 154, palmately three-parted; 155, palmately three-divided +or trisected, leaves.</p> +</div> + +<p><a name="UNIT_141" id="UNIT_141">141.</a> <b>The Mode of Lobing or Division</b> corresponds to that of the veining, +whether <i>pinnately veined</i> or <i>palmately veined</i>. In the former the +notches or incisions, or <i>sinuses</i>, coming between the principal veins +or ribs are directed toward the midrib: in the latter they are directed +toward the apex of the petiole; as the figures show.</p> + +<p>142. So degree and mode of division may be tersely expressed in brief +phrases. Thus, in the four upper figures of pinnately veined leaves, the +first is said to be <i>pinnately lobed</i> (in the special sense), the second +<i>pinnately cleft</i> (or <i>pinnatifid</i> in Latin form), the third <i>pinnately +parted</i>, the fourth <i>pinnately divided</i>, or <i>pinnatisected</i>.</p> + +<p>143. Correspondingly in the lower row, of palmately veined leaves, the +first is <i>palmately lobed</i>, the second <i>palmately cleft</i>, the third +<i>palmately parted</i>, the fourth <i>palmately divided</i>. Or, in other +language of the same meaning (but now less commonly employed), they are +said to be <i>digitately lobed</i>, <i>cleft</i>, <i>parted</i>, or <i>divided</i>.</p> + +<p>144. The number of the divisions or lobes may come into the phrase. Thus +in the four last named figures the leaves are respectively <i>palmately +three-lobed</i>, <i>three-cleft</i> (or <i>trifid</i>), <i>three-parted</i>, +<i>three-divided</i>, or better (in Latin form), <i>trisected</i>. And so for +higher numbers, as <i>five-lobed</i>, <i>five-cleft</i>, <span class="pagenum"><a name="Page_57" id="Page_57">[Pg 57]</a></span>etc., up to +<i>many-lobed</i>, <i>many-cleft</i> or <i>multifid</i>, etc. The same mode of +expression may be used for pinnately lobed leaves, as <i>pinnately +7-lobed</i>, <i>-cleft</i>, <i>-parted</i>, etc.</p> + +<p>145. The divisions, lobes, etc., may themselves be <i>entire</i> (without +teeth or notches), or <i>serrate</i>, or otherwise toothed or incised; or +lobed, cleft, parted, etc.: in the latter cases making <i>twice +pinnatifid</i>, <i>twice palmately</i> or <i>pinnately lobed</i>, <i>parted</i>, or +<i>divided</i> leaves, etc. From these illustrations one will perceive how +the botanist, in two or three words, may describe any one of the almost +endlessly diversified shapes of leaves, so as to give a clear and +definite idea of it.</p> + +<p><a name="UNIT_146" id="UNIT_146">146.</a> <b>Compound Leaves.</b> A compound leaf is one which has its blade in +entirely separate parts, each usually with a stalklet of its own; and +the stalklet is often <i>jointed</i> (or <i>articulated</i>) with the main +leaf-stalk, just as this is jointed with the stem. When this is the +case, there is no doubt that the leaf is compound. But when the pieces +have no stalklets, and are not jointed with the main leaf-stalk, it may +be considered either as a divided simple leaf, or a compound leaf, +according to the circumstances. This is a matter of names where all +intermediate forms may be expected.</p> + +<p>147. While the pieces or projecting parts of a simple leaf-blade are +called <i>Lobes</i>, or in deeply cut leaves, etc., <i>Segments</i>, or +<i>Divisions</i>, the separate pieces or blades of a compound leaf are called +<span class="smcap">Leaflets</span>.</p> + +<p>148. Compound leaves are of two principal kinds, namely, the <i>Pinnate</i> +and the <i>Palmate</i>; answering to the two modes of veining in reticulated +leaves, and to the two sorts of lobed or divided leaves (<a href="#UNIT_141">141</a>).</p> + +<div class="figcenter"> +<a name="Fig156" id="Fig156"></a> +<img src="images/fig156_158.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 156-158. Pinnate leaves, the first with an odd +leaflet (<i>odd-pinnate</i>); the second with a tendril in place of uppermost +leaflets; the third <i>abruptly pinnate</i>, or of even pairs.</p> +</div> + +<p>149. <i>Pinnate</i> leaves are those in which the leaflets are arranged on +the sides of a main leaf-stalk; as in Fig. <a href="#Fig156">156-158</a>. They answer to the +<span class="pagenum"><a name="Page_58" id="Page_58">[Pg 58]</a></span><i>feather-veined</i> (i. e. <i>pinnately-veined</i>) simple leaf; as will be +seen at once on comparing the forms. The <i>leaflets</i> of the former answer +to the <i>lobes</i> or <i>divisions</i> of the latter; and the continuation of the +petiole, along which the leaflets are arranged, answers to the midrib of +the simple leaf.</p> + +<p>150. Three sorts of pinnate leaves are here given. Fig. <a href="#Fig156">156</a> is <i>pinnate +with an odd</i> or <i>end leaflet</i>, as in the Common Locust and the Ash. Fig. +<a href="#Fig156">157</a> is <i>pinnate with a tendril at the end</i>, in place of the odd leaflet, +as in the Vetches and the Pea. Fig. <a href="#Fig156">158</a> is evenly or <i>abruptly pinnate</i>, +as in the Honey-Locust.</p> + +<div class="figcenter"> +<a name="Fig159" id="Fig159"></a> +<img src="images/fig159.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 159. Palmate (or digitate) leaf of five leaflets, of +the Sweet Buckeye.</p> +</div> + +<p>151. <i>Palmate</i> (also named <i>Digitate</i>) leaves are those in which the +leaflets are all borne on the tip of the leaf-stalk, as in the Lupine, +the Common Clover, the Virginia Creeper (Fig. <a href="#Fig93">93</a>), and the +Horse-chestnut and Buckeye (Fig. <a href="#Fig159">159</a>). They evidently answer to the +<i>radiate-veined</i> or <i>palmately-veined</i> simple leaf. That is, the +Clover-leaf of three leaflets is the same as a palmately three-ribbed +leaf cut into three separate leaflets. And such a simple five-lobed leaf +as that of the Sugar Maple, if more cut, so as to separate the parts, +would produce a palmate leaf of five leaflets, like that of the +Horse-chestnut or Buckeye.</p> + +<p>152. Either sort of compound leaf may have any number of leaflets; yet +palmate leaves cannot well have a great many, since they are all crowded +together on the end of the main leaf-stalk. Some Lupines have nine or +eleven; the Horse-chestnut has seven, the Sweet Buckeye more commonly +five, the Clover three. A pinnate leaf often has only seven or five +leaflets, or only three, as in Beans of the genus Phaseolus, etc.; in +some rarer cases only two; in the Orange and Lemon and also in the +common Barberry there is only one! The joint at the place where the +leaflet is united with the petiole distinguishes this last case from a +simple leaf. In other species of these genera the lateral leaflets also +are present.</p> + +<p>153. The leaflets of a compound leaf may be either <i>entire</i> (as in Fig. +<a href="#Fig125">126</a>-<a href="#Fig128">128</a>), or <i>serrate</i>, or lobed, cleft, parted, etc.; in fact, may +present all the variations of simple leaves, and the same terms equally +apply to them.</p> + +<p>154. When the division is carried so far as to separate what would be +one leaflet into two, three, or several, the leaf becomes <i>doubly</i> or +<i>twice compound</i>, either <i>pinnately</i> or <i>palmately</i>, as the case may be. +For example, while the clustered leaves of the Honey-Locust are <i>simply +pinnate</i>, that is, <i>once pinnate</i>, those on new shoots are <i>bipinnate</i>, +or <i>twice pinnate</i>, as in Fig. <a href="#Fig160">160</a>. When these leaflets are again +divided in the same way, the leaf <span class="pagenum"><a name="Page_59" id="Page_59">[Pg 59]</a></span>becomes <i>thrice pinnate</i>, or +<i>tripinnate</i>, as in many Acacias. The first divisions are called +<i>Pinnæ</i>; the others, <i>Pinnules</i>; and the last, or little blades +themselves, <i>Leaflets</i>.</p> + +<div class="figcenter"> +<a name="Fig160" id="Fig160"></a> +<img src="images/fig160.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 160. A twice-pinnate (abruptly) leaf of the +Honey-Locust. +</div> + +<p>155. So the palmate leaf, if again compounded in the same way, becomes +<i>twice palmate</i>, or, as we say when the divisions are in threes, <i>twice +ternate</i> (in Latin form <i>biternate</i>); if a third time compounded, +<i>thrice ternate</i> or <i>triternate</i>. But if the division goes still +further, or if the degree is variable, we simply say that the leaf is +<i>decompound</i>; either palmately or pinnately decompound, as the case may +be. Thus, Fig. <a href="#Fig161">161</a> represents a four times ternately compound (in other +words a <i>ternately decompound</i>) leaf of a common Meadow Rue.</p> + +<div class="figcenter"> +<a name="Fig161" id="Fig161"></a> +<img src="images/fig161.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 161. Ternately decompound leaf of Meadow Rue. +</div> + +<p>156. When the botanist, in describing leaves, wishes to express the +number of the leaflets, he may use terms like these:—</p> + +<p><i>Unifoliolate</i>, for a compound leaf of a single leaflet; from the Latin +<i>unum</i>, one, and <i>foliolum</i>, leaflet.</p> + +<p><i>Bifoliolate</i>, of two leaflets, from the Latin <i>bis</i>, twice, and +<i>foliolum</i>, leaflet.</p> + +<p><i>Trifoliolate</i> (or <i>ternate</i>), of three leaflets, as the Clover; and so +on.</p> + +<p><i>Palmately bifoliolate</i>, <i>trifoliolate</i>, <i>quadrifoliolate</i>, +<i>plurifoliolate</i> (of several leaflets), etc.: or else</p> + +<p><i>Pinnately bi-</i>, <i>tri-</i>, <i>quadri-</i>, or <i>plurifoliolate</i> (that is, of +two, three, four, five, or several leaflets), as the case may be: these +are terse ways of denoting in single phrases both the number of leaflets +and the kind of compounding.</p> + +<p>157. Of foliage-leaves having certain peculiarities in structure, the +following may be noted:—</p> + +<p><span class="pagenum"><a name="Page_60" id="Page_60">[Pg 60]</a></span></p><p><a name="UNIT_158" id="UNIT_158">158.</a> <b>Perfoliate Leaves.</b> In these the stem that bears them seems to run +through the blade of the leaf, more or less above its base. A common +Bellwort (Uvularia perfoliata, Fig. <a href="#Fig162">162</a>) is a familiar illustration. The +lower and earlier leaves show it distinctly. Later, the plant is apt to +produce some leaves merely clasping the stem by the sessile and +heart-shaped base, and the latest may be merely sessile. So the series +explains the peculiarity: in the formation of the leaf the bases, +meeting around the stem, grow together there.</p> + +<div class="figcenter"> +<a name="Fig162" id="Fig162"></a> +<img src="images/fig162_163.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 162. A summer branch of Uvularia perfoliata; lower +leaves perfoliate, upper cordate-clasping, uppermost simply sessile.</p> +<p><span class="smcap">Fig.</span> 163. Branch of a Honeysuckle, with +connate-perfoliate leaves.</p> +</div> + +<p><a name="UNIT_159" id="UNIT_159">159.</a> <b>Connate-perfoliate.</b> Such are the upper leaves of true Honeysuckles. +Here (Fig. <a href="#Fig162">163</a>) of the opposite and sessile leaves, some pairs, +especially the uppermost, in the course of their formation unite around +the stem, which thus seems to run through the disk formed by their +union.</p> + +<div class="figcenter"> +<a name="Fig164" id="Fig164"></a> +<img src="images/fig164_165.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 164. Rootstock and equitant leaves of Iris. 165. A +section across the cluster of leaves at the bottom, showing the +equitation.</p> +</div> + +<p><a name="UNIT_160" id="UNIT_160">160.</a> <b>Equitant Leaves.</b> While ordinary leaves spread horizontally, and +present one face to the sky and the other to the earth, there are some +that present their tip to the sky, and their faces right and left to the +horizon. Among these are the <i>equitant</i> leaves of the Iris or +Flower-de-Luce. Inspection shows that each leaf was formed as if <i>folded +together lengthwise</i>, <span class="pagenum"><a name="Page_61" id="Page_61">[Pg 61]</a></span>so that what would be the upper surface is +within, and all grown together, except next the bottom, where each leaf +covers the next younger one. It was from their straddling over each +other, like a man on horseback (as is seen in the cross-section, Fig. +<a href="#Fig164">165</a>), that Linnæus, with his lively fancy, called these <i>Equitant</i> +leaves.</p> + +<p><a name="UNIT_161" id="UNIT_161">161.</a> <b>Leaves with no distinction of Petiole and Blade.</b> The leaves of Iris +just mentioned show one form of this. The flat but narrow leaves of +Jonquils, Daffodils, and the cylindrical leaf of Onions are other +instances. <i>Needle-shaped</i> leaves, like those of the Pine, Larch, and +Spruce, and the <i>awl-shaped</i> as well as the <i>scale-shaped</i> leaves of +Junipers, Red Cedar, and Arbor-Vitæ (Fig. <a href="#Fig166">166</a>), are examples.</p> + +<div class="figcenter"> +<a name="Fig166" id="Fig166"></a> +<img src="images/fig166.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 166. Branch of Arbor-Vitæ, with awl-shaped and +scale-shaped leaves.</p> +</div> + +<p><a name="UNIT_162" id="UNIT_162">162.</a> <b>Phyllodia.</b> Sometimes an expanded <i>petiole</i> takes the place of the +blade; as in numerous New Holland Acacias, some of which are now common +in greenhouses. Such counterfeit blades are called <i>phyllodia</i>,—meaning +leaf-like bodies. They may be known from true blades by their standing +edgewise, their margins being directed upwards and downwards; while in +true blades the faces look upwards and downwards; excepting in equitant +leaves, as already explained.</p> + +<p><a name="UNIT_163" id="UNIT_163">163.</a> <b>Falsely Vertical Leaves.</b> These are apparent exceptions to the rule, +the blade standing edgewise instead of flatwise to the stem; but this +position comes by a twist of the stalk or the base of the blade. Such +leaves present the two faces about equally to the light. The +Compass-plant (Silphium laciniatum) is an example. So also the leaves of +Boltonia, of Wild Lettuce, and of a vast number of Australian Myrtaceous +shrubs and trees, which much resemble the phyllodia of the Acacias of +the same country. They are familiar in Callistemon, the Bottle-brush +Flower, and in Eucalyptus. But in the latter the leaves of the young +tree have the normal structure and position.</p> + +<div class="figcenter"> +<a name="Fig167" id="Fig167"></a> +<img src="images/fig167_168.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 167. The ambiguous leaf? (cladophyllum) of +Myrsiphyllum.</p> +<p><span class="smcap">Fig.</span> 168. Same of Ruscus, or Butcher's Broom.</p> +</div> + +<p><a name="UNIT_164" id="UNIT_164">164.</a> <b>Cladophylla</b>, meaning <i>branch-leaves</i>. The foliage of Ruscus (the +Butcher's Broom of Europe) and of Myrsiphyllum of South Africa +(cultivated for decoration under the false <span class="pagenum"><a name="Page_62" id="Page_62">[Pg 62]</a></span>name of Smilax) is peculiar +and puzzling. If these blades (Fig. <a href="#Fig167">167, 168</a>) are really leaves, they +are most anomalous in occupying the axil of another leaf, reduced to a +little scale. Yet they have an upper and lower face, as leaves should, +although they soon twist, so as to stand more or less edgewise. If they +are branches which have assumed exactly the form and office of leaves, +they are equally extraordinary in not making any further development. +But in Ruscus, flowers are borne on one face, in the axil of a little +scale: and this would seem to settle that they are branches. In +Asparagus just the same things as to position are thread-shaped and +branch-like.</p> + + +<h3><a name="Subsect_VII_2" id="Subsect_VII_2">§ 2.</a> LEAVES OF SPECIAL CONFORMATION AND USE.</h3> + +<div class="figcenter"> +<a name="Fig169" id="Fig169"></a> +<img src="images/fig169.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 169. A young Agave Americana, or Century-plant; +fleshy-leaved.</p> +</div> + +<p><a name="UNIT_165" id="UNIT_165">165.</a> <b>Leaves for Storage.</b> A leaf may at the same time serve both ordinary +and special uses. Thus in those leaves of Lilies, such as the common +White Lily, which spring from the bulb, the upper and green part serves +for foliage and elaborates nourishment, while the thickened portion or +bud-scale beneath serves for the storage of this nourishment. The +thread-shaped leaf of the Onion fulfils the same office, and the +nourishing matter it prepares is deposited in its sheathing base, +forming one of the concentric layers of the onion. When these layers, so +thick and succulent, have given up their store to the growing parts +within, they are left as thin and dry husks. In a Houseleek, an Aloe or +an Agave, the green color of the surface of the fleshy leaf indicates +that it is doing the work of foliage; the deeper-seated white portion +within is the storehouse of the nourishment which the green surface has +elaborated. So, also, the seed-leaves or cotyledons are commonly used +for storage. Some, as in one of the Maples, the Pea, Horse-chestnut, +Oak, etc., are for nothing else. Others, as in Beech and in our common +<span class="pagenum"><a name="Page_63" id="Page_63">[Pg 63]</a></span>Beans, give faint indications of service as foliage also, chiefly in +vain. Still others, as in the Pumpkin and Flax, having served for +storage, develop into the first efficient foliage. Compare <a href="#UNIT_11">11</a>, <a href="#UNIT_22">22-30</a>, +and the accompanying figures.</p> + +<div class="figcenter"> +<a name="Fig170" id="Fig170"></a> +<img src="images/fig170.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 170. Series of bud-scales and foliage-leaves from a +developing bud of the Low Sweet Buckeye (Æsculus parviflora), showing +nearly complete gradation, from a scale to a compound leaf of five +leaflets; and that the scales answer to reduced petioles.</p> +</div> + +<p><a name="UNIT_166" id="UNIT_166">166.</a> <b>Leaves as Bud-Scales</b> serve to protect the forming parts within. +Having fulfilled this purpose they commonly fall off when the shoot +develops and foliage-leaves appear. Occasionally, as in Fig. <a href="#Fig170">170</a>, there +is a transition of bud-scales to leaves, which reveals the nature of the +former. The Lilac also shows a gradation from bud-scale to simple leaf. +In Cornus florida (the Flowering Dogwood), the four bud-scales which +through the winter protect the head of forming flowers remain until +blossoming, and then the base of each grows out into <span class="pagenum"><a name="Page_64" id="Page_64">[Pg 64]</a></span>a large and very +showy petal-like leaf; the original dry scale is apparent in the notch +at the apex.</p> + +<div class="figcenter"> +<a name="Fig171" id="Fig171"></a> +<img src="images/fig171.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 171. Shoot of common Barberry, showing transition of +foliage-leaves to spines.</p> +</div> + +<p><a name="UNIT_167" id="UNIT_167">167.</a> <b>Leaves as Spines</b> occur in several plants. A familiar instance is +that of the common Barberry (Fig. <a href="#Fig171">171</a>). In almost any summer shoot, most +of the gradations may be seen between the ordinary leaves, with sharp +bristly teeth, and leaves which are reduced to a branching spine or +thorn. The fact that the spines of the Barberry produce a leaf-bud in +their axil also proves them to be leaves.</p> + +<div class="figcenter"> +<a name="Fig172" id="Fig172"></a> +<img src="images/fig172_173.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 172. Leaves of Solanum jasminoides, the petiole +adapted for climbing.</p> +<p><span class="smcap">Fig.</span> 173. Leaf of Lathyrus Aphaca, consisting of a pair +of stipules and a tendril.</p> +</div> + +<p><a name="UNIT_168" id="UNIT_168">168.</a> <b>Leaves for Climbing</b> are various in adaptation. True foliage-leaves +serve this purpose; as in Gloriosa, where the attenuated tip of a simple +leaf (otherwise like that of a Lily) hooks around a supporting object; +or in Solanum jasminoides of the gardens (Fig. <a href="#Fig172">172</a>), and in Maurandia, +etc., where the leaf-stalk coils round and clings to a support; or in +the compound leaves of Clematis and of Adlumia, in which both the +leaflets and their stalks hook or coil around the support.</p> + +<p>169. Or in a compound leaf, as in the Pea and most Vetches, and in +Cobæa, while the lower leaflets serve for foliage, some of the uppermost +are developed as tendrils for climbing (Fig. <a href="#Fig167">167</a>). In the common Pea +this is so with all but one or two pairs of leaflets.</p> + +<p>170. In one European Vetch, the leaflets are wanting and the whole +petiole is a tendril, while the stipules become the only foliage (Fig. +<a href="#Fig172">173</a>).</p> + +<p><a name="UNIT_171" id="UNIT_171">171.</a> <b>Leaves as Pitchers</b>, or hollow tubes, are familiar in the common +Pitcher-plant or Side-saddle Flower (Sarracenia, Fig. <a href="#Fig174">174</a>) of our bogs. +These pitchers are generally half full of water, in which flies and +other insects are drowned, often in such numbers as to make a rich +manure for the plant. More curious are some of the southern species of +Sarracenia, which seem to be specially adapted to the capture and +destruction of flies and other insects.</p> + +<div class="figcenter"> +<a name="Fig174" id="Fig174"></a> +<img src="images/fig174_175.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 174. Leaf of Sarracenia purpurea, entire, and +another with the upper part cut off.</p> +<p><span class="smcap">Fig.</span> 175. Leaf of Nepenthes; foliage, tendril, and +pitcher combined.</p> +</div> + +<p><span class="pagenum"><a name="Page_65" id="Page_65">[Pg 65]</a></span></p><p>172. The leaf of Nepenthes (Fig. <a href="#Fig174">175</a>) combines three structures and +uses. The expanded part below is foliage: this tapers into a tendril for +climbing; and this bears a pitcher with a lid. Insects are caught, and +perhaps digested, in the pitcher.</p> + +<div class="figcenter"> +<a name="Fig176" id="Fig176"></a> +<img src="images/fig176.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 176. Leaves of Dionæa; the trap in one of them open, +in the others closed.</p> +</div> + +<p><a name="UNIT_173" id="UNIT_173">173.</a> <b>Leaves as Fly-traps.</b> Insects are caught in another way, and more +expertly, by the most extraordinary of all the plants of this country, +the Dionæa or Venus's Fly-trap, which grows in the sandy bogs around +Wilmington, North Carolina. Here (Fig. <a href="#Fig176">176</a>) each leaf bears at its +summit an appendage which opens and shuts, in shape something like a +steel-trap, and operating much like one. For when open, no sooner does a +fly alight on its surface, and brush against any one of the two or three +bristles that grow there, than the trap suddenly closes, capturing the +intruder. If the fly escapes, the trap soon slowly opens, and is ready +for another capture. When retained, the insect is after a time moistened +by a secretion from minute glands of the inner surface, and is digested. +In the various species of Drosera or Sundew, insects are caught <span class="pagenum"><a name="Page_66" id="Page_66">[Pg 66]</a></span>by +sticking fast to very viscid glands at the tip of strong bristles, aided +by adjacent gland-tipped bristles which bend slowly toward the captive. +The use of such adaptations and operations may be explained in another +place.</p> + + +<h3><a name="Subsect_VII_3" id="Subsect_VII_3">§ 3.</a> STIPULES.</h3> + +<p>174. A leaf complete in its parts consists of blade, leaf-stalk or +petiole, and a pair of stipules. But most leaves have either fugacious +or minute stipules or none at all; many have no petiole (the blade being +<i>sessile</i> or stalkless); some have no clear distinction of blade and +petiole; and many of these, such as those of the Onion and all phyllodia +(<a href="#UNIT_166">166</a>), consist of petiole only.</p> + +<p>175. The base of the petiole is apt to be broadened and flattened, +sometimes into thin margins, sometimes into a sheath which embraces the +stem at the point of attachment.</p> + +<div class="figcenter"> +<a name="Fig177" id="Fig177"></a> +<img src="images/fig177_180.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 177. Leaf of Red Clover: <i>st</i>, stipules, adhering to +the base of <i>p</i>, the petiole; <i>b</i>, blade of three leaflets.</p> +<p><span class="smcap">Fig.</span> 178. Part of stem and leaf of Prince's-Feather +(Polygonum orientale) with the united sheathing stipules forming a +sheath or <i>ocrea</i>.</p> +<p><span class="smcap">Fig.</span> 179. Terminal winter bud of Magnolia Umbrella, +natural size. 180. Outermost bud-scale (pair of stipules) detached.</p> +</div> + +<p><a name="UNIT_176" id="UNIT_176">176.</a> <b>Stipules</b> are such appendages, either wholly or partly separated +from the petiole. When quite separate they are said to be <i>free</i>, as in +Fig. <a href="#Fig112">112</a>. When attached to the base of the petiole, as in the Rose and +in <span class="pagenum"><a name="Page_67" id="Page_67">[Pg 67]</a></span>Clover (Fig. <a href="#Fig177">177</a>), they are <i>adnate</i>. When the two stipules unite +and sheathe the stem above the insertion, as in Polygonum (Fig. <a href="#Fig177">178</a>), +this sheath is called an <i>Ocrea</i> from its likeness to a greave or +leggin.</p> + +<p>177. In Grasses, when the sheathing base of the leaf may answer to +petiole, the summit of the sheath commonly projects as a thin and short +membrane, like an ocrea: this is called a <span class="smcap">Ligula</span> or <span class="smcap">Ligule</span>.</p> + +<p>178. When stipules are green and leaf-like they act as so much foliage. +In the Pea they make up no small part of the actual foliage. In a +related plant (Lathyrus Aphaca, Fig. <a href="#Fig172">173</a>), they make the whole of it, +the remainder of the leaf being tendril.</p> + +<p>179. In many trees the stipules are the bud-scales, as in the Beech, and +very conspicuously in the Fig-tree, Tulip-tree, and Magnolia (Fig. <a href="#Fig177">179</a>). +These fall off as the leaves unfold.</p> + +<p>180. The stipules are spines or prickles in Locust and several other +Leguminous trees and shrubs; they are tendrils in Smilax or Greenbrier.</p> + + +<h3><a name="Subsect_VII_4" id="Subsect_VII_4">§ 4.</a> THE ARRANGEMENT OF LEAVES.</h3> + +<p><a name="UNIT_181" id="UNIT_181">181.</a> <b>Phyllotaxy</b>, meaning leaf-arrangement, is the study of the position +of leaves, or parts answering to leaves, upon the stem.</p> + +<div class="figcenter"> +<a name="Fig181" id="Fig181"></a> +<img src="images/fig181_182.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 181. Alternate leaves, in Linden, Lime-tree, or +Basswood.</p> +<p><span class="smcap">Fig.</span> 182. Opposite leaves, in Red Maple.</p> +</div> + +<p>182. The technical name for the attachment of leaves to the stem is the +<i>insertion</i>. Leaves (as already noticed, <a href="#UNIT_54">54</a>) are <i>inserted</i> in three +modes. They are</p> + +<p><i>Alternate</i> (Fig. <a href="#Fig181">181</a>), that is, one after another, or in other words, +with only a single leaf to each node;</p> + +<p><span class="pagenum"><a name="Page_68" id="Page_68">[Pg 68]</a></span></p><p><i>Opposite</i> (Fig. <a href="#Fig181">182</a>), when there is a pair to each node, the two +leaves in this case being always on opposite sides of the stem;</p> + +<p><i>Whorled</i> or <i>Verticillate</i> (Fig. <a href="#Fig183">183</a>) when there are more than two +leaves on a node, in which case they divide the circle equally between +them, forming a <i>Verticel</i> or whorl. When there are three leaves in the +whorl, the leaves are one third of the circumference apart; when four, +one quarter, and so on. So the plan of opposite leaves, which is very +common, is merely that of whorled leaves, with the fewest leaves to the +whorl, namely, two.</p> + +<div class="figcenter"> +<a name="Fig183" id="Fig183"></a> +<img src="images/fig183.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 183. Whorled leaves of Galium. +</div> + +<p>183. In both modes and in all their modifications, the arrangement is +such as to distribute the leaves systematically and in a way to give +them a good exposure to the light.</p> + +<div class="figcenter"> +<a name="Fig184" id="Fig184"></a> +<img src="images/fig184_185.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 184. A piece of stem of Larch with two clusters +(fascicles) of numerous leaves.</p> +<p><span class="smcap">Fig.</span> 185. Piece of a branch of Pitch Pine, with three +leaves in a fascicle or bundle, in the axil of a thin scale which +answers to a primary leaf. The bundle is surrounded at the base by a +short sheath, formed of the delicate scales of the axillary bud.</p> +</div> + +<p>184. No two or more leaves ever grow from the same point. The so-called +<i>Fascicled</i> or <i>Clustered</i> leaves are the leaves of a branch the nodes +of which are very close, just as they are in the bud, so keeping the +leaves in a cluster. This is evident in the Larch (Fig. <a href="#Fig184">184</a>), in which +examination shows each cluster to be made up of numerous leaves crowded +on a spur or short axis. In spring there are only such clusters; but in +summer some of them lengthen into ordinary shoots with scattered +alternate leaves. So, likewise, each cluster of two or three +needle-shaped leaves in Pitch Pines (as in Fig. <a href="#Fig184">185</a>), or of five leaves +in White Pine, answers to a similar extremely short branch, springing +from the axil of a thin and slender scale, which represents a leaf of +the main shoot. For Pines produce two kinds of leaves,—1. primary, the +proper leaves of the shoots, not as foliage, but in the shape of +delicate scales in spring, which soon fall away; and 2. secondary, the +<i>fascicled</i> leaves, from buds in the axils of the former, and these form +the actual foliage.</p> + +<p><span class="pagenum"><a name="Page_69" id="Page_69">[Pg 69]</a></span></p><p><a name="UNIT_185" id="UNIT_185">185.</a> <b>Phyllotaxy of Alternate Leaves.</b> Alternate leaves are distributed +along the stem in an order which is uniform for each species. The +arrangement in all its modifications is said to be <i>spiral</i>, because, if +we draw a line from the <i>insertion</i> (i. e. the point of attachment) of +one leaf to that of the next, and so on, this line will wind spirally +around the stem as it rises, and in the same species will always bear +the same number of leaves for each turn round the stem. That is, any two +successive leaves will always be separated from each other by an equal +portion of the circumference of the stem. The distance in <i>height</i> +between any two leaves may vary greatly, even on the same shoot, for +that depends upon the length of the <i>internodes</i>, or spaces between the +leaves; but the distance as measured around the circumference (in other +words, the <i>Angular Divergence</i>, or angle formed by any two successive +leaves) is uniformly the same.</p> + +<p><a name="UNIT_186" id="UNIT_186">186.</a> <b>Two-ranked.</b> The greatest possible divergence is, of course, where +the second leaf stands on exactly the opposite side of the stem from the +first, the third on the side opposite the second, and therefore over the +first, and the fourth over the second. This brings all the leaves into +two ranks, one on one side of the stem and one on the other, and is +therefore called the <i>Two-ranked</i> arrangement. It occurs in all +Grasses,—in Indian Corn, for instance; also, in the Basswood (Fig. +<a href="#Fig181">181</a>). This is the simplest of all arrangements, and the one which most +widely distributes successive leaves, but which therefore gives the +fewest vertical ranks. Next is the</p> + +<p><a name="UNIT_187" id="UNIT_187">187.</a> <b>Three-ranked</b> arrangement,—that of all Sedges, and of White +Hellebore. Here the second leaf is placed one third of the way round the +stem, the third leaf two thirds of the way round, the fourth leaf +accordingly directly over the first, the fifth over the second, and so +on. That is, three leaves occur in each turn round the stem, and they +are separated from each other by one third of the circumference. (Fig. +<a href="#Fig186">186, 187</a>.)</p> + +<div class="figcenter"> +<a name="Fig186" id="Fig186"></a> +<img src="images/fig186_187.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 186. Two-ranked arrangement, shown in a piece of the +stalk of a Sedge, with the leaves cut off above their bases; the leaves +are numbered in order, from 1 to 6. 187. Diagram or cross-section of the +same, in one plane; the leaves similarly numbered; showing two cycles of +three.</p> +</div> + +<p><a name="UNIT_188" id="UNIT_188">188.</a> <b>Five-ranked</b> is the next in the series, and the most common. It is +seen in the Apple (Fig. <a href="#Fig188">188</a>), Cherry, Poplar, and the greater number of +trees and shrubs. In this case the line traced from leaf to leaf will +pass twice round the stem before it reaches a leaf <span class="pagenum"><a name="Page_70" id="Page_70">[Pg 70]</a></span>situated directly +over any below (Fig. <a href="#Fig189">189</a>). Here the sixth leaf is over the first; the +leaves stand in five perpendicular ranks, with equal angular distance +from each other; and this distance between any two successive leaves is +just two fifths of the circumference of the stem.</p> + +<div class="figcenter"> +<a name="Fig188" id="Fig188"></a> +<img src="images/fig188.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 188. Shoot with its leaves 5-ranked, the sixth leaf +over the first; as in the Apple-tree.</p> +</div> + +<div class="figcenter"> +<a name="Fig189" id="Fig189"></a> +<img src="images/fig189_190.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 189. Diagram of this arrangement, with a spiral line +drawn from the attachment of one leaf to the next, and so on; the parts +on the side turned from the eye are fainter.</p> +<p><span class="smcap">Fig.</span> 190. A ground-plan of the same; the section of the +leaves similarly numbered; a dotted line drawn from the edge of one leaf +to that of the next marks out the spiral.</p> +</div> + +<p>189. The five-ranked arrangement is expressed by the fraction 2/5. This +fraction denotes the divergence of the successive leaves, i. e. the +angle they form with each other: the numerator also expresses the number +of turns made round the stem by the spiral line in completing one cycle +or set of leaves, namely, two; and the denominator gives the number of +leaves in each cycle, or the number of perpendicular ranks, namely, +five. In the same way the fraction 1/2 stands for the two-ranked mode, +and 1/3 for the three-ranked: and so these different sorts are expressed +by the series of fractions 1/2, 1/3, 2/5. Other cases follow in the same +numerical progression, the next being the</p> + +<p><a name="UNIT_190" id="UNIT_190">190.</a> <b>Eight-ranked</b> arrangement. In this the ninth leaf stands over the +first, and three turns are made around the stem to reach it; so it is +expressed by the fraction 3/8. This is seen in the Holly, and in the +common Plantain. Then comes the</p> + +<p><a name="UNIT_191" id="UNIT_191">191.</a> <b>Thirteen-ranked</b> arrangement, in which the fourteenth leaf is over +the first, after five turns around the stem. The common Houseleek (Fig. +<a href="#Fig191">191</a>) is a good example.</p> + +<p>192. The series so far, then, is 1/2, 1/3, 2/5, 3/8, 5/13; the numerator +and the denominator of each fraction being those of the two next +preceding ones added together. At this rate the next higher should be +8/21, then 13/34, and so on; and in fact just such cases are met with, +and (commonly) no others. These higher sorts are found in the Pine +Family, both in the leaves and the cones and in many other plants with +small and crowded leaves. But in those the number of the ranks, or of +leaves in each cycle, can only rarely <span class="pagenum"><a name="Page_71" id="Page_71">[Pg 71]</a></span>be made out by direct inspection. +They may be indirectly ascertained, however, by studying the <i>secondary</i> +spirals, as they are called, which usually become conspicuous, at least +two series of them, one turning to the right and one to the left, as +shown in Fig. <a href="#Fig191">191</a>. For an account of the way in which the character of +the phyllotaxy may be deduced from the secondary spirals, see Structural +Botany, Chapter IV.</p> + +<div class="figcenter"> +<a name="Fig191" id="Fig191"></a> +<img src="images/fig191_192.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 191. A young plant of the Houseleek, with the leaves +(not yet expanded) numbered, and exhibiting the 13-ranked arrangement; +and showing secondary spirals.</p> +<p><span class="smcap">Fig.</span> 192. Opposite leaves of Euonymus, or Spindle-tree, +showing the successive pairs crossing each other at right angles.</p> +</div> + +<p><a name="UNIT_193" id="UNIT_193">193.</a> <b>Phyllotaxy of Opposite and whorled Leaves.</b> This is simple and +comparatively uniform. The leaves of each pair or whorl are placed over +the intervals between those of the preceding, and therefore under the +intervals of the pair or whorl next above. The whorls or pairs alternate +or cross each other, usually at right angles, that is, they <i>decussate</i>. +Opposite leaves, that is, whorls of two leaves only, are far commoner +than whorls of three or four or more members. This arrangement in +successive decussating pairs gives an advantageous distribution on the +stem in four vertical ranks. Whorls of three give six vertical ranks, +and so on. Note that in descriptive botany leaves in whorls of two are +simply called <i>opposite</i> leaves; and that the term <i>verticillate</i> or +<i>whorled</i>, is employed only for cases of more than two, unless the +latter number is specified.</p> + +<p><a name="UNIT_194" id="UNIT_194">194.</a> <b>Vernation or Præfoliation</b>, the disposition of the leaf-blades in +the bud, comprises two things; 1st, the way in which each separate leaf +is folded, coiled, or packed up in the bud; and 2d, the arrangement of +the leaves in the bud with respect to one another. The latter of course +depends very much upon the phyllotaxy, i. e. the position and order of +the leaves upon the stem. The same terms are used for it as for the +arrangement of the leaves of the flower in the flower-bud. See, +therefore, <a href="#UNIT_276">"Æstivation, or Præfloration."</a></p> + +<p>195. As to each leaf separately, it is sometimes <i>straight</i> and open in +vernation, but more commonly it is either <i>bent</i>, <i>folded</i>, or <i>rolled +up</i>. When the upper part is bent down upon the lower, as the young blade +in the Tulip-tree is bent upon the leaf-stalk, it is said to be +<i>Inflexed</i> or <i>Reclined</i> in vernation. When folded by the midrib so that +the two halves are placed face to face, it is <i>Conduplicate</i> (Fig. <a href="#Fig193">193</a>), +as in the Magnolia, the Cherry, and the Oak. When folded back and forth +like the plaits of a fan, it is <span class="pagenum"><a name="Page_72" id="Page_72">[Pg 72]</a></span><i>Plicate</i> or <i>Plaited</i> (Fig. <a href="#Fig193">194</a>), as +in the Maple and Currant. If rolled, it may be so either from the tip +downwards, as in Ferns and the Sundew (Fig. <a href="#Fig193">197</a>), when in unrolling it +resembles the head of a crosier, and is said to be <i>Circinate</i>; or it +may be rolled up parallel with the axis, either from one edge into a +coil, when it is <i>Convolute</i> (Fig. <a href="#Fig193">195</a>), as in the Apricot and Plum; or +rolled from both edges towards the midrib,—sometimes inwards, when it +is <i>Involute</i> (Fig. <a href="#Fig193">198</a>), as in the Violet and Water-Lily; sometimes +outwards, when it is <i>Revolute</i> (Fig. <a href="#Fig193">196</a>), in the Rosemary and Azalea. +The figures are diagrams, representing sections through the leaf, in the +way they were represented by Linnæus.</p> + +<div class="figcenter"> +<a name="Fig193" id="Fig193"></a> +<img src="images/fig193_198.jpg" alt="" /><br /> +</div> + + + +<hr class="chap" /> +<h2><a name="Section_VIII" id="Section_VIII"></a><span class="smcap">Section VIII.</span> FLOWERS.</h2> + + +<p>196. Flowers are for the production of seed (<a href="#UNIT_16">16</a>). Stems and branches, +which for a time put forth leaves for vegetation, may at length put +forth flowers for reproduction.</p> + + +<h3><a name="Subsect_VIII_1" id="Subsect_VIII_1">§ 1.</a> POSITION AND ARRANGEMENT OF FLOWERS, OR INFLORESCENCE.</h3> + +<p>197. Flower-buds appear just where leaf-buds appear; that is, they are +either <i>terminal</i> or <i>axillary</i> (<a href="#UNIT_47">47-49</a>). Morphologically, flowers answer +to shoots or branches, and their parts to leaves.</p> + +<p>198. In the same species the flowers are usually from axillary buds +only, or from terminal buds only; but in some they are both axillary and +terminal.</p> + +<p><a name="UNIT_199" id="UNIT_199">199.</a> <b>Inflorescence</b>, which is the name used by Linnæus to signify mode of +flower-arrangement, is accordingly of three classes: namely, +<i>Indeterminate</i>, when the flowers are in the axils of leaves, that is, +are from axillary buds; <i>Determinate</i>, when they are from terminal buds, +and so <i>terminate</i> a stem or branch; and <i>Mixed</i>, when these two are +combined.</p> + +<p><a name="UNIT_200" id="UNIT_200">200.</a> <b>Indeterminate Inflorescence</b> (likewise, and for the same reason, +called <i>indefinite inflorescence</i>) is so named because, as the flowers +all come from axillary buds, the terminal bud may keep on growing and +prolong the stem indefinitely. This is so in Moneywort (Fig. <a href="#Fig199">199</a>).</p> + +<div class="figcenter"> +<a name="Fig199" id="Fig199"></a> +<img src="images/fig199.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 199. Piece of a flowering-stem of Moneywort +(Lysimachia nummularia,) with single flowers successively produced in +the axils of the leaves, from below upwards, as the stem grows on.</p> +</div> + +<p><span class="pagenum"><a name="Page_73" id="Page_73">[Pg 73]</a></span></p><p>201. When flowers thus arise singly from the axils of ordinary leaves, +they are <i>axillary</i> and <i>solitary</i>, not collected into flower-clusters.</p> + +<p>202. But when several or many flowers are produced near each other, the +accompanying leaves are apt to be of smaller size, or of different shape +or character: then they are called <span class="smcap">Bracts</span>, and the flowers thus brought +together form a cluster. The kinds of flower-clusters of the +indeterminate class have received distinct names, according to their +form and disposition. They are principally <i>Raceme</i>, <i>Corymb</i>, <i>Umbel</i>, +<i>Spike</i>, <i>Head</i>, <i>Spadix</i>, <i>Catkin</i>, and <i>Panicle</i>.</p> + +<p>203. In defining these it will be necessary to use some of the following +terms of descriptive botany which relate to inflorescence. If a flower +is stalkless, i. e. sits directly in the axil or other support, it is +said to be <i>sessile</i>. If raised on a naked stalk of its own (as in Fig. +<a href="#Fig199">199</a>) it is <i>pedunculate</i>, and the stalk is a <span class="smcap">Peduncle</span>.</p> + +<p><a name="UNIT_204" id="UNIT_204">204.</a> A peduncle on which a flower-cluster is raised is a <i>Common +peduncle</i>. That which supports each separate flower of the cluster is a +<i>Partial peduncle</i>, and is generally called a <span class="smcap">Pedicel</span>. The portion of +the general stalk along which flowers are disposed is called the <i>Axis +of inflorescence</i>, or, when covered with sessile flowers, the <i>Rhachis</i> +(back-bone), and sometimes the <i>Receptacle</i>. The leaves of a +flower-cluster generally are termed <span class="smcap">Bracts</span>. But when bracts of different +orders are to be distinguished, those on the common peduncle or axis, +and which have a flower in their axil, keep the name of <i>bracts</i>; and +those on the pedicels or partial flower-stalks, if any, that of +<span class="smcap">Bractlets</span> or <i>Bracteoles</i>. The former is the preferable English name.</p> + +<div class="figcenter"> +<a name="Fig200" id="Fig200"></a> +<img src="images/fig200.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 200. A raceme, with a general peduncle (<i>p</i>), +pedicels (<i>p'</i>), bracts (<i>b</i>), and bractlets (<i>b'</i>). Plainly the bracts +here answer to the leaves in Fig. <a href="#Fig199">199</a>.</p> +</div> + +<p><a name="UNIT_205" id="UNIT_205">205.</a> <b>A Raceme</b> (Fig. <a href="#Fig200">200</a>) is that form of flower-cluster in which the +flowers, each on their own foot-stalk or pedicel, are arranged along the +sides of a common stalk or axis of inflorescence; as in the Lily of the +Valley, Currant, Barberry, one section of Cherry, etc. Each flower comes +from the axil of a small leaf, or bract, which, however, is often so +small that it might escape notice, and even sometimes (as in the Mustard +Family) disappears altogether. The lowest blossoms of a <span class="pagenum"><a name="Page_74" id="Page_74">[Pg 74]</a></span>raceme are of +course the oldest, and therefore open first, and the order of blossoming +is <i>ascending</i> from the bottom to the top. The summit, never being +stopped by a terminal flower, may go on to grow, and often does so (as +in the common Shepherd's Purse), producing lateral flowers one after +another for many weeks.</p> + +<div class="figcenter"> +<a name="Fig201" id="Fig201"></a> +<img src="images/fig201_203.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 201. A raceme. 202. A corymb. 203. An umbel. +</div> + +<p><a name="UNIT_206" id="UNIT_206">206.</a> <b>A Corymb</b> (Fig. <a href="#Fig201">202</a>) is the same as a raceme, except that it is flat +and broad, either convex, or level-topped. That is, a raceme becomes a +corymb by lengthening the lower pedicels while the uppermost remain +shorter. The axis of a corymb is short in proportion to the lower +pedicels. By extreme shortening of the axis the corymb may be converted +into</p> + +<p><a name="UNIT_207" id="UNIT_207">207.</a> <b>An Umbel</b> (Fig. <a href="#Fig201">203</a>) as in the Milkweed, a sort of flower-cluster +where the pedicels all spring apparently from the same point, from the +top of the peduncle, so as to resemble, when spreading, the rays of an +umbrella; whence the name. Here the pedicels are sometimes called the +<i>Rays</i> of the umbel. And the bracts, when brought in this way into a +cluster or circle, form what is called an <span class="smcap">Involucre</span>.</p> + +<p>208. The corymb and the umbel being more or less level-topped, bringing +the flowers into a horizontal plane or a convex form, the ascending +order of development appears as <i>Centripetal</i>. That is, the flowering +proceeds from the margin or circumference regularly towards the centre; +the lower flowers of the former answering to the outer ones of the +latter.</p> + +<p>209. In these three kinds of flower-clusters, the flowers are raised on +conspicuous <i>pedicels</i> (<a href="#UNIT_204">204</a>) or stalks of their own. The shortening of +these pedicels, so as to render the flowers <i>sessile</i> or nearly so, +converts a raceme into a <i>Spike</i>, and a corymb or an umbel into a +<i>Head</i>.</p> + +<p><a name="UNIT_210" id="UNIT_210">210.</a> <b>A Spike</b> is a flower cluster with a more or less lengthened axis, +along which the flowers are sessile or nearly so; as in the Plantain +(Fig. <a href="#Fig204">204</a>).</p> + +<div class="figcenter"> +<a name="Fig204" id="Fig204"></a> +<img src="images/fig204.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 204. Spike of the common Plantain or Ribwort. +</div> + +<p><a name="UNIT_211" id="UNIT_211">211.</a> <b>A Head</b> (<i>Capitulum</i>) is a round or roundish cluster of flowers, +<span class="pagenum"><a name="Page_75" id="Page_75">[Pg 75]</a></span>which are sessile on a very short axis or receptacle, as in the +Button-ball, Button-bush (Fig. <a href="#Fig205">205</a>), and Red Clover. It is just what a +spike would become if its axis were shortened; or an umbel, if its +pedicels were all shortened until the flowers became sessile. The head +of the Button-bush is naked; but that of the Thistle, of the Dandelion, +and the like, is surrounded by empty bracts, which form an <i>Involucre</i>. +Two particular forms of the spike and the head have received particular +names, namely, the <i>Spadix</i> and the <i>Catkin</i>.</p> + +<div class="figcenter"> +<a name="Fig205" id="Fig205"></a> +<img src="images/fig205_206.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 205. Head of the Button-bush (Cephalanthus).</p> +<p><span class="smcap">Fig.</span> 206. Spadix and spathe of the Indian Turnip; the +latter cut through below.</p> +</div> + +<p><a name="UNIT_212" id="UNIT_212">212.</a> <b>A Spadix</b> is a fleshy spike or head, with small and often imperfect +flowers, as in the Calla, Indian Turnip, (Fig. <a href="#Fig205">206</a>), Sweet Flag, etc. It +is commonly surrounded or embraced by a peculiar enveloping leaf, called +a <span class="smcap">Spathe</span>.</p> + +<p><a name="UNIT_213" id="UNIT_213">213.</a> <b>A Catkin, or Ament</b>, is the name given to the scaly sort of spike of +the Birch (Fig. <a href="#Fig207">207</a>) and Alder, the Willow and Poplar, and one sort of +flower-clusters of the Oak, Hickory, and the like,—the so-called +<i>Amentaceous</i> trees.</p> + +<div class="figcenter"> +<a name="Fig207" id="Fig207"></a> +<img src="images/fig207.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 207. Catkin, or Ament, of Birch. +</div> + +<p>214. <i>Compound</i> flower-clusters of these kinds are not uncommon. When +the stalks which in the simple umbel are the pedicels of single flowers +themselves branch into an umbel, a <i>Compound Umbel</i> is formed. <span class="pagenum"><a name="Page_76" id="Page_76">[Pg 76]</a></span>This is +the inflorescence of Caraway (Fig. <a href="#Fig208">208</a>), Parsnip, and almost all of the +great family of Umbelliferous (umbel-bearing) plants.</p> + +<div class="figcenter"> +<a name="Fig208" id="Fig208"></a> +<img src="images/fig208.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 208. Compound Umbel of Caraway. +</div> + +<p>215. The secondary or partial umbels of a compound umbel are <span class="smcap">Umbellets</span>. +When the umbellets are subtended by an involucre, this secondary +involucre is called an <span class="smcap">Involucel</span>.</p> + +<p>216. A <i>Compound raceme</i> is a cluster of racemes racemosely arranged, as +in Smilacina racemosa. A <i>compound corymb</i> is a corymb some branches of +which branch again in the same way, as in Mountain Ash. A <i>compound +spike</i> is a spicately disposed cluster of spikes.</p> + +<div class="figcenter"> +<a name="Fig209" id="Fig209"></a> +<img src="images/fig209.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 209. Diagram of a simple panicle. +</div> + +<p><a name="UNIT_217" id="UNIT_217">217.</a> <b>A Panicle</b>, such as that of Oats and many Grasses, is a compound +flower-cluster of a more or less open sort which branches with apparent +irregularity, neither into corymbs nor racemes. Fig. <a href="#Fig209">209</a> represents the +simplest panicle. It is, as it were, a raceme of which some of the +pedicels have branched so as to bear a few flowers on pedicels of their +own, while others remain simple. A <i>compound panicle</i> is one that +branches in this way again and again.</p> + +<div class="figcenter"> +<a name="Fig210" id="Fig210"></a> +<img src="images/fig210_212.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 210. Diagram of an opposite-leaved plant, with a +single terminal flower. 211. Same, with a cyme of three flowers; <i>a</i>, +the first flower, of the main axis; <i>b b</i>, those of branches. 212. Same, +with flowers also of the third order, <i>c c</i>.</p> +</div> + +<p><a name="UNIT_218" id="UNIT_218">218.</a> <b>Determinate Inflorescence</b> is that in which the flowers are from +terminal buds. The simplest case is that of a solitary terminal flower, +as in Fig. <a href="#Fig210">210</a>. This stops the growth of the stem; for its terminal bud, +becoming a blossom, can no more lengthen in the manner of a leaf-bud. +Any <span class="pagenum"><a name="Page_77" id="Page_77">[Pg 77]</a></span>further growth must be from axillary buds developing into branches. +If such branches are leafy shoots, at length terminated by single +blossoms, the inflorescence still consists of solitary flowers at the +summit of stem and branches. But if the flowering branches bear only +bracts in place of ordinary leaves, the result is the kind of +flower-cluster called</p> + +<p><a name="UNIT_219" id="UNIT_219">219.</a> <b>A Cyme.</b> This is commonly a flat-topped or convex flower-cluster, +like a corymb, only the blossoms are from terminal buds. Fig. <a href="#Fig210">211</a> +illustrates the simplest cyme in a plant with opposite leaves, namely, +with three flowers. The middle flower, <i>a</i>, terminates the stem; the two +others, <i>b b</i>, terminate branches, one from the axil of each of the +uppermost leaves; and being later than the middle one, the flowering +proceeds from the centre outwards, or is <i>Centrifugal</i>. This is the +opposite of the indeterminate mode, or that where all the flower-buds +are axillary. If flowering branches appear from the axils below, the +lower ones are the later, so that the order of blossoming continues +<i>centrifugal</i> or, which is the same thing, <i>descending</i>, as in Fig. <a href="#Fig213">213</a>, +making a sort of reversed raceme or <i>false raceme</i>,—a kind of cluster +which is to the true raceme just what the flat cyme is to the corymb.</p> + +<div class="figcenter"> +<a name="Fig213" id="Fig213"></a> +<img src="images/fig213.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 213. Diagram of a simple cyme in which the axis +lengthens, so as to take the form of a raceme.</p> +</div> + +<p>220. Wherever there are bracts or leaves, buds may be produced from +their axils and appear as flowers. Fig. <a href="#Fig210">212</a> represents the case where +the branches, <i>b b</i>, of Fig. <a href="#Fig210">211</a>, each with a pair of small leaves or +bracts about their middle, have branched again, and produced the +branchlets and flowers <i>c c</i>, on each side. It is the continued +repetition of this which forms the full or compound cyme, such as that +of the Laurestinus, Hobble-bush, Dogwood, and Hydrangea (Fig. <a href="#Fig214">214</a>).</p> + +<div class="figcenter"> +<a name="Fig214" id="Fig214"></a> +<img src="images/fig214.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 214. Compound cyme of Hydrangea arborescens, with +neutral enlarged flowers round the circumference.</p> +</div> + +<p><a name="UNIT_221" id="UNIT_221">221.</a> <b>A Fascicle</b> (meaning a bundle), like that of the Sweet William and +Lychnis of the gardens, is only a cyme with the flowers much crowded.</p> + +<p><a name="UNIT_222" id="UNIT_222">222.</a> <b>A Glomerule</b> is a cyme still more compacted, so as to imitate a +head. It may be known from a true head by the flowers not expanding +centripetally, that is, not from the circumference towards the centre.</p> + +<p>223. The illustrations of determinate or <i>cymose</i> inflorescence have +been taken from plants with opposite leaves, which give rise to the most +regular cymes. But the Rose, Cinquefoil, Buttercup, etc., with alternate +leaves, furnish also good examples of cymose inflorescence.</p> + +<p><a name="UNIT_224" id="UNIT_224">224.</a> <b>A Cymule</b> (or diminutive cyme) is either a reduced small cyme of few +flowers, or a branch of a compound cyme, i. e. a partial cyme.</p> + +<p><a name="UNIT_225" id="UNIT_225">225.</a> <b>Scorpioid</b> or <b>Helicoid Cymes</b>, of various sorts, are forms of +determinate inflorescence (often puzzling to the student) in which one +half of the ramification fails to appear. So that they may be called +<i>incomplete cymes</i>. The commoner forms may be understood by comparing a +complete <span class="pagenum"><a name="Page_78" id="Page_78">[Pg 78]</a></span>cyme, like that of Fig. <a href="#Fig215">215</a> with Fig. <a href="#Fig216">216</a>, the diagram of a +cyme of an opposite-leaved plant, having a series of terminal flowers +and the axis continued by the development of a branch in the axil of +only one of the leaves at each node. The dotted lines on the left +indicate the place of the wanting branches, which if present would +convert this <i>scorpioid cyme</i> into the complete one of Fig. <a href="#Fig215">215</a>. Fig. +<a href="#Fig216">217</a> is a diagram of similar inflorescence with alternate leaves. Both +are kinds of <i>false racemes</i> (<a href="#UNIT_219">219</a>). When the bracts are also wanting in +such cases, as in many Borragineous plants, the true nature of the +inflorescence is very much disguised.</p> + +<div class="figcenter"> +<a name="Fig215" id="Fig215"></a> +<img src="images/fig215.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 215. A complete forking cyme of an Arenaria, or +Chickweed.</p> +</div> + +<div class="figcenter"> +<a name="Fig216" id="Fig216"></a> +<img src="images/fig216_217.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 216. Diagram of a scorpioid cyme, with opposite +leaves or bracts.</p> +<p><span class="smcap">Fig.</span> 217. Diagram of analogous scorpioid cyme, with +alternate leaves or bracts.</p> +</div> + +<p><span class="pagenum"><a name="Page_79" id="Page_79">[Pg 79]</a></span></p><p>226. These distinctions between determinate and indeterminate +inflorescence, between corymbs and cymes, and between the true and the +false raceme and spike, were not recognized by botanists much more than +half a century ago, and even now are not always attended to in +descriptions. It is still usual and convenient to describe rounded or +flat-topped and open ramification as <i>corymbose</i>, even when essentially +cymose; also to call the reversed or false racemes or spikes by these +(strictly incorrect) names.</p> + +<p><a name="UNIT_227" id="UNIT_227">227.</a> <b>Mixed Inflorescence</b> is that in which the two plans are mixed or +combined in compound clusters. A <i>mixed panicle</i> is one in which, while +the primary ramification is of the indeterminate order, the secondary or +ultimate is wholly or partly of the determinate order. A contracted or +elongated inflorescence of this sort is called a <span class="smcap">Thyrsus</span>. Lilac and +Horse-chestnut afford common examples of mixed inflorescence of this +sort. When loose and open such flower-clusters are called by the general +name of <i>Panicles</i>. The heads of Compositæ are centripetal; but the +branches or peduncles which bear the heads are usually of centrifugal +order.</p> + + +<h3><a name="Subsect_VIII_2" id="Subsect_VIII_2">§ 2.</a> PARTS OR ORGANS OF THE FLOWER.</h3> + +<p>228. These were simply indicated in Section II. <a href="#UNIT_16">16</a>. Some parts are +necessary to seed-bearing; these are <i>Essential Organs</i>, namely, the +<i>Stamens</i> and <i>Pistils</i>. Others serve for protection or for attraction, +often for both. Such are the leaves of the Flower, or the <i>Floral +Envelopes</i>.</p> + +<p><a name="UNIT_229" id="UNIT_229">229.</a> <b>The Floral Envelopes</b>, taken together, are sometimes called the +<span class="smcap">Perianth</span>, also <i>Perigone</i>, in Latin form <i>Perigonium</i>. In a flower which +possesses its full number of organs, the floral envelopes are of two +kinds, namely, an outer circle, the <span class="smcap">Calyx</span>, and an inner, the <span class="smcap">Corolla</span>.</p> + +<p><a name="UNIT_230" id="UNIT_230">230.</a> <b>The Calyx</b> is commonly a circle of green or greenish leaves, but not +always. It may be the most brightly colored part of the blossom. Each +calyx-leaf or piece is called a <span class="smcap">Sepal</span>.</p> + +<p><a name="UNIT_231" id="UNIT_231">231.</a> <b>The Corolla</b> is the inner circle of floral envelopes or +flower-leaves, usually of delicate texture and <i>colored</i>, that is, of +some other color than green. Each corolla-leaf is called a <span class="smcap">Petal</span>.</p> + +<p>232. There are flowers in abundance which consist wholly of floral +envelopes. Such are the so-called full <i>double flowers</i>, of which the +choicer roses and camellias of the cultivator are familiar examples. In +them, under the gardener's care and selection, petals have taken the +place of both stamens and pistils. These are monstrous or unnatural +flowers, incapable of producing seed, and subservient only to human +gratification. Their common name of <i>double</i> flowers is not a sensible +one: except that it is fixed by custom, it were better to translate +their Latin name, <i>flores pleni</i>, and call them <i>full flowers</i>, meaning +full of leaves.</p> + +<p>233. Moreover, certain plants regularly produce <i>neutral flowers</i>, +consisting of floral envelopes only. In Fig. <a href="#Fig214">214</a>, some are seen around +the margin <span class="pagenum"><a name="Page_80" id="Page_80">[Pg 80]</a></span>of the cyme in Hydrangea. They are likewise familiar in the +Hobble-bush and in Wild-Cranberry tree, Viburnum Oxycoccus; where they +form an attractive setting to the cluster of small and comparatively +inconspicuous perfect flowers which they adorn. In the Guelder Rose, or +Snow-ball of ornamental cultivation, all or most of the blossoms of this +same shrub are transformed into neutral flowers.</p> + +<div class="figcenter"> +<a name="Fig218" id="Fig218"></a> +<img src="images/fig218.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 218. A <i>flos plenus</i>, namely, a full double flower +of Rose.</p> +</div> + +<p><a name="UNIT_234" id="UNIT_234">234.</a> <b>The Essential Organs</b> are likewise of two kinds, placed one above or +within the other; namely, first, the <span class="smcap">Stamens</span> or fertilizing organs, and +second, the <span class="smcap">Pistils</span>, which are to be fertilized and bear the seeds.</p> + +<div class="figcenter"> +<a name="Fig219" id="Fig219"></a> +<img src="images/fig219_220.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 219. A stamen: <i>a</i>, filament; <i>b</i>, anther, +discharging pollen.</p> +<p><span class="smcap">Fig.</span> 220. A pistil; with ovary, <i>a</i>, half cut away, to +show the contained ovules; <i>b</i>, style; <i>c</i>, stigma.</p> +</div> + +<p><a name="UNIT_235" id="UNIT_235">235.</a> <b>A Stamen</b> consists of two parts, namely, the <span class="smcap">Filament</span> or stalk (Fig. +<a href="#Fig219">219</a> <i>a</i>), and the <span class="smcap">Anther</span> (<i>b</i>). The latter is the only essential part. +It is a case, commonly with two lobes or cells, each opening lengthwise +by a slit, at the proper time, and discharging a powder or dust-like +substance, usually of a yellow color. This powder is the <span class="smcap">Pollen</span>, or +fertilizing matter, to produce which is the office of the stamen.</p> + +<p><a name="UNIT_236" id="UNIT_236">236.</a> <b>A Pistil</b> (Fig. <a href="#Fig219">220</a>, <a href="#Fig221">221</a>) when complete, has three parts; <span class="smcap">Ovary</span>, +<span class="smcap">Style</span>, and <span class="smcap">Stigma</span>. The <i>Ovary</i>, at base, is the hollow portion, which +contains one or more <span class="smcap">Ovules</span> or rudimentary seeds. The <i>Style</i> is the +tapering <span class="pagenum"><a name="Page_81" id="Page_81">[Pg 81]</a></span>portion above: the <i>Stigma</i> is a portion of the style, usually +its tip, with moist naked surface, upon which grains of pollen may lodge +and adhere, and thence make a growth which extends down to the ovules. +When there is no style then the stigma occupies the tip of the ovary.</p> + +<div class="figcenter"> +<a name="Fig221" id="Fig221"></a> +<img src="images/fig221.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 221. Model of a simple pistil, with ovary cut across +and slightly opened ventrally, to show the ovules and their attachment.</p> +</div> + +<p><a name="UNIT_237" id="UNIT_237">237.</a> <b>The Torus</b> or <b>Receptacle</b> is the end of the flower-stalk, or the +portion of axis or stem out of which the several organs of the flower +grow, upon which they are borne (Fig. <a href="#Fig222">223</a>).</p> + +<div class="figcenter"> +<a name="Fig222" id="Fig222"></a> +<img src="images/fig222_223.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 222. Flower of Sedum ternatum, a Stonecrop.</p> +<p><span class="smcap">Fig.</span> 223. Parts of same, two of each kind, separated and +displayed; the torus or receptacle in the centre; <i>a</i>, a sepal; <i>b</i>, a +petal; <i>c</i>, a stamen; <i>d</i>, a pistil.</p> +</div> + +<p>238. The parts of the flower are thus disposed on the receptacle or axis +essentially as are leaves upon a very short stem; first the sepals, or +outer floral leaves; then the petals or inner floral leaves; then the +stamens; lastly, at summit or centre, the pistils, when there are two or +more of them, or the single pistil, when only one. Fig. <a href="#Fig222">223</a> shows the +organs displayed, two of each kind, of such a simple and symmetrical +flower as that of a Sedum or Stonecrop, Fig. <a href="#Fig222">222</a>.</p> + + +<h3><a name="Subsect_VIII_3" id="Subsect_VIII_3">§ 3.</a> PLAN OF FLOWER.</h3> + +<p><a name="UNIT_239" id="UNIT_239">239.</a> All flowers are formed upon one general plan, but with almost +infinite variations, and many disguises. This common plan is best +understood by taking for a type, or standard for comparison, some +<i>perfect</i>, <i>complete</i>, <i>regular</i>, and <i>symmetrical</i> blossom, and one as +simple as such a blossom could well be. Flowers are said to be</p> + +<p><i>Perfect</i> (<i>hermaphrodite</i>), when provided with both kinds of essential +organs, i. e. with both stamens and pistils.</p> + +<p><i>Complete</i>, when, besides, they have the two sets of floral envelopes, +namely, <span class="pagenum"><a name="Page_82" id="Page_82">[Pg 82]</a></span>calyx and corolla. Such are completely furnished with all that +belongs to a flower.</p> + +<p><i>Regular</i>, when all the parts of each set are alike in shape and size.</p> + +<p><i>Symmetrical</i>, when there is an equal number of parts in each set or +circle of organs.</p> + +<p>240. Flax-flowers were taken for a pattern in Section II. <a href="#UNIT_16">16</a>. But in +them the five pistils have their ovaries as it were consolidated into +one body. Sedum, Fig. <a href="#Fig222">222</a>, has the pistils and all the other parts free +from such combination. The flower is perfect, complete, regular, and +symmetrical, but is not quite as simple as it might be; for there are +twice as many stamens as there are of the other organs. Crassula, a +relative of Sedum, cultivated in the conservatories for winter +blossoming (Fig. <a href="#Fig224">224</a>) is simpler, being <i>isostemonous</i>, or with just as +many stamens as petals or sepals, while Sedum is <i>diplostemonous</i>, +having double that number: it has, indeed, two sets of stamens.</p> + +<div class="figcenter"> +<a name="Fig224" id="Fig224"></a> +<img src="images/fig224_225.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 224. Flower of a Crassula. 225. Diagram or +ground-plan of same.</p> +</div> + +<p><a name="UNIT_241" id="UNIT_241">241.</a> <b>Numerical Plan.</b> A certain number either runs through the flower or +is discernible in some of its parts. This number is most commonly either +five or three, not very rarely four, occasionally two. Thus the +<i>ground-plan</i> of the flowers thus far used for illustration is five. +That of Trillium (Fig. <a href="#Fig226">226, 227</a>) is three, as it likewise is as really, +if not as plainly, in Tulips and Lilies, Crocus, Iris, and all that +class of blossoms. In some Sedums all the flowers are in fours. In +others the first flowers are on the plan of five, the rest mostly on the +plan of four, that is, with four sepals, four petals, eight stamens (i. +e. twice four), and four pistils. Whatever the ground number may be, it +runs through the whole in symmetrical blossoms.</p> + +<div class="figcenter"> +<a name="Fig226" id="Fig226"></a> +<img src="images/fig226_227.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 226. Flower of a Trillium; its parts in threes.</p> +<p><span class="smcap">Fig.</span> 227. Diagram of flower of Trillium. In this, as in +all such diagrams of cross-section of blossoms, the parts of the outer +circle represent the calyx; the next, corolla; within, stamens (here in +two circles of three each, and the cross-section is through the +anthers); in the centre, section of three ovaries joined into a compound +one of three cells.</p> +</div> + +<p><a name="UNIT_242" id="UNIT_242">242.</a> <b>Alternation of the successive Circles.</b> In these flowers the parts +of the successive circles <i>alternate</i>; and such is the rule. That is, +<span class="pagenum"><a name="Page_83" id="Page_83">[Pg 83]</a></span>the petals stand over the intervals between the sepals; the stamens, +when of the same number, stand over the intervals between the petals; or +when twice as many, as in the Trillium, the outer set alternates with +the petals, and the inner set, alternating with the other, of course +stands before the petals; and the pistils alternate with these. This is +just as it should be on the theory that the circles of the blossom +answer to whorls of leaves, which alternate in this way. While in such +flowers the circles are to be regarded as whorls, in others they are +rather to be regarded as condensed spirals of alternate leaves. But, +however this may be, in the mind of a morphological botanist,</p> + +<p><a name="UNIT_243" id="UNIT_243">243.</a> <b>Flowers are altered Branches</b>, and their parts, therefore, altered +leaves. That is, certain buds, which might have grown and lengthened +into a leafy branch, do, under other circumstances and to accomplish +other purposes, develop into blossoms. In these the axis remains short, +nearly as it is in the bud; the leaves therefore remain close together +in sets or circles; the outer ones, those of the calyx, generally +partake more or less of the character of foliage; the next set are more +delicate, and form the corolla, while the rest, the stamens and pistils, +appear under forms very different from those of ordinary leaves, and are +concerned in the production of seed. This view gives to Botany an +interest which one who merely notices the shape and counts the parts of +blossoms, without understanding their plan, has no conception of.</p> + +<p>244. That flowers answer to branches may be shown, first, from their +position. As explained in the section on Inflorescence, flowers arise +from the same places as branches, and from no other; flower-buds, like +leaf-buds, appear either on the summit of a stem, that is, as a terminal +bud, or in the axil of a leaf, as an axillary bud. And, as the plan of a +symmetrical flower shows, the arrangement of the parts on their axis or +receptacle is that of leaves upon the stem.</p> + +<p>245. That the sepals and petals are of the nature of leaves is evident +from their appearance; they are commonly called the leaves of the +flower. The calyx is most generally green in color, and foliaceous +(leaf-like) in texture. And though the corolla is rarely green, yet +neither are proper leaves always green. In our wild Painted-cup, and in +some scarlet Sages, common in gardens, the leaves just under the flowers +are of the brightest red or scarlet, often much brighter-colored than +the corolla itself. And sometimes (as in many Cactuses, and in Carolina +Allspice) there is such a regular gradation from the last leaves of the +plant (bracts or bractlets) into the leaves of the calyx, that it is +impossible to say where the one ends and the other begins. If sepals are +leaves, so also are petals; for there is no clearly fixed limit between +them. Not only in the Carolina Allspice and Cactus (Fig. <a href="#Fig229">229</a>), but in +the Water-Lily (Fig. <a href="#Fig228">228</a>) and in a variety of flowers with more than one +row of petals, there is such a complete transition between calyx and +corolla that no one can surely tell how many of the leaves belong to the +one and how many to the other.</p> + +<p><span class="pagenum"><a name="Page_84" id="Page_84">[Pg 84]</a></span></p><div class="figcenter"> +<a name="Fig228" id="Fig228"></a> +<img src="images/fig228.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 228. Series of sepals, petals, and stamens of White +Water-Lily, showing the transitions.</p> +</div> + +<div class="figcenter"> +<a name="Fig229" id="Fig229"></a> +<img src="images/fig229.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 229. A Cactus blossom. +</div> + +<p>246. That stamens are of the same general nature as petals, and +therefore a modification of leaves, is shown by the gradual transitions +that occur between the one and the other in many blossoms; especially in +cultivated flowers, such as Roses and Camellias, when they begin to +<i>double</i>, that is, to change their stamens into petals. Some wild and +natural flowers show the same interesting transitions. The Carolina +Allspice and the White Water-Lily exhibit complete gradations not only +between sepals and petals, but between petals and stamens. The sepals of +our Water-Lily are green outside, but white and petal-like on the +inside; the petals, in many rows, gradually grow narrower towards the +centre of the flower; some of these are tipped with a trace of a yellow +anther, but still are petals; the next are more contracted and +stamen-like, but with a flat petal-like filament; and a further +narrowing of this completes the genuine stamen.</p> + +<p>247. Pistils and stamens now and then change into each other in some +Willows; pistils often turn into petals in cultivated flowers; and in +the Double Cherry they are occasionally replaced by small green leaves. +Sometimes a whole blossom changes into a cluster of green leaves, as in +the "green roses" occasionally noticed in gardens, and sometimes it +degenerates into a leafy branch. So the botanist regards pistils also as +answering to leaves; that is, to single leaves when simple and separate, +to a whorl of leaves when conjoined.</p> + + +<p><span class="pagenum"><a name="Page_85" id="Page_85">[Pg 85]</a></span></p><h3><a name="Subsect_VIII_4" id="Subsect_VIII_4">§ 4.</a> MODIFICATIONS OF THE TYPE.</h3> + +<p><a name="UNIT_248" id="UNIT_248">248.</a> <b>The Deviations</b>, as they may be called, from the assumed type or +pattern of flower are most various and extensive. The differences +between one species and another of the same genus are comparatively +insignificant; those between different genera are more striking; those +between different families and classes of plants more and more profound. +They represent different adaptations to conditions or modes of life, +some of which have obvious or probable utilities, although others are +beyond particular explanation. The principal modifications may be +conveniently classified. First those which in place of perfect +(otherwise called <i>hermaphrodite</i> or bisexual) flowers, give origin to</p> + +<p><a name="UNIT_249" id="UNIT_249">249.</a> <b>Unisexual, or Separated, or Diclinous Flowers</b>, <i>imperfect</i> flowers, +as they have been called in contradistinction to perfect flowers; but +that term is too ambiguous. In these some flowers want the stamens, +while others want the pistils. Taking hermaphrodite flowers as the +pattern, it is natural to say that the missing organs are <i>suppressed</i>. +This expression is justified by the very numerous cases in which the +missing parts are <i>abortive</i>, that is, are represented by rudiments or +vestiges, which serve to exemplify the plan, although useless as to +office. Unisexual flowers are</p> + +<p><i>Monœcious</i> (or <i>Monoicous</i>, i. e. of one household), when flowers of +both sorts or sexes are produced by the same individual plant, as in the +Ricinus or Castor-oil Plant, Fig. <a href="#Fig230">230</a>.</p> + +<p><i>Diœcious</i> (or <i>Dioicous</i>, i. e. of separate households), when the +two kinds are borne on different plants; as in Willows, Poplars, Hemp, +and Moonseed, Fig. <a href="#Fig231">231, 232</a>.</p> + +<p><i>Polygamous</i>, when the flowers are some of them perfect, and some +staminate or pistillate only.</p> + +<div class="figcenter"> +<a name="Fig230" id="Fig230"></a> +<img src="images/fig230.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 230. Unisexual flowers of Castor-oil plant: <i>s</i>, +staminate flower; <i>p</i>, pistillate flower.</p> +</div> + +<div class="figcenter"> +<a name="Fig231" id="Fig231"></a> +<img src="images/fig231_232.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 231, staminate, and 232, pistillate flower of +Moonseed.</p> +</div> + +<p><span class="pagenum"><a name="Page_86" id="Page_86">[Pg 86]</a></span></p><p>250. A blossom having stamens and no pistil is a <i>Staminate</i> or <i>Male</i> +flower. Sometimes it is called a <i>Sterile</i> flower, not appropriately, +for other flowers may equally be sterile. One having pistil but no +stamens is a <i>Pistillate</i> or <i>Female</i> flower.</p> + +<div class="figcenter"> +<a name="Fig233" id="Fig233"></a> +<img src="images/fig233.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 233. Flower of Anemone Pennsylvanica; apetalous, +hermaphrodite.</p> +</div> + +<div class="figcenter"> +<a name="Fig234" id="Fig234"></a> +<img src="images/fig234.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 234. Flower of Saururus or Lizard's-tail; naked, but +hermaphrodite.</p> +</div> + +<p><a name="UNIT_251" id="UNIT_251">251.</a> <b>Incomplete Flowers</b> are so named in contradistinction to complete: +they want either one or both of the floral envelopes. Those of Fig. <a href="#Fig230">230</a> +are incomplete, having calyx but no corolla. So is the flower of Anemone +(Fig. <a href="#Fig233">233</a>), although its calyx is colored like a corolla. The flowers of +Saururus or Lizard's-tail, although perfect, have neither calyx nor +corolla (Fig. <a href="#Fig234">234</a>). Incomplete flowers, accordingly, are</p> + +<p><i>Naked</i> or <i>Achlamydeous</i>, destitute of both floral envelopes, as in +Fig. <a href="#Fig234">234</a>, or</p> + +<p><i>Apetalous</i>, when wanting only the corolla. The case of corolla present +and calyx wholly wanting is extremely rare, although there are seeming +instances. In fact, a single or simple perianth is taken to be a calyx, +unless the absence or abortion of a calyx can be made evident.</p> + +<div class="figcenter"> +<a name="Fig235" id="Fig235"></a> +<img src="images/fig235_238.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 235. Flower of Mustard. 236. Its stamens and pistil +separate and enlarged.</p> +<p><span class="smcap">Fig.</span> 237. Flower of a Violet. 238. Its calyx and corolla +displayed: the five smaller parts are the sepals; the five intervening +larger ones are the petals.</p> +</div> + +<p>252. In contradistinction to regular and symmetrical, very many flowers +are</p> + +<p><i>Irregular</i>, that is, with the members of some or all of the floral +circles unequal or dissimilar, and</p> + +<p><i>Unsymmetrical</i>, that is, when the circles of the flower or some of them +differ in the number of their members. (Symmetrical and unsymmetrical +are used in a different sense in some recent books, but the older use +should be adhered to). Want of numerical symmetry and irregularity +commonly go together; and both are common. Indeed, few flowers are +entirely <span class="pagenum"><a name="Page_87" id="Page_87">[Pg 87]</a></span>symmetrical beyond calyx, corolla, and perhaps stamens; and +probably no irregular blossoms are quite symmetrical.</p> + +<p><a name="UNIT_253" id="UNIT_253">253.</a> <b>Irregular and Unsymmetrical Flowers</b> may therefore be illustrated +together, beginning with cases which are comparatively free from other +complications. The blossom of Mustard, and of all the very natural +family which it represents (Fig. <a href="#Fig235">235, 236</a>), is regular but unsymmetrical +in the stamens. There are four equal sepals, four equal petals; but six +stamens, and only two members in the pistil, which for the present may +<span class="pagenum"><a name="Page_88" id="Page_88">[Pg 88]</a></span>be left out of view. The want of symmetry is in the stamens. These are +in two circles, an outer and an inner. The outer circle consists of two +stamens only; the inner has its proper number of four. The flower of +Violet, which is on the plan of five, is symmetrical in calyx, corolla, +and stamens, inasmuch as each of these circles consists of five members; +but it is conspicuously irregular in the corolla, one of the petals +being very different from the rest.</p> + +<div class="figcenter"> +<a name="Fig239" id="Fig239"></a> +<img src="images/fig239_244.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 239. Flower of a Larkspur. 240. Its calyx and +corolla displayed; the five larger parts are the sepals; the four +smaller, of two shapes, are the petals; the place of the fifth petal is +vacant. 241. Diagram of the same; the place for the missing petal marked +by a dotted line.</p> +<p><span class="smcap">Fig.</span> 242. Flower of a Monkshood. 243. Its parts +displayed; five sepals, the upper forming the hood; the two lateral +alike, broad and flat; the two lower small. The two pieces under the +hood represent the corolla, reduced to two odd-shaped petals; in centre +the numerous stamens and three pistils. 244. Diagram of the calyx and +corolla; the three dotted lines in the place of missing petals.</p> +</div> + +<p>254. The flowers of Larkspur, and of Monkshood or Aconite, which are +nearly related, are both strikingly irregular in calyx and corolla, and +considerably unsymmetrical. In Larkspur (Fig. <a href="#Fig239">239-241</a>) the irregular +calyx consists of five sepals, one of which, larger than the rest, is +prolonged behind into a large sac or spur; but the corolla is of only +four petals (of two shapes),—the fifth, needed to complete the +symmetry, being left out. And the Monkshood (Fig. <a href="#Fig239">242-244</a>) has five very +dissimilar sepals, and a corolla of only two very small and +curiously-shaped petals,—the three needed to make up the symmetry being +left out. The stamens in both are out of symmetry with the ground-plan, +being numerous. So are the pistils, which are usually diminished to +three, sometimes to two or to one.</p> + +<p><a name="UNIT_255" id="UNIT_255">255.</a> <b>Flowers with Multiplication of Parts</b> are very common. The stamens +are indefinitely numerous in Larkspur and in Monkshood (Fig. <a href="#Fig239">242, 243</a>), +while the pistils are fewer than the ground-plan suggests. Most +Cactus-flowers have all the organs much increased in number (Fig. <a href="#Fig229">229</a>), +and so of the Water-Lily. In Anemone (Fig. <a href="#Fig233">233</a>) the stamens and pistils +are multiplied while the petals are left out. In Buttercups or Crowfoot, +while the sepals and petals conform to the ground-plan of five, both +stamens and pistils are indefinitely multiplied (Fig. <a href="#Fig245">245</a>).</p> + +<div class="figcenter"> +<a name="Fig245" id="Fig245"></a> +<img src="images/fig245.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 245. Flower of Ranunculus bulbosus, or Buttercup, in +section.</p> +</div> + +<p><a name="UNIT_256" id="UNIT_256">256.</a> <b>Flowers modified by Union of Parts</b>, so that these parts more or +less lose the appearance of separate leaves or other organs growing out +of the end of the stem or receptacle, are extremely common. There are +two kinds of such union, namely:—</p> + +<p><i>Coalescence</i> of parts of the same circle by their contiguous margins; +and</p> + +<p><i>Adnation</i>, or the union of adjacent circles or unlike parts.</p> + +<p><a name="UNIT_257" id="UNIT_257">257.</a> <b>Coalescence</b> is not rare in leaves, as in the upper pairs of +Honeysuckles, Fig. <a href="#Fig162">163</a>. It may all the more be expected in the crowded +circles or whorls of flower-leaves. Datura or Stramonium (Fig. <a href="#Fig246">246</a>) +shows this coalescence both in calyx and corolla, the five sepals and +the five petals being thus united to near their tips, each into a tube +or long and narrow cup. These unions make needful the following terms:—</p> + +<p><span class="pagenum"><a name="Page_89" id="Page_89">[Pg 89]</a></span></p><p><i>Gamopetalous</i>, said of a corolla the petals of which are thus +coalescent into one body, whether only at base or higher. The union may +extend to the very summit, as in Morning Glory and the like (Fig. <a href="#Fig247">247</a>), +so that the number of petals in it may not be apparent. The old name for +this was <i>Monopetalous</i>, but that means "one-petalled;" while +gamopetalous means "petals united," and therefore is the proper term.</p> + +<div class="figcenter"> +<a name="Fig246" id="Fig246"></a> +<img src="images/fig246.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 246. Flower of Datura Stramonium; gamosepalous and +gamopetalous.</p> +</div> + +<div class="figcenter"> +<a name="Fig247" id="Fig247"></a> +<img src="images/fig247.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 247. Funnelform corolla of a common Morning Glory, +detached from its polysepalous calyx.</p> +</div> + +<p><i>Polypetalous</i> is the counterpart term, to denote a corolla of +<i>distinct</i>, that is, separate petals. As it means "many petalled," it is +not the best possible name, but it is the old one and in almost +universal use.</p> + +<p><i>Gamosepalous</i> applies to the calyx when the sepals are in this way +united.</p> + +<p><i>Polysepalous</i>, to the calyx when of separate sepals or calyx-leaves.</p> + +<p><a name="UNIT_258" id="UNIT_258">258.</a> Degree of union or of separation in descriptive botany is expressed +in the same way as is the lobing of leaves (<a href="#UNIT_139">139</a>). See Fig. <a href="#Fig248">249</a>-<a href="#Fig252">253</a>, and +the explanations.</p> + +<p>259. A corolla when gamopetalous commonly shows a distinction (well +marked in Fig. <a href="#Fig248">249-251</a>) between a contracted tubular portion below, the +<span class="smcap">Tube</span>, and the spreading part above, the <span class="smcap">Border</span> or <span class="smcap">Limb</span>. The junction +between tube and limb, or a more or less enlarged upper portion of the +tube between the two, is the <span class="smcap">Throat</span>. The same is true of the calyx.</p> + +<p>260. Some names are given to particular forms of the gamopetalous +corolla, applicable also to a gamosepalous calyx, such as</p> + +<p><i>Wheel-shaped</i>, or <i>Rotate</i>; when spreading out at once, without a tube +or with a very short one, something in the shape of a wheel or of its +diverging spokes, Fig. <a href="#Fig252">252, 253</a>.</p> + +<p><i>Salver-shaped</i>, or <i>Salver-form</i>; when a flat-spreading border is +raised on <span class="pagenum"><a name="Page_90" id="Page_90">[Pg 90]</a></span>a narrow tube, from which it diverges at right angles, like +the salver represented in old pictures, with a slender handle beneath, +Fig. <a href="#Fig248">249-251</a>, <a href="#Fig254">255</a>.</p> + +<div class="figcenter"> +<a name="Fig248" id="Fig248"></a> +<img src="images/fig248_251.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 248. Polypetalous corolla of Soapwort, of five +petals with long claws or stalk-like bases.</p> +<p><span class="smcap">Fig.</span> 249. Flower of Standing Cypress (Gilia +coronopifolia); gamopetalous: the tube answering to the long claws in +248, except that they are coalescent: the limb or border (the spreading +part above) is <i>five-parted</i>, that is, the petals not there united +except at very base.</p> +<p><span class="smcap">Fig.</span> 250. Flower of Cypress-vine (Ipomœa Quamoclit); +like preceding, but limb <i>five-lobed</i>.</p> +<p><span class="smcap">Fig.</span> 251. Flower of Ipomœa coccinea; limb almost +<i>entire</i>.</p> +</div> + +<div class="figcenter"> +<a name="Fig252" id="Fig252"></a> +<img src="images/fig252_253.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 252. Wheel-shaped or rotate and five-parted corolla +of Bittersweet, Solanum Dulcamara. 253. Wheel-shaped and five-lobed +corolla of Potato.</p> +</div> + +<p><i>Bell-shaped</i>, or <i>Campanulate</i>; where a short and broad tube widens +upward, in the shape of a bell, as in Fig. <a href="#Fig254">254</a>.</p> + +<div class="figcenter"> +<a name="Fig254" id="Fig254"></a> +<img src="images/fig254_258.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 254. Flower of a Campanula or Harebell, with a +campanulate or bell-shaped corolla; 255, of a Phlox, with salver-shaped +corolla; 256, of Dead Nettle (Lamium), with labiate <i>ringent</i> (or +gaping) corolla; 257, of Snapdragon, with labiate <i>personate</i> corolla; +258, of Toad-Flax, with a similar corolla spurred at the base.</p> +</div> + +<p><i>Funnel-shaped</i>, or <i>Funnelform</i>; gradually spreading at the summit of a +tube which is narrow below, in the shape of a funnel or tunnel, as in +the corolla of the common Morning Glory (Fig. <a href="#Fig247">247</a>) and of the Stramonium +(Fig. <a href="#Fig246">246</a>).</p> + +<p><span class="pagenum"><a name="Page_91" id="Page_91">[Pg 91]</a></span></p><p><i>Tubular</i>; when prolonged into a tube, with little or no spreading at +the border, as in the corolla of the Trumpet Honeysuckle, the calyx of +Stramonium (Fig. <a href="#Fig246">246</a>), etc.</p> + +<p>261. Although sepals and petals are usually all blade or lamina (<a href="#UNIT_123">123</a>), +like a sessile leaf, yet they may have a contracted and stalk-like base, +answering to petiole. This is called its <span class="smcap">Claw</span>, in Latin <i>Unguis</i>. +<i>Unguiculate</i> petals are universal and strongly marked in the Pink +tribe, as in Soapwort (Fig. <a href="#Fig248">248</a>).</p> + +<div class="figcenter"> +<a name="Fig259" id="Fig259"></a> +<img src="images/fig259.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 259. Unguiculate (clawed) petal of a Silene; with a +two-parted crown.</p> +</div> + +<div class="figcenter"> +<a name="Fig260" id="Fig260"></a> +<img src="images/fig260.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 260. A small Passion-flower, with crown of slender +threads.</p> +</div> + +<p>262. Such petals, and various others, may have an outgrowth of the inner +face into an appendage or fringe, as in Soapwort, and in Silene (Fig. +<a href="#Fig259">259</a>), where it is at the junction of claw and blade. This is called a +<span class="smcap">Crown</span>, or <i>Corona</i>. In Passion-flowers (Fig. <a href="#Fig260">260</a>) the crown consists of +numerous threads on the base of each petal.</p> + +<div class="figcenter"> +<a name="Fig261" id="Fig261"></a> +<img src="images/fig261_262.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 261. Front view of a papilionaceous corolla. 262. +The parts of the same, displayed: <i>s</i>, Standard, or Vexillum; <i>w</i>, +Wings, or Alæ; <i>k</i>, Keel, or Carina.</p> +</div> + +<p><a name="UNIT_263" id="UNIT_263">263.</a> <b>Irregular Flowers</b> may be polypetalous, or nearly so, as in the +papilionaceous corolla; but most of them are irregular through +coalescence, which often much disguises the numerical symmetry also. As +affecting the corolla the following forms have received particular +names:</p> + +<p><a name="UNIT_264" id="UNIT_264">264.</a> <b>Papilionaceous Corolla</b>, Fig. <a href="#Fig261">261, 262</a>. This is polypetalous, except +that two of the petals cohere, usually but slightly. It belongs only to +the Leguminous or Pulse family. The name means butterfly-like; but the +likeness is hardly obvious. The names of the five petals of the +<i>papilionaceous</i> corolla are curiously incongruous. They are,</p> + +<p><span class="pagenum"><a name="Page_92" id="Page_92">[Pg 92]</a></span></p><p>The <span class="smcap">Standard</span> or <i>Banner</i> (<i>Vexillum</i>), the large upper petal which is +external in the bud and wrapped around the others.</p> + +<p>The <span class="smcap">Wings</span> (<i>Alæ</i>), the pair of side petals, of quite different shape +from the standard.</p> + +<p>The <span class="smcap">Keel</span> (<i>Carina</i>), the two lower and usually smallest petals; these +are lightly coalescent into a body which bears some likeness, not to the +keel, but to the prow of a boat; and this encloses the stamens and +pistil. A Pea-blossom is a typical example; the present illustration is +from a species of Locust, Robinia hispida.</p> + +<p><a name="UNIT_265" id="UNIT_265">265.</a> <b>Labiate Corolla</b> (Fig. <a href="#Fig254">256-258</a>), which would more properly have been +called <i>Bilabiate</i>, that is, two-lipped. This is a common form of +gamopetalous corolla; and the calyx is often bilabiate also. These +flowers are all on the plan of five; and the irregularity in the corolla +is owing to unequal union of the petals as well as to diversity of form. +The two petals of the upper or posterior side of the flower unite with +each other higher up than with the lateral petals (in Fig. <a href="#Fig254">256</a>, quite to +the top), forming the <i>Upper lip</i>: the lateral and the lower similarly +unite to form the <i>Lower lip</i>. The single notch which is generally found +at the summit of the upper lip, and the two notches of the lower lip, or +in other words the two lobes of the upper and the three of the lower +lip, reveal the real composition. So also does the alternation of these +five parts with those of the calyx outside. When the calyx is also +bilabiate, as in the Sage, this alternation gives three lobes or sepals +to the upper and two to the lower lip. Two forms of the labiate corolla +have been designated, viz.:—</p> + +<p><i>Ringent</i> or <i>Gaping</i>, when the orifice is wide open, as in Fig. <a href="#Fig254">256</a>.</p> + +<p><i>Personate</i> or <i>Masked</i>, when a protuberance or intrusion of the base of +the lower lip (called a <i>Palate</i>) projects over or closes the orifice, +as in Snapdragon and Toad-Flax, Fig. <a href="#Fig254">257, 258</a>.</p> + +<div class="figcenter"> +<a name="Fig263" id="Fig263"></a> +<img src="images/fig263_265.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 263. Corolla of a purple Gerardia laid open, showing +the four stamens; the cross shows where the fifth stamen would be, if +present.</p> +<p><span class="smcap">Fig.</span> 264. Corolla, laid open, and stamens of Pentstemon +grandiflorus, with a sterile filament in the place of the fifth stamen, +and representing it.</p> +<p><span class="smcap">Fig.</span> 265. Corolla of Catalpa laid open, displaying two +good stamens and three abortive ones or vestiges.</p> +</div> + +<p><span class="pagenum"><a name="Page_93" id="Page_93">[Pg 93]</a></span></p><p>266. There are all gradations between labiate and regular corollas. In +those of Gerardia, of some species of Pentstemon, and of Catalpa (Fig. +<a href="#Fig263">263-265</a>), the labiate character is slight, but is manifest on close +inspection. In almost all such flowers the plan of five, which is +obvious or ascertainable in the calyx and corolla, is obscured in the +stamens by the abortion or suppression of one or three of their number.</p> + +<div class="figcenter"> +<a name="Fig266" id="Fig266"></a> +<img src="images/fig266.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 266. Two flower-heads of Chiccory. +</div> + +<div class="figcenter"> +<a name="Fig267" id="Fig267"></a> +<img src="images/fig267.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 267. One of them half cut away, better showing some +of the flowers.</p> +</div> + +<p><a name="UNIT_267" id="UNIT_267">267.</a> <b>Ligulate Corolla.</b> The ligulate or <i>Strap-shaped</i> corolla mainly +belongs to the family of Compositæ, in which numerous small flowers are +gathered into a head, within an involucre that imitates a calyx. It is +best exemplified in the Dandelion and in Chiccory (Fig. <a href="#Fig266">266</a>). Each one +of these straps or <i>Ligules</i>, looking like so many petals, is the +corolla of a distinct flower: the base is a short tube, which opens out +into the ligule: the five minute teeth at the end indicate the number of +constituent petals. So this is a kind of gamopetalous corolla, which is +open along one side nearly <span class="pagenum"><a name="Page_94" id="Page_94">[Pg 94]</a></span>to the base, and outspread. The nature of +such a corolla (and of the stamens also, to be explained in the next +section) is illustrated by the flower of a Lobelia, Fig. <a href="#Fig285">285</a>.</p> + +<div class="figcenter"> +<a name="Fig268" id="Fig268"></a> +<img src="images/fig268.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 268. Head of flowers of a Coreopsis, divided +lengthwise. +</div> + +<p>268. In Asters, Daisies, Sunflower, Coreopsis (Fig. <a href="#Fig268">268</a>), and the like, +only the marginal (or <i>Ray</i>) corollas are ligulate; the rest (those of +the <i>Disk</i>) are regularly gamopetalous, tubular, and five-lobed at +summit; but they are small and individually inconspicuous, only the +<i>ray-flowers</i> making a show. In fact, those of Coreopsis and of +Sunflower are simply for show, these ray-flowers being not only sterile, +but <i>neutral</i>, that is, having neither stamens nor pistil. But in +Asters, Daisies, Golden-rods, and the like, these ray-flowers are +pistillate and fertile, serving therefore for seed-bearing as well as +for show. Let it not be supposed that the show is useless. See Section +<a href="#Section_XIII">XIII</a>.</p> + +<div class="figcenter"> +<a name="Fig269" id="Fig269"></a> +<img src="images/fig269.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 269. A slice of the preceding more enlarged, with +one tubular perfect flower (<i>a</i>) left standing on the receptacle, with +its bractlet or chaff (<i>b</i>), one ligulate and neutral ray-flower (<i>cc</i>) +and part of another; <i>dd</i>, section of bracts or leaves of the +involucre.</p> +</div> + +<p><a name="UNIT_269" id="UNIT_269">269.</a> <b>Adnation, or Consolidation</b>, is the union of the members of parts +belonging to different circles of the flower (<a href="#UNIT_256">256</a>). It is of course +understood that in this (as likewise in coalescence) the parts are not +formed and then conjoined, but are produced in union. They are born +united, as the term <i>adnate</i> implies. To illustrate this kind of union, +take the accompanying series of flowers (Fig. <a href="#Fig270">270</a>-<a href="#Fig273">274</a>), shown in +vertical section. In the first, Fig. <a href="#Fig270">270</a>, Flax-flower, there is no +adnation; sepals, petals, and stamens, are <i>free</i> as well as distinct, +being separately borne on the receptacle, one circle within or above the +next; only the five pistils have their ovaries coalescent. In Fig. <a href="#Fig270">271</a>, +a Cherry-flower, the petals and stamens are borne on the throat of the +calyx-tube; that is, the sepals are coalescent into a cup, and the +petals and stamens are adnate to the inner face of this; in other +<span class="pagenum"><a name="Page_95" id="Page_95">[Pg 95]</a></span>words, the sepals, petals, and stamens are all consolidated up to a +certain height. In Fig. <a href="#Fig270">272</a>, a Purslane-flower, the same parts are +adnate to or consolidated with the ovary up to its middle. In Fig. <a href="#Fig273">273</a>, +a Hawthorn-flower, the consolidation has extended over the whole ovary; +and petals and stamens are adnate to the calyx still further. In Fig. +<a href="#Fig273">274</a>, a Cranberry-blossom, it is the same except that all the parts are +free at the same height; all seem to arise from the top of the ovary.</p> + +<p>270. In botanical description, to express tersely such differences in +the relation of these organs to the pistil, they are said to be</p> + +<p><i>Hypogynous</i> (i. e. under the pistil) when they are all <i>free</i>, that is, +not adnate to pistil nor connate with each other, as in Fig. <a href="#Fig270">270</a>.</p> + +<div class="figcenter"> +<a name="Fig270" id="Fig270"></a> +<img src="images/fig270_272.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 270. Flax-flower in section; the parts all +free,—hypogynous.</p> +<p><span class="smcap">Fig.</span> 271. Cherry-flower in section; petals and stamens +adnate to tube of calyx,—perigynous.</p> +<p><span class="smcap">Fig.</span> 272. Purslane-flower in section; calyx, petals, +stamens, all adnate to lower half of ovary,—perigynous.</p> +</div> + +<p><i>Perigynous</i> (around the pistil) when connate with each other, that is, +when petals and stamens are <i>inserted</i> or borne on the calyx, whether as +in Cherry-flowers (Fig. <a href="#Fig270">271</a>) they are free from the pistil, or as in +Purslane and Hawthorn (Fig. <a href="#Fig270">272</a>, <a href="#Fig273">273</a>) they are also adnate below to the +ovary.</p> + +<p><i>Epigynous</i> (on the ovary) when so adnate that all these parts appear to +arise from the very summit of the ovary, as in Fig. <a href="#Fig273">274</a>. The last two +terms are not very definitely distinguished.</p> + +<p>271. Another and a simpler form of expression is to describe parts of +the flower as being</p> + +<p><i>Free</i>, when not united with or <i>inserted</i> upon other parts.</p> + +<p><i>Distinct</i>, when parts of the same kind are not united. This term is the +counterpart of coalescent, as free is the counterpart of adnate. Many +writers use the term "free" indiscriminately for both; but it is better +to distinguish them.</p> + +<p><span class="pagenum"><a name="Page_96" id="Page_96">[Pg 96]</a></span></p><p><i>Connate</i> is a term common for either not free or not distinct, that +is, for parts united congenitally, whether of same or of different +kinds.</p> + +<p><i>Adnate</i>, as properly used, relates to the union of dissimilar parts.</p> + +<p>272. In still another form of expression, the terms superior and +inferior have been much used in the sense of above and below.</p> + +<p><i>Superior</i> is said of the ovary of Flax-flower, Cherry, etc., because +above the other parts; it is equivalent to "ovary free." Or it is said +of the calyx, etc., when above the ovary, as in Fig. <a href="#Fig273">273</a>-<a href="#Fig275">275</a>.</p> + +<p><i>Inferior</i>, when applied to the ovary, means the same as "calyx adnate;" +when applied to the floral envelopes, it means that they are free.</p> + +<div class="figcenter"> +<a name="Fig273" id="Fig273"></a> +<img src="images/fig273_274.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 273. Hawthorn-blossom in section; parts adnate to +whole face of ovary, and with each other beyond; another grade of +perigynous.</p> +<p><span class="smcap">Fig.</span> 274. Cranberry-blossom in section; parts epigynous.</p> +</div> + +<p><a name="UNIT_273" id="UNIT_273">273.</a> <b>Position of Flower or of its Parts.</b> The terms superior and +inferior, or upper and lower, are also used to indicate the relative +position of the parts of a flower in reference to the axis of +inflorescence. An axillary flower stands between the bract or leaf which +subtends it and the axis or stem which bears this bract or leaf. This is +represented in sectional diagrams (as in Fig. <a href="#Fig275">275, 276</a>) by a transverse +line for the bract, and a small circle for the axis of inflorescence. +Now the side of the blossom which faces the bract is the</p> + +<p><i>Anterior</i>, or <i>Inferior</i>, or <i>Lower</i> side; while the side next the axis +is the</p> + +<p><i>Posterior</i>, or <i>Superior</i>, or <i>Upper</i> side of the flower.</p> + +<div class="figcenter"> +<a name="Fig275" id="Fig275"></a> +<img src="images/fig275_276.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 275. Diagram of papilionaceous flower (Robinia, Fig. +<a href="#Fig261">261</a>), with bract below; axis of inflorescence above.</p> +<p><span class="smcap">Fig.</span> 276. Diagram of Violet-flower; showing the relation +of parts to bract and axis.</p> +</div> + +<p>274. So, in the labiate corolla (Fig. <a href="#Fig254">256-258</a>), the lip which is +composed of three of the five petals is the <i>anterior</i>, or <i>inferior</i>, +or <i>lower</i> lip; the other is the <i>posterior</i>, or <i>superior</i>, or <i>upper</i> +lip.</p> + +<p><span class="pagenum"><a name="Page_97" id="Page_97">[Pg 97]</a></span></p><p>275. In Violets (Fig. <a href="#Fig235">238</a>, <a href="#Fig275">276</a>), the odd sepal is posterior (next the +axis); the odd petal is therefore anterior, or next the subtending leaf. +In the papilionaceous flower (Fig. <a href="#Fig261">261</a>, and diagram, Fig. <a href="#Fig275">275</a>), the odd +sepal is anterior, and so two sepals are posterior; consequently, by the +alternation, the odd petal (the standard) is posterior or upper, and the +two petals forming the keel are anterior or lower.</p> + + +<h3><a name="Subsect_VIII_5" id="Subsect_VIII_5">§ 5.</a> ARRANGEMENT OF PARTS IN THE BUD.</h3> + +<p><a name="UNIT_276" id="UNIT_276">276.</a> <b>Æstivation</b> was the fanciful name given by Linnæus to denote the +disposition of the parts, especially the leaves of the flower, before +<i>Anthesis</i>, i. e. before the blossom opens. <i>Præfloration</i>, a better +term, is sometimes used. This is of importance in distinguishing +different families or genera of plants, being generally uniform in each. +The æstivation is best seen by making a slice across the flower-bud; and +it may be expressed in diagrams, as in the accompanying figures.</p> + +<p>277. The pieces of the calyx or the corolla either overlap each other in +the bud, or they do not. When they do not overlap, the æstivation is</p> + +<p><i>Valvate</i>, when the pieces meet each other by their abrupt edges, +without any infolding or overlapping; as the calyx of the Linden or +Basswood (Fig. <a href="#Fig277">277</a>).</p> + +<div class="figcenter"> +<a name="Fig277" id="Fig277"></a> +<img src="images/fig277.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 277. Diagram of a flower of Linden, showing the +calyx valvate and corolla imbricate in the bud, etc.</p> +</div> + +<p><i>Induplicate</i>, which is valvate with the margins of each piece +projecting inwards, as in the calyx of a common Virgin's-bower, Fig. +<a href="#Fig278">278</a>, or</p> + +<div class="figcenter"> +<a name="Fig278" id="Fig278"></a> +<img src="images/fig278_279.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 278. Valvate-induplicate æstivation of calyx of +common Virgin's-bower.</p> +<p><span class="smcap">Fig.</span> 279. Valvate-involute æstivation of same in +Vine-bower, Clematis Vitialla.</p> +</div> + +<p><i>Involute</i>, which is the same but the margins rolled inward, as in most +of the large-flowered species of Clematis, Fig. <a href="#Fig278">279</a>.</p> + +<p><i>Reduplicate</i>, a rarer modification of valvate, is similar but with +margins projecting outward.</p> + +<p><i>Open</i>, the parts not touching in the bud, as the calyx of Mignonette.</p> + +<p>278. When the pieces overlap in the bud, it is in one of two ways; +either every piece has one edge in and one edge out, or some pieces are +wholly outside and others wholly inside. In the first case the +æstivation is</p> + +<p><i>Convolute</i>, also named <i>Contorted</i> or <i>Twisted</i>, as in Fig. <a href="#Fig280">280</a>, a +cross-section of a corolla very strongly thus convolute or rolled up +together, and in the corolla of a Flax-flower (Fig. <a href="#Fig281">281</a>), where the +petals only moderately overlap in this way. Here one edge of every petal +covers the next before <span class="pagenum"><a name="Page_98" id="Page_98">[Pg 98]</a></span>it, while its other edge is covered by the next +behind it. The other mode is the</p> + +<div class="figcenter"> +<a name="Fig280" id="Fig280"></a> +<img src="images/fig280.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 280. Convolute æstivation, as in the corolla-lobes +of Oleander.</p> +</div> + +<div class="figcenter"> +<a name="Fig281" id="Fig281"></a> +<img src="images/fig281.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 281. Diagram of a Flax-flower; calyx imbricated and +corolla convolute in the bud.</p> +</div> + +<p><i>Imbricate</i> or <i>Imbricated</i>, in which the outer parts cover or overlap +the inner so as to "break joints," like tiles or shingles on a roof; +whence the name. When the parts are three, the first or outermost is +wholly external, the third wholly internal, the second has one margin +covered by the first while the other overlaps the third or innermost +piece: this is the arrangement of alternate three-ranked leaves (<a href="#UNIT_187">187</a>). +When there are five pieces, as in the corolla of Fig. <a href="#Fig224">225</a>, and calyx of +Fig. <a href="#Fig281">281</a>, as also of Fig. <a href="#Fig239">241</a>, <a href="#Fig275">276</a>, two are external, two are internal, +and one (the third in the spiral) has one edge covered by the outermost, +while its other edge covers the innermost; which is just the five-ranked +arrangement of alternate leaves (<a href="#UNIT_188">188</a>). When the pieces are four, two are +outer and two are inner; which answers to the arrangement of opposite +leaves.</p> + +<p>279. The imbricate and the convolute modes sometimes vary one into the +other, especially in the corolla.</p> + +<p>280. In a gamopetalous corolla or gamosepalous calyx, the shape of the +tube in the bud may sometimes be noticeable. It may be</p> + +<p><i>Plicate</i> or <i>Plaited</i>, that is, folded lengthwise; and the plaits may +either be turned outwards, forming projecting ridges, as in the corolla +of Campanula; or turned inwards, as in that of Gentian Belladonna; or</p> + +<p><i>Supervolute</i>, when the plaits are convolutely wrapped round each other, +as in the corolla of Morning Glory and of Stramonium, Fig. <a href="#Fig282">282</a>.</p> + +<div class="figcenter"> +<a name="Fig282" id="Fig282"></a> +<img src="images/fig282.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 282. Upper part of corolla of Datura Stramonium in +the bud; and below a section showing the convolution of the plaits.</p> +</div> + + + +<hr class="chap" /> +<h2><a name="Section_IX" id="Section_IX"></a><span class="smcap">Section IX.</span> STAMENS IN PARTICULAR.</h2> + + +<p><a name="UNIT_281" id="UNIT_281">281.</a> <b>Andrœcium</b> is a technical name for the staminate system of a +flower (that is, for the stamens taken together), which it is sometimes +convenient to use. The preceding section has dealt with modifications of +the flower pertaining mainly to calyx and corolla. Those relating to the +stamens are now to be indicated. First as to</p> + +<p><span class="pagenum"><a name="Page_99" id="Page_99">[Pg 99]</a></span></p><p>282. Insertion, or place of attachment. The stamens usually go with the +petals. Not rarely they are at base</p> + +<p><i>Epipetalous</i>, that is, inserted on (or adnate to) the corolla, as in +Fig. <a href="#Fig283">283</a>. When free from the corolla, they may be</p> + +<div class="figcenter"> +<a name="Fig283" id="Fig283"></a> +<img src="images/fig283.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 283. Corolla of Morning Glory laid open, to show the +five stamens inserted on it, near the base.</p> +</div> + +<p><i>Hypogynous</i>, inserted on the receptacle under the pistil or +gynœcium.</p> + +<p><i>Perigynous</i>, inserted on the calyx, that is, with the lower part of +filament adnate to the calyx-tube.</p> + +<p><i>Epigynous</i>, borne apparently on the top of the ovary; all which is +explained in Fig. <a href="#Fig270">270</a>-<a href="#Fig273">274</a>.</p> + +<div class="figcenter"> +<a name="Fig284" id="Fig284"></a> +<img src="images/fig284.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 284. Style of a Lady's Slipper (Cypripedium), and +stamens united with it; <i>a</i>, <i>a</i>, the anthers of the two good stamens; +<i>st</i>, an abortive stamen, what should be its anther changed into a +petal-like body; <i>stig</i>, the stigma.</p> +</div> + +<p><i>Gynandrous</i> is another term relating to insertion of rarer occurrence, +that is, where the stamens are inserted on (in other words, adnate to) +the style, as in Lady's Slipper (Fig. <a href="#Fig284">284</a>), and in the Orchis family +generally.</p> + +<p><a name="UNIT_283" id="UNIT_283">283.</a> <b>In Relation to each Other</b>, stamens are more commonly</p> + +<p><i>Distinct</i>, that is, without any union with each other. But when united, +the following technical terms of long use indicate their modes of mutual +connection:—</p> + +<p><i>Monadelphous</i> (from two Greek words, meaning "in one brotherhood"), +when united by their filaments into one set, usually into a ring or cup +below, or into a tube, as in the Mallow Family (Fig. <a href="#Fig286">286</a>), the +Passion-flower (Fig. <a href="#Fig260">260</a>), the Lupine (Fig. <a href="#Fig286">287</a>), and in Lobelia (Fig. +<a href="#Fig285">285</a>).</p> + +<div class="figcenter"> +<a name="Fig285" id="Fig285"></a> +<img src="images/fig285.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 285. Flower of Lobelia cardinalis, Cardinal flower; +corolla making approach to the ligulate form; filaments (<i>st</i>) +monadelphous, and anthers (<i>a</i>) syngenesious.</p> +</div> + +<div class="figcenter"> +<a name="Fig286" id="Fig286"></a> +<img src="images/fig286_288.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 286. Flower of a Mallow, with calyx and corolla cut +away; showing monadelphous stamens.</p> +<p><span class="smcap">Fig.</span> 287. Monadelphous stamens of Lupine. 288. +Diadelphous stamens (9 and 1) of a Pea-blossom.</p> +</div> + +<p><i>Diadelphous</i> (meaning in two brotherhoods), when united by the +filaments into two sets, as in the Pea and most of its near relatives +(Fig. <a href="#Fig286">288</a>), usually nine in one set, and one in the other.</p> + +<p><i>Triadelphous</i> (three brotherhoods), when the filaments are united in +three sets or clusters, as in most species of Hypericum.</p> + +<p><span class="pagenum"><a name="Page_100" id="Page_100">[Pg 100]</a></span></p><p><i>Pentadelphous</i> (five brotherhoods), when in five sets, as in some +species of Hypericum and in American Linden (Fig. <a href="#Fig277">277</a>, <a href="#Fig289">289</a>).</p> + +<p><i>Polyadelphous</i> (many or several brotherhoods) is the term generally +employed when these sets are several, or even more than two, and the +particular number is left unspecified. These terms all relate to the +filaments.</p> + +<p><i>Syngenesious</i> is the term to denote that stamens have their anthers +united, coalescent into a ring or tube; as in Lobelia (Fig. <a href="#Fig285">285</a>), in +Violets, and in all of the great family of Compositæ.</p> + +<p><a name="UNIT_284" id="UNIT_284">284.</a> <b>Their Number</b> in a flower is commonly expressed directly, but +sometimes adjectively, by a series of terms which were the name of +classes in the Linnæan artificial system, of which the following names, +as also the preceding, are a survival:—</p> + +<p><i>Monandrous</i>, i. e. solitary-stamened, when the flower has only one +stamen,</p> + +<p><i>Diandrous</i>, when it has two stamens only,</p> + +<p><i>Triandrous</i>, when it has three stamens,</p> + +<p><i>Tetrandrous</i>, when it has four stamens,</p> + +<p><i>Pentandrous</i>, when it has five stamens,</p> + +<p><i>Hexandrous</i>, when with six stamens, and so on to</p> + +<p><i>Polyandrous</i>, when it has many stamens, or more than a dozen.</p> + +<div class="figcenter"> +<a name="Fig289" id="Fig289"></a> +<img src="images/fig289_291.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 289. One of the five stamen-clusters of the flower +of American Linden, with accompanying scale. The five clusters are shown +in section in the diagram of this flower, Fig. <a href="#Fig277">277</a>.</p> +<p><span class="smcap">Fig.</span> 290. Five syngenesious stamens of a Coreopsis. 291. +Same, with tube laid open and displayed.</p> +</div> + +<p>285. For which terms, see the Glossary. They are all Greek numerals +prefixed to <i>-andria</i> (from the Greek), which Linnæus used for +<i>andrœcium</i>, and are made into an English adjective, <i>-androus</i>. Two +other terms, of same origin, designate particular cases of number (four +or six) in connection with unequal length. Namely, the stamens are</p> + +<p><i>Didynamous</i>, when, being only four, they form two pairs, one pair +longer than the other, as in the Trumpet Creeper, in Gerardia (Fig. +<a href="#Fig263">263</a>), etc.</p> + +<p><span class="pagenum"><a name="Page_101" id="Page_101">[Pg 101]</a></span></p><p><i>Tetradynamous</i>, when, being only six, four of them surpass the other +two, as in the Mustard-flower and all the Cruciferous family, Fig. <a href="#Fig235">235</a>.</p> + +<p><a name="UNIT_286" id="UNIT_286">286.</a> <b>The Filament</b> is a kind of stalk to the anther, commonly slender or +thread-like: it is to the anther nearly what the petiole is to the blade +of a leaf. Therefore it is not an essential part. As a leaf may be +without a stalk, so the anther may be <i>Sessile</i>, or without a filament.</p> + +<div class="figcenter"> +<a name="Fig292" id="Fig292"></a> +<img src="images/fig292_294.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 292. Stamen of Isopyrum, with innate anther. 293. Of +Tulip-tree, with adnate (and extrorse) anther. 294. Of Evening Primrose, +with versatile anther.</p> +</div> + +<p><a name="UNIT_287" id="UNIT_287">287.</a> <b>The Anther</b> is the essential part of the stamen. It is a sort of +case, filled with a fine powder, <i>the Pollen</i>, which serves to fertilize +the pistil, so that it may perfect seeds. The anther is said to be</p> + +<p><i>Innate</i> (as in Fig. <a href="#Fig292">292</a>), when it is attached by its base to the very +apex of the filament, turning neither inward nor outward;</p> + +<p><i>Adnate</i> (as in Fig. <a href="#Fig292">293</a>), when attached as it were by one face, usually +for its whole length, to the side of a continuation of the filament; and</p> + +<p><i>Versatile</i> (as in Fig. <a href="#Fig292">294</a>), when fixed by or near its middle only to +the very point of the filament, so as to swing loosely, as in the Lily, +in Grasses, etc. Versatile or adnate anthers are</p> + +<p><i>Introrse</i>, or <i>Incumbent</i>, when facing inward, that is, toward the +centre of the flower, as in Magnolia, Water-Lily, etc.</p> + +<p><i>Extrorse</i>, when facing outwardly, as in the Tulip-tree.</p> + +<p>288. Rarely does a stamen bear any resemblance to a leaf, or even to a +petal or flower-leaf. Nevertheless, the botanist's idea of a stamen is +that it answers to a leaf developed in a peculiar form and for a special +purpose. In the filament he sees the stalk of the leaf; in the anther, +the blade. The blade of a leaf consists of two similar sides; so the +anther consists of two <span class="smcap">Lobes</span> or <span class="smcap">Cells</span>, one answering to the left, the +other to the right, side of the blade. The two lobes are often connected +by a prolongation of the filament, which answers to the midrib of a +leaf; this is called the <span class="smcap">Connective</span>. This is conspicuous in Fig. <a href="#Fig292">292</a>, +where the connective is so broad that it separates the two cells of the +anther to some distance.</p> + +<div class="figcenter"> +<a name="Fig295" id="Fig295"></a> +<img src="images/fig295.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 295. Diagram of the lower part of an anther, cut +across above, and the upper part of a leaf, to show how the one answers +to the other; the filament to petiole, the connective to midrib; the two +cells to the right and left halves of the blade.</p> +</div> + +<p>289. A simple conception of the morphological relation of an anther to a +leaf is given in Fig. <a href="#Fig295">295</a>, an ideal figure, the lower part representing +a stamen with the top of its anther cut away; the upper, the +corresponding upper part of a leaf.</p> + +<p><span class="pagenum"><a name="Page_102" id="Page_102">[Pg 102]</a></span></p><p>290. So anthers are generally <i>two-celled</i>. But as the pollen begins to +form in two parts of each cell (the anterior and the posterior), +sometimes these two strata are not confluent, and the anther even at +maturity may be <i>four-celled</i>, as in Moonseed (Fig. <a href="#Fig296">296</a>); or rather, in +that case (the word <i>cell</i> being used for each lateral half of the +organ), it is <i>two-celled</i>, but the cells <i>bilocellate</i>.</p> + +<div class="figcenter"> +<a name="Fig296" id="Fig296"></a> +<img src="images/fig296_299.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 296. Stamen of Moonseed, with anther cut across; +this 4-celled, or rather 4-locellate.</p> +<p><span class="smcap">Fig.</span> 297. Stamen of Pentstemon pubescens; the two +anther-cells diverging, and almost confluent.</p> +<p><span class="smcap">Fig.</span> 298. Stamen of Mallow; the anther supposed to answer +to that of Fig. 297, but the cells completely confluent into one.</p> +<p><span class="smcap">Fig.</span> 299. Stamen of Globe Amaranth; very short filament +bearing a single anther-cell; it is open from top to bottom, showing the +pollen within.</p> +</div> + +<div class="figcenter"> +<a name="Fig300" id="Fig300"></a> +<img src="images/fig300_305.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 300-305. Stamens of several plants of the Labiate or +Mint Family. <span class="smcap">Fig.</span> 300. Of a Monarda: the two anther-cells with bases +divergent so that they are transverse to the filament, and their +contiguous tips confluent, so as to form one cell opening by a +continuous line. <span class="smcap">Fig.</span> 301. Of a Calamintha: the broad connective +separating the two cells. <span class="smcap">Fig.</span> 302. Of a Sage (Salvia Texana); with long +and slender connective resembling forks of the filament, one bearing a +good anther-cell; the other an abortive or poor one. <span class="smcap">Fig.</span> 303. Another +Sage (S. coccinea), with connective longer and more thread-shaped, the +lower fork having its anther-cell wholly wanting. <span class="smcap">Fig.</span> 304. Of a White +Sage, Audibertia grandiflora; the lower fork of connective a mere +vestige. <span class="smcap">Fig.</span> 305. Of another White Sage (A. stachyoides), the lower +fork of connective suppressed.</p> +</div> + +<p>291. But anthers may become <i>one-celled</i>, and that either by confluence +or by suppression.</p> + +<p>292. By confluence, when the two cells run together into one, as they +nearly do in most species of Pentstemon (Fig. <a href="#Fig296">297</a>), more so in Monarda +(Fig. <a href="#Fig300">300</a>), and completely in the Mallow (Fig. <a href="#Fig296">298</a>) and all the Mallow +family.</p> + +<p><span class="pagenum"><a name="Page_103" id="Page_103">[Pg 103]</a></span></p><p>293. By suppression in certain cases the anther may be reduced to one +cell or halved. In Globe Amaranth (Fig. <a href="#Fig296">299</a>) there is a single cell +without vestige of any other. Different species of Sage and of the White +Sages of California show various grades of abortion of one of the +anther-cells, along with a singular lengthening of the connective (Fig. +<a href="#Fig300">302-305</a>).</p> + +<p>294. The splitting open of an anther for the discharge of its pollen is +termed its <i>Dehiscence</i>.</p> + +<div class="figcenter"> +<a name="Fig306" id="Fig306"></a> +<img src="images/fig306_308.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 306. Stamen with the usual dehiscence of anther down +the side of each cell.</p> +<p><span class="smcap">Fig.</span> 307. Stamen of Pyrola; cells opening by a terminal +hole.</p> +<p><span class="smcap">Fig.</span> 308. Stamen of Barberry; cells of anther each +opening by an uplifted valve.</p> +</div> + +<p>295. As the figures show, this is commonly by a line along the whole +length of each cell, either lateral or, when the anthers are extrorse, +often along the outer face, and when introrse, along the inner face of +each cell. Sometimes the opening is only by a chink, hole, or pore at +the top, as in the Azalea, Pyrola (Fig. <a href="#Fig306">307</a>), etc.; sometimes a part of +the face separates as a sort of trap-door (or valve), hinged at the top, +and opening to allow the escape of the pollen, as in the Sassafras, +Spice-bush, and Barberry (Fig. <a href="#Fig306">308</a>).</p> + +<p><a name="UNIT_296" id="UNIT_296">296.</a> <b>Pollen.</b> This is the powdery matter, commonly of a yellow color, +which fills the cells of the anther, and is discharged during +blossoming, after which the stamens generally fall or wither away. Under +the microscope it is found to consist of grains, usually round or oval, +and all alike in the same species, but very different in different +plants. So that the plant may sometimes be recognized from the pollen +alone. Several forms are shown in the accompanying figures.</p> + +<div class="figcenter"> +<a name="Fig309" id="Fig309"></a> +<img src="images/fig309_313.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 309. Magnified pollen of a Lily, smooth and oval; +310, of Echinocystis, grooved lengthwise; 311, of Sicyos, with bristly +points and smooth bands; 312, of Musk Plant (Mimulus), with spiral +grooves; 313, of Succory, twelve-sided and dotted.</p> +</div> + +<p><span class="pagenum"><a name="Page_104" id="Page_104">[Pg 104]</a></span></p><p>297. An ordinary pollen-grain has two coats; the outer coat thickish, +but weak, and frequently adorned with lines or bands, or studded with +points; the inner coat is extremely thin and delicate, but extensible, +and its cavity when fresh contains a thickish protoplasmic fluid, often +rendered turbid by an immense number of minute particles that float in +it. As the pollen matures this fluid usually dries up, but the +protoplasm does not lose its vitality. When the grain is wetted it +absorbs water, swells up, and is apt to burst, discharging the contents. +But when weak syrup is used it absorbs this slowly, and the tough inner +coat will sometimes break through the outer and begin a kind of growth, +like that which takes place when the pollen is placed upon the stigma.</p> + +<div class="figcenter"> +<a name="Fig314" id="Fig314"></a> +<img src="images/fig314_318.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 314. Magnified pollen of Hibiscus and other +Mallow-plants, beset with prickly projections; 315, of Circæa, with +angles bearing little lobes; 316, of Evening Primrose, the three lobes +as large as the central body; 317, of Kalmia, four grains united, as in +most of the Heath family; 318, of Pine, as it were of three grains or +cells united; the lateral empty and light.</p> +</div> + +<p>298. Some pollen-grains are, as it were, lobed (as in Fig. <a href="#Fig314">315, 316</a>), or +formed of four grains united (as in the Heath family, Fig. <a href="#Fig314">317</a>): that of +Pine (Fig. <a href="#Fig314">318</a>) has a large rounded and empty bladder-like expansion +upon each side. This renders such pollen very buoyant, and capable of +being transported to a great distance by the wind.</p> + +<p>299. In species of Acacia simple grains lightly cohere into globular +pellets. In Milkweeds and in most Orchids all the pollen of an +anther-cell is compacted or coherent into one mass, called a +<i>Pollen-mass</i>, or <span class="smcap">Pollinium</span>, plural <span class="smcap">Pollinia</span>. (Fig. <a href="#Fig319">319</a>-<a href="#Fig322">322</a>.)</p> + +<div class="figcenter"> +<a name="Fig319" id="Fig319"></a> +<img src="images/fig319.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 319. Pollen, a pair of pollinia of a Milkweed, +Asclepias, attached by stalks to a gland; moderately magnified.</p> +</div> + +<div class="figcenter"> +<a name="Fig320" id="Fig320"></a> +<img src="images/fig320_321.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 320. Pollinium of an Orchis (Habenaria), with its +stalk attached to a sticky gland; magnified. 321. Some of the packets or +partial pollinia, of which Fig. 320 is made up, more magnified.</p> +</div> + +<div class="figcenter"> +<a name="Fig322" id="Fig322"></a> +<img src="images/fig322.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 322. One of the partial pollinia, torn up at top to +show the grains (which are each composed of four), and highly +magnified.</p> +</div> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_105" id="Page_105">[Pg 105]</a></span></p> +<h2><a name="Section_X" id="Section_X"></a><span class="smcap">Section X.</span> PISTILS IN PARTICULAR.</h2> + + +<h3><a name="Subsect_X_1" id="Subsect_X_1">§ 1.</a> ANGIOSPERMOUS OR ORDINARY GYNŒCIUM.</h3> + +<p><a name="UNIT_300" id="UNIT_300">300.</a> <b>Gynœcium</b> is the technical name for the pistil or pistils of a +flower taken collectively, or for whatever stands in place of these. The +various modifications of the gynœcium and the terms which relate to +them require particular attention.</p> + +<p><a name="UNIT_301" id="UNIT_301">301.</a> The <span class="smcap">Pistil</span>, when only one, occupies the centre of the flower; when +there are two pistils, they stand facing each other in the centre of the +flower; when several, they commonly form a ring or circle; and when very +numerous, they are generally crowded in rows or spirals on the surface +of a more or less enlarged or elongated receptacle. Their number gives +rise to certain terms, the counterpart of those used for stamens (<a href="#UNIT_284">284</a>), +which are survivals of the names of orders in the Linnæan artificial +system. The names were coined by prefixing Greek numerals to <i>-gynia</i> +used for gynœcium, and changed into adjectives in the form of +<i>-gynous</i>. That is, a flower is</p> + +<p><i>Monogynous</i>, when it has a single pistil, whether that be simple or +compound;</p> + +<p><i>Digynous</i>, when it has only two pistils; <i>Trigynous</i>, when with three; +<i>Tetragynous</i>, with four; <i>Pentagynous</i>, with five; <i>Hexagynous</i>, with +six; and so on to <i>Polygynous</i>, with many pistils.</p> + +<p><a name="UNIT_302" id="UNIT_302">302.</a> <b>The Parts of a Complete Pistil</b>, as already twice explained (<a href="#UNIT_16">16</a>, +<a href="#UNIT_236">236</a>), are the <span class="smcap">Ovary</span>, the <span class="smcap">Style</span>, and the <span class="smcap">Stigma</span>. The ovary is one +essential part: it contains the rudiments of seeds, called <span class="smcap">Ovules</span>. The +stigma at the summit is also essential: it receives the pollen, which +fertilizes the ovules in order that they may become seeds. But the +style, commonly a tapering or slender column borne on the summit of the +ovary, and bearing the stigma on its apex or its side, is no more +necessary to a pistil than the filament is to the stamen. Accordingly, +there is no style in many pistils: in these the stigma is <i>sessile</i>, +that is, rests directly on the ovary (as in Fig. <a href="#Fig326">326</a>). The stigma is +very various in shape and appearance, being sometimes a little knob (as +in the Cherry, Fig. <a href="#Fig270">271</a>), sometimes a point or small surface of bare +tissue (as in Fig. <a href="#Fig326">327-330</a>), and sometimes a longitudinal crest or line +(as in Fig. <a href="#Fig323">324</a>, <a href="#Fig341">341-343</a>), or it may occupy the whole length of the +style, as in Fig. <a href="#Fig331">331</a>.</p> + +<p>303. The word Pistil (Latin, <i>Pistillum</i>) means a pestle. It came into +use in the first place for such flowers as those of Crown Imperial, or +Lily, in which the pistil in the centre was likened to the pestle, and +the perianth around it to the mortar, of the apothecary.</p> + +<p>304. A pistil is either <i>simple</i> or <i>compound</i>. It is simple when it +answers to a single flower-leaf, compound when it answers to two or +three, or a fuller circle of such leaves conjoined.</p> + +<p><span class="pagenum"><a name="Page_106" id="Page_106">[Pg 106]</a></span></p><p><a name="UNIT_305" id="UNIT_305">305.</a> <b>Carpels.</b> It is convenient to have a name for each flower-leaf of +the gynœcium; so it is called a <i>Carpel</i>, in Latin <i>Carpellum</i> or +<i>Carpidium</i>. A simple pistil is a carpel. Each component flower-leaf of +a compound pistil is likewise a carpel. When a flower has two or more +pistils, these of course are simple pistils, that is, separate carpels +or pistil-leaves. There may be only a single simple pistil to the +flower, as in a Pea or Cherry blossom (Fig. <a href="#Fig270">271</a>); there may be two such, +as in many Saxifrages; or many, as in the Strawberry. More commonly the +single pistil in the centre of a blossom is a compound one. Then there +is seldom much difficulty in ascertaining the number of carpels or +pistil-leaves that compose it.</p> + +<p><a name="UNIT_306" id="UNIT_306">306.</a> <b>The Simple Pistil</b>, viewed morphologically, answers to a leaf-blade +with margins incurved and united where they meet, so forming a closed +case or pod (the ovary), and bearing ovules at the suture or junction of +these margins: a tapering upper portion with margins similarly inrolled, +is supposed to form the style; and these same margins, exposed at the +tip or for a portion of the length, become the stigma. Compare, under +this view, the three accompanying figures.</p> + +<div class="figcenter"> +<a name="Fig323" id="Fig323"></a> +<img src="images/fig323_325.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 323. An inrolled small leaf, such as in +double-flowered Cherry blossoms is often seen to occupy the place of a +pistil.</p> +<p><span class="smcap">Fig.</span> 324. A simple pistil (of Isopyrum), with ovary cut +across; the inner (ventral) face turned toward the eye: the ovules seem +to be borne on the ventral suture, answering to leaf-margins: the stigma +above seen also to answer to leaf-margins.</p> +<p><span class="smcap">Fig.</span> 325. Pod or simple pistil of Caltha or +Marsh-Marigold, which has opened, and shed its seeds.</p> +</div> + +<p>307. So a simple pistil should have a one-celled ovary, only one line of +attachment for the ovules, a single style, and a single stigma. Certain +variations from this normal condition which sometimes occur do not +invalidate this morphological conception. For instance, the stigma may +become two-lobed or two-ridged, because it consists of two leaf-margins, +as Fig. <a href="#Fig323">324</a> shows; it may become 2-locellate by the turning or growing +inward of one of the sutures, so as to divide the cavity.</p> + +<p>308. There are two or three terms which primarily relate to the parts of +a simple pistil or carpel, and are thence carried on to the compound +pistil, viz.:—</p> + +<p><span class="smcap">Ventral Suture</span>, the line which answers to the united margins of the +carpel-leaf, therefore naturally called a suture or seam, and the +ventral or inner one, because in the circle of carpel-leaves it looks +inward or to the centre of the flower.</p> + +<p><span class="smcap">Dorsal Suture</span> is the line down the back of the carpel, answering to <span class="pagenum"><a name="Page_107" id="Page_107">[Pg 107]</a></span>the +midrib of the leaf,—not a seam therefore; but at maturity many fruits, +such as pea-pods, open by this dorsal as well as by the ventral line.</p> + +<p><span class="smcap">Placenta</span>, a name given to the surface, whatever it be, which bears the +ovules and seeds. The name may be needless when the ovules grow directly +on the ventral suture, or from its top or bottom; but when there are +many ovules there is usually some expansion of an ovule-bearing or +seed-bearing surface; as is seen in our Mandrake or Podophyllum, Fig. +<a href="#Fig326">326</a>.</p> + +<div class="figcenter"> +<a name="Fig326" id="Fig326"></a> +<img src="images/fig326_330.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 326. Simple pistil of Podophyllum, cut across, +showing ovules borne on placenta.</p> +<p><span class="smcap">Fig.</span> 327. Pistil of a Saxifrage, of two simple carpels or +pistil-leaves, united at the base only, cut across both above and +below.</p> +<p><span class="smcap">Fig.</span> 328. Compound 3-carpellary pistil of common St. +John's-wort, cut across: the three styles separate.</p> +<p><span class="smcap">Fig.</span> 329. The same of shrubby St. John's-wort; the three +styles as well as ovaries here united into one.</p> +<p><span class="smcap">Fig.</span> 330. Compound 3-carpellary pistil of Tradescantia or +Spiderwort; the three stigmas as well as styles and ovary completely +coalescent into one.</p> +</div> + +<p><a name="UNIT_309" id="UNIT_309">309.</a> <b>A Compound Pistil</b> is a combination of two, three, or a greater +number of pistil-leaves or carpels in a circle, united into one body, at +least by their ovaries. The annexed figures should make it clear. A +series of Saxifrages might be selected the gynœcium of which would +show every gradation between two simple pistils, or separate carpels, +and their complete coalescence into one compound and two-celled ovary. +Even when the constituent styles and stigmas are completely coalescent +into one, the nature of the combination is usually revealed by some +external lines or grooves, or (as in Fig. <a href="#Fig326">328-330</a>) by the internal +partitions, or the number of the placentæ. The simplest case of compound +pistil is that</p> + +<p><a name="UNIT_310" id="UNIT_310">310.</a> <b>With two or more Cells and Axile Placentæ</b>, namely, with as many +cells as there are carpels, that have united to compose the organ. <span class="pagenum"><a name="Page_108" id="Page_108">[Pg 108]</a></span>Such +a pistil is just what would be formed if the simple pistils (two, three, +or five in a circle, as the case may be), like those of a Pæony or +Stonecrop (Fig. <a href="#Fig224">224, 225</a>), pressed together in the centre of the flower, +were to cohere by their contiguous parts. In such a case the placentæ +are naturally <i>axile</i>, or all brought together in the axis or centre; +and the ovary has as many <span class="smcap">Dissepiments</span>, or internal <i>Partitions</i>, as +there are carpels in its composition. For these are the contiguous and +coalescent walls or sides of the component carpels. When such pistils +ripen into pods, they often separate along these lines into their +elementary carpels.</p> + +<div class="figcenter"> +<a name="Fig331" id="Fig331"></a> +<img src="images/fig331_332.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 331, 332. Pistil of a Sandwort, with vertical and +transverse section of the ovary: free central placenta.</p> +</div> + +<p><a name="UNIT_311" id="UNIT_311">311.</a> <b>One-celled, with free Central Placenta.</b> The commoner case is that +of Purslane (Fig. <a href="#Fig270">272</a>) and of the Pink and Chickweed families (Fig. <a href="#Fig331">331, +332</a>). This is explained by supposing that the partitions (such as those +of Fig. <a href="#Fig326">329</a>) have early vanished or have been suppressed. Indeed, traces +of them may often be detected in Pinks. On the other hand, it is equally +supposable that in the Primula family the free central is derived from +parietal placentation by the carpels bearing ovules only at base, and +forming a consolidated common placenta in the axis. Mitella and Dionæa +help out this conception.</p> + +<div class="figcenter"> +<a name="Fig333" id="Fig333"></a> +<img src="images/fig333.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 333. Plan of a one-celled ovary of three +carpel-leaves, with parietal placentæ, cut across below, where it is +complete; the upper part showing the top of the three leaves it is +composed of, approaching, but not united.</p> +</div> + +<div class="figcenter"> +<a name="Fig334" id="Fig334"></a> +<img src="images/fig334_336.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 334. Cross section of the ovary of Frost weed +(Helianthemum), with three parietal placentæ, bearing ovules.</p> +<p><span class="smcap">Fig.</span> 335. Cross section of an ovary of Hypericum +graveolens, the three large placentæ meeting in the centre, so as to +form a three-celled ovary. 336. Same in fruit, the placentæ now separate +and rounded.</p> +</div> + +<p><a name="UNIT_312" id="UNIT_312">312.</a> <b>One-celled, with Parietal Placentæ.</b> In this not uncommon case it is +conceived that the two or three or more carpel-leaves of such a compound +pistil coalesce by their adjacent edges, just as sepal-leaves do to form +a gamosepalous calyx, or petals to form a gamopetalous corolla, and as +is shown in the diagram, Fig. <a href="#Fig333">333</a>, and in an actual cross-section, Fig. +<a href="#Fig334">334</a>. Here each carpel is an open leaf, or with some introflexion, +bearing ovules along its margins; and each placenta consists of the +contiguous <span class="pagenum"><a name="Page_109" id="Page_109">[Pg 109]</a></span>margins of two pistil-leaves grown together. There is every +gradation between this and the three-celled ovary with the placentæ in +the axis, even in the same genus, sometimes even in different stages in +the same pistil (Fig. <a href="#Fig334">335, 336</a>).</p> + + +<h3><a name="Subsect_X_2" id="Subsect_X_2">§ 2.</a> GYMNOSPERMOUS GYNŒCIUM.</h3> + +<p><a name="UNIT_313" id="UNIT_313">313.</a> The ordinary pistil has a closed ovary, and accordingly the pollen +can act upon the contained ovules only indirectly, through the stigma. +This is expressed in a term of Greek derivation, viz.:—</p> + +<p><i>Angiospermous</i>, meaning that the seeds are borne in a sac or closed +vessel. The counterpart term is</p> + +<p><i>Gymnospermous</i>, meaning naked-seeded. This kind of pistil, or +gynœcium, the simplest of all, yet the most peculiar, characterizes +the Pine family and its relatives.</p> + +<div class="figcenter"> +<a name="Fig337" id="Fig337"></a> +<img src="images/fig337.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 337. A pistil, that is, a scale of the cone, of a +Larch, at the time of flowering; inside view, showing its pair of naked +ovules.</p> +</div> + +<div class="figcenter"> +<a name="Fig338" id="Fig338"></a> +<img src="images/fig338_339.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 338. Branchlet of the American Arbor-Vitæ, +considerably larger than in nature, terminated by its pistillate +flowers, each consisting of a single scale (an open pistil), together +forming a small cone.</p> +<p><span class="smcap">Fig.</span> 339. One of the scales or carpels of the last, +removed and more enlarged, the inside exposed to view, showing a pair of +ovules on its base.</p> +</div> + +<p>314. While the ordinary simple pistil is conceived by the botanist to be +a leaf rolled together into a closed pod (<a href="#UNIT_306">306</a>), those of the Pine, Larch +(Fig. <a href="#Fig337">337</a>), Cedar, and Arbor-Vitæ (Fig. <a href="#Fig338">338, 339</a>) are open leaves, in +the form of scales, each bearing two or more ovules on the inner face, +next the base. At the time of blossoming, these pistil-leaves of the +young cone diverge, and the pollen, so abundantly shed from the +staminate blossoms, falls directly upon the exposed ovules. Afterward +the scales close over each other until the seeds are ripe. Then they +separate that the seeds may be shed. As the pollen acts directly on the +ovules, such pistil (or organ acting as pistil) has no stigma.</p> + +<p>315. In the Yew, and in Torreya and Gingko, the gynœcium is reduced +to extremest simplicity, that is, to a naked ovule, without any visible +carpel.</p> + +<p>316. In Cycas the large naked ovules are borne on the margins or lobes +of an obvious open leaf. All <span class="smcap">Gymnospermous</span> plants have other +peculiarities, also distinguishing them, as a class, from <span class="smcap">Angiospermous</span> +plants.</p> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_110" id="Page_110">[Pg 110]</a></span></p> +<h2><a name="Section_XI" id="Section_XI"></a><span class="smcap">Section XI.</span> OVULES.</h2> + + +<p><a name="UNIT_317" id="UNIT_317">317.</a> <b>Ovule</b> (from the Latin, meaning a little egg) is the technical name +of that which in the flower answers to and becomes the seed.</p> + +<div class="figcenter"> +<a name="Fig340" id="Fig340"></a> +<img src="images/fig340.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 340. A cluster of ovules, pendulous on their +funicles.</p> +</div> + +<p>318. Ovules are <i>naked</i> in gymnospermous plants (as just described), in +all others they are enclosed in the ovary. They may be produced along +the whole length of the cell or cells of the ovary, and then they are +apt to be numerous, or only from some part of it, generally the top or +the bottom. In this case they are usually few or single (<i>solitary</i>, as +in Fig. <a href="#Fig341">341-343</a>). They may be <i>sessile</i>, i. e. without stalk, or they +may be attached by a distinct stalk, the <span class="smcap">Funicle</span> or <span class="smcap">Funiculus</span> (Fig. +<a href="#Fig340">340</a>).</p> + +<div class="figcenter"> +<a name="Fig341" id="Fig341"></a> +<img src="images/fig341_343.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 341. Section of the ovary of a Buttercup, +lengthwise, showing its ascending ovule.</p> +<p><span class="smcap">Fig.</span> 342. Section of the ovary of Buckwheat, showing the +erect ovule.</p> +<p><span class="smcap">Fig.</span> 343. Section of the ovary of Anemone, showing its +suspended ovule.</p> +</div> + +<p>319. Considered as to then position and direction in the ovary, they are</p> + +<p><i>Horizontal</i>, when they are neither turned upward nor downward, as in +Podophyllum (Fig. <a href="#Fig326">326</a>),</p> + +<p><i>Ascending</i>, when rising obliquely upwards, usually from the side of the +cell, not from its very base, as in the Buttercup (Fig. <a href="#Fig341">341</a>), and the +Purslane (Fig. <a href="#Fig270">272</a>),</p> + +<p><i>Erect</i>, when rising upright from the very base of the cell, as in the +Buckwheat (Fig. <a href="#Fig341">342</a>),</p> + +<p><i>Pendulous</i>, when hanging from the side or from near the top, as in the +Flax (Fig. <a href="#Fig270">270</a>), and</p> + +<p><i>Suspended</i>, when hanging perpendicularly from the very summit of the +cell, as in the Anemone (Fig. <a href="#Fig341">343</a>). All these terms equally apply to +seeds.</p> + +<p><a name="UNIT_320" id="UNIT_320">320.</a> In structure an ovule is a pulpy mass of tissue, usually with one +or two coats or coverings. The following parts are to be noted, viz.—</p> + +<p><span class="smcap">Kernel</span> or <span class="smcap">Nucleus</span>, the body of the ovule. In the Mistletoe and some +related plants, there is only this nucleus, the coats being wanting.</p> + +<p><span class="smcap">Teguments</span>, or coats, sometimes only one, more commonly two. When two, +one has been called <span class="smcap">Primine</span>, the other <span class="smcap">Secundine</span>. It will serve all +purposes to call them simply outer and inner ovule coats.</p> + +<p><span class="smcap">Orifice</span>, or <span class="smcap">Foramen</span>, an opening through the coats at the organic apex of +the ovule. In the seed it is <i>Micropyle</i>.</p> + +<p><span class="smcap">Chalaza</span>, the place where the coats and the kernel of the ovule blend.</p> + +<p><span class="smcap">Hilum</span>, the place of junction of the funiculus with the body of the +ovule.</p> + +<p><span class="pagenum"><a name="Page_111" id="Page_111">[Pg 111]</a></span></p><div class="figcenter"> +<a name="Fig344" id="Fig344"></a> +<img src="images/fig344_347.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 344. Orthotropous ovule of Buckwheat: <i>c</i>, hilum +and chalaza; <i>f</i>, orifice.</p> +<p><span class="smcap">Fig.</span> 345. Campylotropous ovule of a Chickweed: <i>c</i>, hilum +and chalaza; <i>f</i>, orifice.</p> +<p><span class="smcap">Fig.</span> 346. Amphitropous ovule of Mallow: <i>f</i>, orifice; +<i>h</i>, hilum; <i>r</i>, rhaphe; <i>c</i>, chalaza.</p> +<p><span class="smcap">Fig.</span> 347. Anatropous ovule of a Violet, the parts +lettered as in the last.</p> +</div> + +<p><a name="UNIT_321" id="UNIT_321">321.</a> <b>The Kinds of Ovules.</b> The ovules in their growth develop in three or +four different ways and thereby are distinguished into</p> + +<p><i>Orthotropous</i> or <i>Straight</i>, those which develop without curving or +turning, as in Fig. <a href="#Fig344">344</a>. The chalaza is at the insertion or base, the +foramen or orifice is at the apex. This is the simplest, but the least +common kind of ovule.</p> + +<p><i>Campylotropous</i> or <i>Incurved</i>, in which, by the greater growth of one +side, the ovule curves into a kidney-shaped outline, so bringing the +orifice down close to the base or chalaza; as in Fig. <a href="#Fig344">345</a>.</p> + +<p><i>Amphitropous</i> or <i>Half Inverted</i>, Fig. <a href="#Fig344">346</a>. Here the forming ovule, +instead of curving perceptibly, keeps its axis nearly straight, and, as +it grows, turns round upon its base so far as to become transverse to +its funiculus, and adnate to its upper part for some distance. Therefore +in this case the attachment of the funiculus or stalk is about the +middle, the chalaza is at one end, the orifice at the other.</p> + +<div class="figcenter"> +<a name="Fig348" id="Fig348"></a> +<img src="images/fig348_354.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 348-350. Three early stages in the growth of ovule +of a Magnolia, showing the forming outer and inner coats which even in +the later figure have not yet completely enclosed the nucleus; 351, +further advanced, and 352, completely anatropous ovule.</p> +<p><span class="smcap">Fig.</span> 353. Longitudinal section, and 354, transverse +section of 352.</p> +</div> + +<div class="figcenter"> +<a name="Fig355" id="Fig355"></a> +<img src="images/fig355.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 355. Same as 353, enlarged showing the parts in +section: <i>a</i>, outer coat; <i>b</i>, inner coat; <i>c</i>, nucleus; <i>d</i>, rhaphe.</p> +</div> + +<p><i>Anatropous</i> or <i>Inverted</i>, as in Fig. <a href="#Fig344">347</a>, the commonest kind, so +called because in its growth it has as it were turned over upon its +stalk, to which it has continued adnate. The organic base, or chalaza, +thus becomes the apparent summit, and the <span class="pagenum"><a name="Page_112" id="Page_112">[Pg 112]</a></span>orifice is at the base, by +the side of the hilum or place of attachment. The adnate portion of the +funiculus, which appears as a ridge or cord extending from the hilum to +the chalaza, and which distinguishes this kind of ovule, is called the +<span class="smcap">Rhaphe</span>. The amphitropous ovule (Fig. <a href="#Fig344">346</a>) has a short or incomplete +rhaphe.</p> + +<p>322. Fig. <a href="#Fig348">348-352</a> show the stages through which an ovule becomes +anatropous in the course of its growth. The annexed two figures are +sections of such an ovule at maturity; and Fig. <a href="#Fig355">355</a> is Fig. <a href="#Fig348">353</a> +enlarged, with the parts lettered.</p> + + + +<hr class="chap" /> +<h2><a name="Section_XII" id="Section_XII"></a><span class="smcap">Section XII.</span> MODIFICATIONS OF THE RECEPTACLE.</h2> + + +<div class="figcenter"> +<a name="Fig356" id="Fig356"></a> +<img src="images/fig356_359.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 356. Longitudinal section of flower of Silene +Pennsylvanica, showing stipe between calyx and corolla.</p> +<p><span class="smcap">Fig.</span> 357. Flower of a Cleome of the section Gynandropsis, +showing broadened receptacle to bear petals, lengthened stipe below the +stamens, and another between these and pistil.</p> +<p><span class="smcap">Fig.</span> 358. Pistil of Geranium or Cranesbill.</p> +<p><span class="smcap">Fig.</span> 359. The same, ripe, with the five carpels splitting +away from the long beak (carpophore), and hanging from its top by their +recurving styles.</p> +</div> + +<p><a name="UNIT_323" id="UNIT_323">323.</a> <b>The Torus</b> or Receptacle of the flower (<a href="#UNIT_237">237</a>, Fig. <a href="#Fig222">223</a>) is the +portion which belongs to the stem or axis. In all preceding +illustrations it is small and short. But it sometimes lengthens, +sometimes thickens or variously enlarges, and takes on various forms. +Some of these have received special names, very few of which are in +common use. A lengthened portion of the receptacle is called</p> + +<p>A <span class="smcap">Stipe</span>. This name, which means simply a trunk or stalk, is used in +botany for various stalks, even for the leaf-stalk in Ferns. It is also +applied to the stalk or petiole of a carpel, in the rare cases when +there is any, as in <span class="pagenum"><a name="Page_113" id="Page_113">[Pg 113]</a></span>Goldthread. Then it is technically distinguished as +a <span class="smcap">Thecaphore</span>. When there is a stalk, or lengthened internode of +receptacle, directly under a compound pistil, as in Stanleya and some +other Cruciferæ, it is called a <span class="smcap">Gynophore</span>. When the stalk is developed +below the stamens, as in most species of Silene (Fig. <a href="#Fig356">356</a>), it has been +called an <span class="smcap">Anthophore</span> or <span class="smcap">Gonophore</span>. In Fig. <a href="#Fig356">357</a> the torus is dilated +above the calyx where it bears the petals, then there is a long +internode (gonophore) between it and the stamens; then a shorter one +(gynophore) between these and the pistil.</p> + +<p><a name="UNIT_324" id="UNIT_324">324.</a> <b>A Carpophore</b> is a prolongation of receptacle or axis between the +carpels and bearing them. Umbelliferous plants and Geranium (Fig. <a href="#Fig356">358, +359</a>) afford characteristic examples.</p> + +<div class="figcenter"> +<a name="Fig360" id="Fig360"></a> +<img src="images/fig360_362.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 360. Longitudinal section of a young strawberry, +enlarged.</p> +<p><span class="smcap">Fig.</span> 361. Similar section of a young Rose-hip.</p> +<p><span class="smcap">Fig.</span> 362. Enlarged and top-shaped receptacle of +Nelumbium, at maturity.</p> +</div> + +<p>325. Flowers with very numerous simple pistils generally have the +receptacle enlarged so as to give them room; sometimes becoming broad +and flat, as in the Flowering Raspberry, sometimes elongated, as in the +Blackberry, the Magnolia, etc. It is the receptacle in the Strawberry +(Fig. <a href="#Fig360">360</a>), much enlarged and pulpy when ripe, which forms the eatable +part of the fruit, and bears the small seed-like pistils on its surface. +In the Rose (Fig. <a href="#Fig360">361</a>), instead of being convex or conical, the +receptacle is deeply concave, or urn-shaped. Indeed, a Rose-hip may be +likened to a strawberry turned inside out, like the finger of a glove +reversed, and the whole covered by the adherent tube of the calyx. The +calyx remains beneath in the strawberry.</p> + +<p>326. In Nelumbium, of the Water-Lily family, the singular and greatly +enlarged receptacle is shaped like a top, and bears the small pistils +immersed in separate cavities of its flat upper surface (Fig. <a href="#Fig360">362</a>).</p> + +<div class="figcenter"> +<a name="Fig363" id="Fig363"></a> +<img src="images/fig363.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 363. Hypogynous disk in Orange. +</div> + +<p><a name="UNIT_327" id="UNIT_327">327.</a> <b>A Disk</b> is an enlarged low receptacle or an outgrowth from it, +<i>hypogynous</i> when underneath the pistil, as in Rue and the Orange (Fig. +<a href="#Fig363">363</a>), and <i>perigynous</i> when adnate to calyx-tube (as in Buckthorn, Fig. +<a href="#Fig364">364, 365</a>), and Cherry (Fig. <a href="#Fig270">271</a>), or <span class="pagenum"><a name="Page_114" id="Page_114">[Pg 114]</a></span>to both calyx-tube and ovary, as +in Hawthorn (Fig. <a href="#Fig273">273</a>). A flattened hypogynous disk, underlying the +ovary or ovaries, and from which they fall away at maturity, is +sometimes called a <span class="smcap">Gynobase</span>, as in the Rue family. In some Borragineous +flowers, such as Houndstongue, the gynobase runs up in the centre +between the carpels into a carpophore. The so-called <i>epigynous</i> disk +(or <span class="smcap">Stylopodium</span>) crowning the summit of the ovary in flowers of +Umbelliferæ, etc., cannot be said to belong to the receptacle.</p> + +<div class="figcenter"> +<a name="Fig364" id="Fig364"></a> +<img src="images/fig364_365.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 364. Flower of a Buckthorn showing a conspicuous +perigynous disk.</p> +<p><span class="smcap">Fig.</span> 365. Vertical section of same flower.</p> +</div> + + + +<hr class="chap" /> +<h2><a name="Section_XIII" id="Section_XIII"></a><span class="smcap">Section XIII.</span> FERTILIZATION.</h2> + + +<p>328. The end of the flower is attained when the ovules become seeds. A +flower remains for a certain time (longer or shorter according to the +species) in <i>anthesis</i>, that is, in the proper state for the fulfilment +of this end. During anthesis, the ovules have to be fertilized by the +pollen; or at least some pollen has to reach the stigma, or in +gymnospermy the ovule itself, and to set up the peculiar growth upon its +moist and permeable tissue, which has for result the production of an +embryo in the ovules. By this the ovules are said to be <i>fertilized</i>. +The first step is <i>pollination</i>, or, so to say, the sowing of the proper +pollen upon the stigma, where it is to germinate.</p> + + +<h3><a name="Subsect_XIII_1" id="Subsect_XIII_1">§ 1.</a> ADAPTATIONS FOR POLLINATION OF THE STIGMA.</h3> + +<p>329. These various and ever-interesting adaptations and processes are +illustrated in the "Botanical Text Book, Structural Botany," chap. VI. +sect. iv., also in a brief and simple way in "Botany for Young People, +How Plants Behave." So mere outlines only are given here.</p> + +<p>330. Sometimes the application of pollen to the stigma is left to +chance, as in diœcious wind-fertilized flowers; sometimes it is +rendered very sure, as in flowers that are fertilized in the bud; +sometimes the pollen is prevented from reaching the stigma of the same +flower, although placed very near to it, but then there are always +arrangements for its transference to the stigma of some other blossom of +the kind. It is among these last that the most exquisite adaptations are +met with.</p> + +<p><a name="UNIT_331" id="UNIT_331">331.</a> Accordingly, some flowers are particularly adapted to close or +self-fertilization; others to cross fertilization; some for either, +according to circumstances.</p> + +<p><span class="pagenum"><a name="Page_115" id="Page_115">[Pg 115]</a></span></p><p><i>Close Fertilization</i> occurs when the pollen reaches and acts upon a +stigma of the very same flower (this is also called self-fertilization), +or, less closely, upon other blossoms of the same cluster or the same +individual plant.</p> + +<p><i>Cross Fertilization</i> occurs when ovules are fertilized by pollen of +other individuals of the same species.</p> + +<p><i>Hybridization</i> occurs when ovules are fertilized by pollen of some +other (necessarily some nearly related) species.</p> + +<p><a name="UNIT_332" id="UNIT_332">332.</a> <b>Close Fertilization</b> would seem to be the natural result in ordinary +hermaphrodite flowers; but it is by no means so in all of them. More +commonly the arrangements are such that it takes place only after some +opportunity for cross fertilization has been afforded. But close +fertilization is inevitable in what are called</p> + +<p><i>Cleistogamous Flowers</i>, that is, in those which are fertilized in the +flower-bud, while still unopened. Most flowers of this kind, indeed, +never open at all; but the closed floral coverings are forced off by the +growth of the precociously fertilized pistil. Common examples of this +are found in the earlier blossoms of Specularia perfoliata, in the later +ones of most Violets, especially the stemless species, in our wild Jewel +weeds or Impatiens, in the subterranean shoots of Amphicarpæa. Every +plant which produces these cleistogamous or bud-fertilized flowers bears +also more conspicuous and open flowers, usually of bright colors. The +latter very commonly fail to set seed, but the former are prolific.</p> + +<p><a name="UNIT_333" id="UNIT_333">333.</a> <b>Cross Fertilization</b> is naturally provided for in diœcious plants +(<a href="#UNIT_249">249</a>), is much favored in monœcious plants (<a href="#UNIT_249">249</a>), and hardly less so +in dichogamous and in heterogonous flowers (<a href="#UNIT_338">338</a>). Cross fertilization +depends upon the transportation of pollen; and the two principal agents +of conveyance are winds and insects. Most flowers are in their whole +structure adapted either to the one or to the other.</p> + +<p><a name="UNIT_334" id="UNIT_334">334.</a> <b>Wind-fertilizable or Anemophilous</b> flowers are more commonly +diœcious or monœcious, as in Pines and all coniferous trees, Oaks, +and Birches, and Sedges; yet sometimes hermaphrodite, as in Plantains +and most Grasses; they produce a superabundance of very light pollen, +adapted to be wind-borne; and they offer neither nectar to feed winged +insects, nor fragrance nor bright colors to attract them.</p> + +<p><a name="UNIT_335" id="UNIT_335">335.</a> <b>Insect-fertilizable or Entomophilous</b> flowers are those which are +sought by insects, for pollen or for nectar, or for both. Through their +visits pollen is conveyed from one flower and from one plant to another. +Insects are attracted to such blossoms by their bright colors, or their +fragrance, or by the nectar (the material of honey) there provided for +them. While supplying their own needs, they carry pollen from anthers to +stigmas and from plant to plant, thus bringing about a certain amount of +cross fertilization. Willows and some other diœcious flowers are so +fertilized, chiefly by bees. But most insect-visited flowers have the +stamens and pistils associated either in the same or in contiguous +blossoms. Even when in the same blossom, anthers and stigmas are very +commonly so situated <span class="pagenum"><a name="Page_116" id="Page_116">[Pg 116]</a></span>that under insect-visitation, some pollen is more +likely to be deposited upon other than upon own stigmas, so giving a +chance for cross as well as for close fertilization. On the other hand, +numerous flowers, of very various kinds, have their parts so arranged +that they must almost necessarily be cross-fertilized or be barren, and +are therefore dependent upon the aid of insects. This aid is secured by +different exquisite adaptations and contrivances, which would need a +volume for full illustration. Indeed, there is a good number of volumes +devoted to this subject.<a name="FNanchor_1_2" id="FNanchor_1_2"></a><a href="#Footnote_1_2" class="fnanchor">[1]</a></p> + +<p>336. Some of the adaptations which favor or ensure cross fertilization +are peculiar to the particular kind of blossom. Orchids, Milkweeds, +Kalmia, Iris, and papilionaceous flowers each have their own special +contrivances, quite different for each.</p> + +<p>337. Irregular flowers (<a href="#UNIT_253">253</a>) and especially irregular corollas are +usually adaptations to insect-visitation. So are all <i>Nectaries</i>, +whether hollow spurs, sacs, or other concavities in which nectar is +secreted, and all <i>nectariferous glands</i>.</p> + +<p><a name="UNIT_338" id="UNIT_338">338.</a> Moreover, there are two arrangements for cross fertilization common +to hermaphrodite flowers in various different families of plants, which +have received special names, <i>Dichogamy</i> and <i>Heterogony</i>.</p> + +<p><a name="UNIT_339" id="UNIT_339">339.</a> <b>Dichogamy</b> is the commoner case. Flowers are <i>dichogamous</i> when the +anthers discharge their pollen either before or after the stigmas of +that flower are in a condition to receive it. Such flowers are</p> + +<p><i>Proterandrous</i>, when the anthers are earlier than the stigmas, as in +Gentians, Campanula, Epilobium, etc.</p> + +<p><i>Proterogynous</i>, when the stigmas are mature and moistened for the +reception of pollen, before the anthers of that blossom are ready to +supply it, and are withered before that pollen can be supplied. +Plantains or Ribworts (mostly wind-fertilized) are strikingly +proterogynous: so is Amorpha, our Papaws, Scrophularia, and in a less +degree the blossom of Pears, Hawthorns, and Horse-chestnut.</p> + +<p>340. In Sabbatia, the large-flowered species of Epilobium, and +strikingly in Clerodendron, the dichogamy is supplemented and perfected +by movements of the stamens and style, one or both, adjusted to make +sure of cross fertilization.</p> + +<p><a name="UNIT_341" id="UNIT_341">341.</a> <b>Heterogony.</b> This is the case in which hermaphrodite and fertile +flowers of two sorts are produced on different individuals of the same +species; one sort having higher anthers and lower stigmas, the other +having higher stigmas and lower anthers. Thus reciprocally disposed, a +visiting insect carries pollen from the high anthers of the one to the +high stigma of the other, and from the low anthers of the one to the low +stigma of the other. These plants are practically as if diœcious, +with the advantage that <span class="pagenum"><a name="Page_117" id="Page_117">[Pg 117]</a></span>both kinds are fruitful. Houstonia and +Mitchella, or Partridge-berry, are excellent and familiar examples. +These are cases of</p> + +<p><i>Heterogone Dimorphism</i>, the relative lengths being only short and long +reciprocally.</p> + +<p><i>Heterogone Trimorphism</i>, in which there is a mid-length as well as a +long and a short set of stamens and style; occurs in Lythrum Salicaria +and some species of Oxalis.</p> + +<p>342. There must be some essential advantage in cross fertilization or +cross breeding. Otherwise all these various, elaborate, and exquisitely +adjusted adaptations would be aimless. Doubtless the advantage is the +same as that which is realized in all the higher animals by the +distinction of sexes.</p> + + +<h3><a name="Subsect_XIII_2" id="Subsect_XIII_2">§ 2.</a> ACTION OF POLLEN, AND FORMATION OF THE EMBRYO.</h3> + +<p><a name="UNIT_343" id="UNIT_343">343.</a> <b>Pollen-growth.</b> A grain of pollen may be justly likened to one of +the simple bodies (<i>spores</i>) which answer for seeds in Cryptogamous +plants. Like one of these, it is capable of germination. When deposited +upon the moist surface of the stigma (or in some cases even when at a +certain distance) it grows from some point, its living inner coat +breaking through the inert outer coat, and protruding in the form of a +delicate tube. This as it lengthens penetrates the loose tissue of the +stigma and of a loose conducting tissue in the style, feeds upon the +nourishing liquid matter there provided, reaches the cavity of the +ovary, enters the orifice of an ovule, and attaches its extremity to a +sac, or the lining of a definite cavity, in the ovule, called the +<i>Embryo-Sac</i>.</p> + +<p><a name="UNIT_344" id="UNIT_344">344.</a> <b>Origination of the Embryo.</b> A globule of living matter in the +embryo-sac is formed, and is in some way placed in close proximity to +the apex of the pollen tube; it probably absorbs the contents of the +latter; it then sets up a special growth, and the <i>Embryo</i> (<a href="#UNIT_8">8-10</a>) or +rudimentary plantlet in the seed is the result.</p> + + +<div class="footnotes"><h3>FOOTNOTES:</h3> + +<div class="footnote"><p><a name="Footnote_1_2" id="Footnote_1_2"></a><a href="#FNanchor_1_2"><span class="label">[1]</span></a> Beginning with one by C. C. Sprengel in 1793, and again in +our day with Darwin, "On the Various Contrivances by which Orchids are +fertilized by Insects," and in succeeding works.</p></div> +</div> + + +<hr class="chap" /> +<h2><a name="Section_XIV" id="Section_XIV"></a><span class="smcap">Section XIV.</span> THE FRUIT.</h2> + + +<p><a name="UNIT_345" id="UNIT_345">345.</a> <b>Its Nature.</b> The ovary matures into the Fruit. In the strictest +sense the fruit is the seed-vessel, technically named the <span class="smcap">Pericarp</span>. But +practically it may include other parts organically connected with the +pericarp. Especially the calyx, or a part of it, is often incorporated +with the ovary, so as to be undistinguishably a portion of the pericarp, +and it even forms along with the receptacle the whole bulk of such +edible fruits as apples and pears. The receptacle is an obvious part in +blackberries, and is the whole edible portion in the strawberry.</p> + +<p>346. Also a cluster of distinct carpels may, in ripening, be +consolidated or compacted, so as practically to be taken for one fruit. +Such are raspberries, <span class="pagenum"><a name="Page_118" id="Page_118">[Pg 118]</a></span>blackberries, the Magnolia fruit, etc. Moreover, +the ripened product of many flowers may be compacted or grown together +so as to form a single compound fruit.</p> + +<p><a name="UNIT_347" id="UNIT_347">347.</a> <b>Its kinds</b> have therefore to be distinguished. Also various names of +common use in descriptive botany have to be mentioned and defined.</p> + +<p>348. In respect to composition, accordingly, fruits may be classified +into</p> + +<p><i>Simple</i>, those which result from the ripening of a single pistil, and +consist only of the matured ovary, either by itself, as in a cherry, or +with calyx-tube completely incorporated with it, as in a gooseberry or +cranberry.</p> + +<p><i>Aggregate</i>, when a cluster of carpels of the same flower are crowded +into a mass; as in raspberries and blackberries.</p> + +<div class="figcenter"> +<a name="Fig366" id="Fig366"></a> +<img src="images/fig366_367.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 366. Forming fruit (capsule) of Gaultheria, with +calyx thickening around its base. 367. Section of same mature, the +berry-like calyx nearly enclosing the capsule.</p> +</div> + +<div class="figcenter"> +<a name="Fig368" id="Fig368"></a> +<img src="images/fig368_370.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 368. Section of a part of a strawberry. Compare with +Fig. <a href="#Fig360">360</a>.</p> +<p><span class="smcap">Fig.</span> 369. Similar section of part of a blackberry. 370. +One of its component simple fruits (drupe) in section, showing the pulp, +stone, and contained seed; more enlarged. Compare with Fig. <a href="#Fig375">375</a>.</p> +</div> + +<p><i>Accessory</i> or <i>Anthocarpous</i>, when the surroundings or supports of the +pistil make up a part of the mass; as does the loose calyx changed into +a fleshy and berry-like envelope of our Wintergreen (Gaultheria, Fig. +<a href="#Fig366">366, 367</a>) and Buffalo-berry, which are otherwise simple fruits. In an +aggregate fruit such as the strawberry the great mass is receptacle +(Fig. <a href="#Fig360">360</a>, <a href="#Fig368">368</a>); and in the blackberry (Fig. <a href="#Fig368">369</a>) the juicy receptacle +forms the central part of the savory mass.</p> + +<p><i>Multiple</i> or <i>Collective</i>, when formed from several flowers +consolidated into one mass, of which the common receptacle or axis of +inflorescence, the floral envelopes, and even the bracts, etc., make a +part. A mulberry (Fig. <a href="#Fig405">408</a>, which superficially much resembles a +blackberry) is of this multiple sort. A pine-apple is another example.</p> + +<p>349. In respect to texture or consistence, fruits may be distinguished +into three kinds, viz.—</p> + +<p><i>Fleshy Fruits</i>, those which are more or less soft and juicy throughout;</p> + +<p><span class="pagenum"><a name="Page_119" id="Page_119">[Pg 119]</a></span></p><p><i>Stone Fruits</i>, or <i>Drupaceous</i>, the outer part fleshy like a berry, +the inner hard or stony, like a nut; and</p> + +<p><i>Dry Fruits</i>, those which have no flesh or pulp.</p> + +<p>350. In reference to the way of disseminating the contained seed, fruits +are said to be</p> + +<p><i>Indehiscent</i> when they do not open at maturity. Fleshy fruits and stone +fruits are of course indehiscent. The seed becomes free only through +decay or by being fed upon by animals. Those which escape digestion are +thus disseminated by the latter. Of dry fruits many are indehiscent; and +these are variously arranged to be transported by animals. Some burst +irregularly; many are</p> + +<p><i>Dehiscent</i>, that is, they split open regularly along certain lines, and +discharge the seeds. A dehiscent fruit almost always contains many or +several seeds, or at least more than one seed.</p> + +<div class="figcenter"> +<a name="Fig371" id="Fig371"></a> +<img src="images/fig371_374.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 371. Leafy shoot and berry (cut across) of the +larger Cranberry, Vaccinium macrocarpon.</p> +<p><span class="smcap">Fig.</span> 372, 373. Pepo of Gourd, in section. 373. One carpel +of same in diagram.</p> +<p><span class="smcap">Fig.</span> 374. Longitudinal and transverse sections of a pear +(pome).</p> +</div> + +<p>351. The principal kinds of fruit which have received substantive names +and are of common use in descriptive botany are the following. Of fleshy +fruits the leading kind is</p> + +<p><a name="UNIT_352" id="UNIT_352">352.</a> <b>The Berry</b>, such as the gooseberry and currant, the blueberry and +cranberry (Fig. <a href="#Fig371">371</a>), the tomato, and the grape. Here the whole flesh is +soft throughout. The orange is a berry with a leathery rind.</p> + +<p><a name="UNIT_353" id="UNIT_353">353.</a> <b>The Pepo</b>, or <i>Gourd-fruit</i>, is a hard-rinded berry, belonging to +the Gourd family, such as the pumpkin, squash, cucumber, and melon, Fig. +<a href="#Fig371">372, 373</a>.</p> + +<p><a name="UNIT_354" id="UNIT_354">354.</a> <b>The Pome</b> is a name applied to the apple, pear (Fig. <a href="#Fig371">374</a>), and +quince; fleshy fruits, like a berry, but the principal thickness is +calyx, only <span class="pagenum"><a name="Page_120" id="Page_120">[Pg 120]</a></span>the papery pods arranged like a star in the core really +belonging to the carpels. The fruit of the Hawthorn is a drupaceous +pome, something between pome and drupe.</p> + +<p>355. Of fruits which are externally fleshy and internally hard the +leading kind is</p> + +<p><a name="UNIT_356" id="UNIT_356">356.</a> <b>The Drupe</b>, or <i>Stone-fruit</i>; of which the cherry, plum, and peach +(Fig. <a href="#Fig375">375</a>) are familiar examples. In this the outer part of the +thickness of the pericarp becomes fleshy, or softens like a berry, while +the inner hardens, like a nut. From the way in which the pistil is +constructed, it is evident that the fleshy part here answers to the +lower, and the stone to the upper face of the component leaf. The layers +or concentric portions of a drupe, or of any pericarp which is thus +separable, are named, when thus distinguishable into three portions,—</p> + +<p><i>Epicarp</i>, the external layer, often the mere skin of the fruit,</p> + +<p><i>Mesocarp</i>, the middle layer, which is commonly the fleshy part, and</p> + +<p><i>Endocarp</i>, the innermost layer, the stone. But more commonly only two +portions of a drupe are distinguished, and are named, the outer one</p> + +<p><i>Sarcocarp</i> or <i>Exocarp</i>, for the flesh, the first name referring to the +fleshy character, the second to its being an external layer; and</p> + +<p><i>Putamen</i> or <i>Endocarp</i>, the <i>Stone</i>, within.</p> + +<div class="figcenter"> +<a name="Fig375" id="Fig375"></a> +<img src="images/fig375.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 375. Longitudinal section of a peach, showing flesh, +stone, and seed.</p> +</div> + +<p>357. The typical or true drupe is of a single carpel. But, not to +multiply technical names, this name is extended to all such fruits when +fleshy without and stony within, although of compound pistil,—even to +those having several or separable stones, such as the fruit of Holly. +These stones in such drupes, or drupaceous fruits, are called <i>Pyrenæ</i>, +or <i>Nucules</i>, or simply <i>Nutlets</i> of the drupe.</p> + +<p>358. Of Dry fruits, there is a greater diversity of kinds having +distinct names. The indehiscent sorts are commonly one-seeded.</p> + +<div class="figcenter"> +<a name="Fig376" id="Fig376"></a> +<img src="images/fig376_378.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 376. Akene of a Buttercup. 377. The same, divided +lengthwise, to show the contained seed.</p> +<p><span class="smcap">Fig.</span> 378. Akene of Virgin's-bower, retaining the +feathered style, which aids in dissemination.</p> +</div> + +<p><a name="UNIT_359" id="UNIT_359">359.</a> <b>The Akene or Achenium</b> is a small, dry and indehiscent one-seeded +fruit, often so seed-like in appearance that it is popularly taken for a +naked seed. The fruit of the Buttercup or Crowfoot is a good example, +Fig. <a href="#Fig376">376, 377</a>. Its nature, as a ripened pistil (in this <span class="pagenum"><a name="Page_121" id="Page_121">[Pg 121]</a></span>case a simple +carpel), is apparent by its bearing the remains of a style or stigma, or +a scar from which this has fallen. It may retain the style and use it in +various ways for dissemination (Fig. <a href="#Fig376">378</a>).</p> + +<p>360. The fruit of Compositæ (though not of a single carpel) is also an +akene. In this case the pericarp is invested by an adherent calyx-tube; +the limb of which, when it has any, is called the <span class="smcap">Pappus</span>. This name was +first given to the down like that of the Thistle, but is applied to all +forms under which the limb of the calyx of the "compound flower" +appears. In Lettuce, Dandelion (Fig. <a href="#Fig379">384</a>), and the like, the achenium as +it matures tapers upwards into a slender beak, like a stalk to the +pappus.</p> + +<div class="figcenter"> +<a name="Fig379" id="Fig379"></a> +<img src="images/fig379_384.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 379. Akene of Mayweed (no pappus). 380. That of +Succory (its pappus a shallow cup). 381. Of Sunflower (pappus of two +deciduous scales). 382. Of Sneezeweed (Helenium), with its pappus of +five scales. 383. Of Sow-Thistle, with its pappus of delicate downy +hairs. 384. Of the Dandelion, its pappus raised on a long beak.</p> +</div> + +<p><a name="UNIT_361" id="UNIT_361">361.</a> <b>A Cremocarp</b> (Fig. <a href="#Fig385">385</a>), a name given to the fruit of Umbelliferæ, +consists as it were of a pair of akenes united completely in the +blossom, but splitting apart when ripe into the two closed carpels. Each +of these is a <i>Mericarp</i> or <i>Hemicarp</i>, names seldom used.</p> + +<p><a name="UNIT_362" id="UNIT_362">362.</a> <b>A Utricle</b> is the same as an akene, but with a thin and bladdery +loose pericarp; like that of the Goosefoot or Pigweed (Fig. <a href="#Fig386">386</a>). When +ripe it may burst open irregularly to discharge the seed; or it may open +by a circular line all round, the upper part falling off like a lid; as +in the Amaranth (Fig. <a href="#Fig386">387</a>).</p> + +<div class="figcenter"> +<a name="Fig385" id="Fig385"></a> +<img src="images/fig385.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 385. Fruit (cremocarp) of Osmorrhiza; the two +akene-like ripe carpels separating at maturity from a slender axis or +carpophore.</p> +</div> + +<div class="figcenter"> +<a name="Fig386" id="Fig386"></a> +<img src="images/fig386_387.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 386. Utricle of the common Pigweed (Chenopodium +album).</p> +<p><span class="smcap">Fig.</span> 387. Utricle (pyxis) of Amaranth, opening all round +(circumscissile).</p> +</div> + +<p><a name="UNIT_363" id="UNIT_363">363.</a> <b>A Caryopsis, or Grain</b>, is like an akene with the seed adhering to +the thin pericarp throughout, so that fruit and seed are incorporated +into one body; as in wheat, Indian corn, and other kinds of grain.</p> + +<p><a name="UNIT_364" id="UNIT_364">364.</a> <b>A Nut</b> is a dry and indehiscent fruit, commonly one-celled and +one-seeded, <span class="pagenum"><a name="Page_122" id="Page_122">[Pg 122]</a></span>with a hard, crustaceous, or bony wall, such as the +cocoa-nut, hazelnut, chestnut, and the acorn (Fig. <a href="#Fig36">37</a>, <a href="#Fig388">388</a>.) Here the +involucre, in the form of a cup at the base, is called the <span class="smcap">Cupule</span>. In +the Chestnut the cupule forms the bur; in the Hazel, a leafy husk.</p> + +<div class="figcenter"> +<a name="Fig388" id="Fig388"></a> +<img src="images/fig388.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 388. Nut (acorn) of the Oak, with its cup or +cupule.</p> +</div> + +<p><a name="UNIT_365" id="UNIT_365">365.</a> <b>A Samara, or Key-fruit</b>, is either a nut or an akene, or any other +indehiscent fruit, furnished with a wing, like that of Ash (Fig. <a href="#Fig389">389</a>), +and Elm (Fig. <a href="#Fig389">390</a>). The Maple-fruit is a pair of keys (Fig. <a href="#Fig389">391</a>).</p> + +<p>366. Dehiscent Fruits, or Pods, are of two classes, viz., those of a +simple pistil or carpel, and those of a compound pistil. Two common +sorts of the first are named as follows:—</p> + +<p><a name="UNIT_367" id="UNIT_367">367.</a> <b>The Follicle</b> is a fruit of a simple carpel, which dehisces down one +side only, i. e. by the inner or ventral suture. The fruits of Marsh +Marigold (Fig. <a href="#Fig389">392</a>), Pæony, Larkspur, and Milkweed are of this kind.</p> + +<div class="figcenter"> +<a name="Fig389" id="Fig389"></a> +<img src="images/fig389_394.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 389. Samara or key of the White Ash, winged at end. +390. Samara of the American Elm, winged all round.</p> +<p><span class="smcap">Fig.</span> 391. Pair of samaras of Sugar Maple.</p> +<p><span class="smcap">Fig.</span> 392. Follicle of Marsh Marigold (Caltha palustris).</p> +<p><span class="smcap">Fig.</span> 393. Legume of a Sweet Pea, opened.</p> +<p><span class="smcap">Fig.</span> 394. Loment or jointed legume of a Tick-Trefoil +(Desmodium).</p> +</div> + +<p><a name="UNIT_368" id="UNIT_368">368.</a> <b>The Legume</b> or true Pod, such as the peapod (Fig. <a href="#Fig389">393</a>), and the +fruit of the Leguminous or Pulse family generally, is one which opens +along the dorsal as well as the ventral suture. The two pieces into +which it splits are called <span class="smcap">Valves</span>. A <span class="smcap">Loment</span> is a legume which is +constricted between the seeds, and at length breaks up crosswise into +distinct joints, as in Fig. <a href="#Fig389">394</a>.</p> + +<p>369. The pods or dehiscent fruits belonging to a compound ovary have +several technical names: but they all may be regarded as kinds of</p> + +<p><a name="UNIT_370" id="UNIT_370">370.</a> <b>The Capsule</b>, the dry and dehiscent fruit of any compound pistil. +The capsule may discharge its seeds through chinks or pores, as in the +<span class="pagenum"><a name="Page_123" id="Page_123">[Pg 123]</a></span>Poppy, or burst irregularly in some part, as in Lobelia and the +Snapdragon; but commonly it splits open (or is <i>dehiscent</i>) lengthwise +into regular pieces, called <span class="smcap">Valves</span>.</p> + +<div class="figcenter"> +<a name="Fig395" id="Fig395"></a> +<img src="images/fig395_396.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 395. Capsule of Iris, with loculicidal dehiscence; +below, cut across.</p> +<p><span class="smcap">Fig.</span> 396. Pod of a Marsh St. John's-wort, with septicidal +dehiscence.</p> +</div> + +<p><a name="UNIT_371" id="UNIT_371">371.</a> Regular <i>Dehiscence</i> in a capsule takes place in two ways, which +are best illustrated in pods of two or three cells. It is either</p> + +<p><i>Loculicidal</i>, or, splitting directly into the <i>loculi</i> or cells, that +is, down the back (or the dorsal suture) of each cell or carpel, as in +Iris (Fig. <a href="#Fig395">395</a>); or</p> + +<p><i>Septicidal</i>, that is, splitting through the partitions or <i>septa</i>, as +in St. John's-wort (Fig. <a href="#Fig395">396</a>), Rhododendron, etc. This divides the +capsule into its component carpels, which then open by their ventral +suture.</p> + +<div class="figcenter"> +<a name="Fig397" id="Fig397"></a> +<img src="images/fig397_400.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 397, 398. Diagrams of the two modes.</p> +<p><span class="smcap">Fig.</span> 399. Diagram of septifragal dehiscence of the +loculicidal type. 400. Same of the septicidal or <i>marginicidal</i> type.</p> +</div> + +<p>372. In loculicidal dehiscence the valves naturally bear the partitions +on their middle; in the septicidal, half the thickness of a partition is +borne on the margin of each valve. See the annexed diagrams. A variation +of either mode occurs when the valves break away from the partitions, +these remaining attached in the axis of the fruit. This is called +<i>Septifragal</i> dehiscence. One form is seen in the Morning-Glory (Fig. +<a href="#Fig397">400</a>).</p> + +<p>373. The capsules of Rue, Spurge, and some others, are both loculicidal +and septicidal, and so split into half-carpellary valves or pieces.</p> + +<p><a name="UNIT_374" id="UNIT_374">374.</a> <b>The Silique</b> (Fig. <a href="#Fig401">401</a>) is the technical name of the peculiar pod of +the Mustard family; which is two-celled by a false partition stretched +across between two parietal placentæ. It generally opens by two valves +from below upward, and the placentæ with the partition are left behind +when the valves fall off.</p> + +<p><a name="UNIT_375" id="UNIT_375">375.</a> <b>A Silicle or Pouch</b> is only a short and broad silique, like that of +the Shepherd's Purse, Fig. <a href="#Fig401">402, 403</a>.</p> + +<p><span class="pagenum"><a name="Page_124" id="Page_124">[Pg 124]</a></span></p><div class="figcenter"> +<a name="Fig401" id="Fig401"></a> +<img src="images/fig401_404.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 401. Silique of a Cadamine or Spring Cress.</p> +<p><span class="smcap">Fig.</span> 402. Silicle of Shepherd's Purse.</p> +<p><span class="smcap">Fig.</span> 403. Same, with one valve removed.</p> +<p><span class="smcap">Fig.</span> 404. Pyxis of Purslane, the lid detaching.</p> +</div> + +<p><a name="UNIT_376" id="UNIT_376">376.</a> <b>The Pyxis</b> is a pod which opens by a circular horizontal line, the +upper part forming a lid, as in Purslane (Fig. <a href="#Fig401">404</a>), the Plantain, +Henbane, etc. In these the dehiscence extends all round, or is +<i>circumscissile</i>. So it does in Amaranth (Fig. <a href="#Fig386">387</a>), forming a +one-seeded utricular pyxis. In Jeffersonia, the line does not separate +quite round, but leaves a portion for a hinge to the lid.</p> + +<p>377. Of Multiple or Collective Fruits, which are properly masses of +fruits aggregated into one body (as is seen in the Mulberry (Fig. <a href="#Fig405">408</a>), +Pine-apple, etc.), there are two kinds with special names and of +peculiar structure.</p> + +<div class="figcenter"> +<a name="Fig405" id="Fig405"></a> +<img src="images/fig405_410.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 405. A fig-fruit when young. 406. Same in section. +407. Magnified portion, a slice, showing some of the flowers.</p> +<p><span class="smcap">Fig.</span> 408. A mulberry. 409. One of the grains younger, +enlarged; seen to be a pistillate flower with calyx becoming fleshy. +410. Same, with fleshy calyx cut across.</p> +</div> + +<p><a name="UNIT_378" id="UNIT_378">378.</a> <b>The Syconium or Fig-fruit</b> (Fig. <a href="#Fig405">405, 406</a>) is a fleshy axis or +summit of stem, hollowed out, and lined within by a multitude of minute +flowers, the whole becoming pulpy, and in the common fig, luscious.</p> + +<p><a name="UNIT_379" id="UNIT_379">379.</a> <b>The Strobile or Cone</b> (Fig. <a href="#Fig411">411</a>), is the peculiar multiple fruit of +Pines, Cypresses, and the like; hence named <i>Coniferæ</i>, viz. +cone-bearing <span class="pagenum"><a name="Page_125" id="Page_125">[Pg 125]</a></span>plants. As already shown (<a href="#UNIT_313">313</a>), these cones are <i>open +pistils</i>, mostly in the form of flat scales, regularly overlying each +other, and pressed together in a spike or head. Each scale bears one or +two naked seeds on its inner face. When ripe and dry, the scales turn +back or diverge, and in the Pine the seed peels off and falls, generally +carrying with it a wing, a part of the lining of the scale, which +facilitates the dispersion of the seeds by the wind (Fig. <a href="#Fig411">412, 413</a>). In +Arbor-Vitæ, the scales of the small cone are few, and not very unlike +the leaves. In Cypress they are very thick at the top and narrow at the +base, so as to make a peculiar sort of closed cone. In Juniper and Red +Cedar, the few scales of the very small cone become fleshy, and ripen +into a fruit which closely resembles a berry.</p> + +<div class="figcenter"> +<a name="Fig411" id="Fig411"></a> +<img src="images/fig411_413.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 411. Cone of a common Pitch Pine. 412. Inside view +of a separated scale or open carpel; one seed in place: 413, the other +seed.</p> +</div> + + + +<hr class="chap" /> +<h2><a name="Section_XV" id="Section_XV"></a><span class="smcap">Section XV.</span> THE SEED.</h2> + + +<p>380. Seeds are the final product of the flower, to which all its parts +and offices are subservient. Like the ovule from which it originates, a +seed consists of coats and kernel.</p> + +<div class="figcenter"> +<a name="Fig414" id="Fig414"></a> +<img src="images/fig414.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 414. Seed of a Linden or Basswood cut through +lengthwise, and magnified, the parts lettered: <i>a</i>, the hilum or scar; +<i>b</i>, the outer coat; <i>c</i>, the inner; <i>d</i>, the albumen; <i>e</i>, the embryo.</p> +</div> + +<p><a name="UNIT_381" id="UNIT_381">381.</a> <b>The Seed-coats</b> are commonly two (<a href="#UNIT_320">320</a>), the outer and the inner. +Fig. <a href="#Fig414">414</a> shows the two, in a seed cut through lengthwise. The outer coat +is often hard or crustaceous, whence it is called the <i>Testa</i>, or shell +of the seed; the inner is almost always thin and delicate.</p> + +<div class="figcenter"> +<a name="Fig415" id="Fig415"></a> +<img src="images/fig415_416.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 415. A winged seed of the Trumpet-Creeper.</p> +<p><span class="smcap">Fig.</span> 416. One of Catalpa, the kernel cut to show the +embryo.</p> +</div> + +<div class="figcenter"> +<a name="Fig417" id="Fig417"></a> +<img src="images/fig417.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 417. Seed of Milkweed, with a <i>Coma</i> or tuft of long +silky hairs at one end.</p> +</div> + +<p>382. The shape and the markings, so various in different seeds, depend +mostly on the outer coat. Sometimes this fits the kernel closely; +sometimes it is expanded into a <i>wing</i>, as in the Trumpet-Creeper (Fig. +<a href="#Fig415">415</a>), and occasionally this wing is cut up into shreds or tufts, as in +the Catalpa (Fig. <a href="#Fig415">416</a>); or instead of a wing it may bear a <i>Coma</i>, or +tuft of long and soft hairs, as in the Milkweed or Silkweed (Fig. <a href="#Fig417">417</a>). +The use of wings, or downy tufts is to render the seeds buoyant <span class="pagenum"><a name="Page_126" id="Page_126">[Pg 126]</a></span>for +dispersion by the winds. This is clear, not only from their evident +adaptation to this purpose, but also from the fact that winged and +tufted seeds are found only in fruits that split open at maturity, never +in those that remain closed. The coat of some seeds is beset with long +hairs or wool. <i>Cotton</i>, one of the most important vegetable products, +since it forms the principal clothing of the larger part of the human +race, consists of the long and woolly hairs which thickly cover the +whole surface of the seed. There are also crests or other appendages of +various sorts on certain seeds. A few seeds have an additional, but more +or less incomplete covering, outside of the real seed-coats called an</p> + +<p><a name="UNIT_383" id="UNIT_383">383.</a> <b>Aril, or Arillus.</b> The loose and transparent bag which encloses the +seed of the White Water-Lily (Fig. <a href="#Fig418">418</a>) is of this kind. So is the +<i>mace</i> of the nutmeg; and also the scarlet pulp around the seeds of the +Waxwork (Celastrus) and Strawberry-bush (Euonymus). The aril is a growth +from the extremity of the seed-stalk, or from the placenta when there is +no seed-stalk.</p> + +<div class="figcenter"> +<a name="Fig418" id="Fig418"></a> +<img src="images/fig418.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 418. Seed of White Water Lily, enclosed in its +aril.</p> +</div> + +<p>384. A short and thickish appendage at or close to the hilum in certain +seeds is called a <span class="smcap">Caruncle</span> or <span class="smcap">Strophiole</span> (Fig. <a href="#Fig419">419</a>).</p> + +<div class="figcenter"> +<a name="Fig419" id="Fig419"></a> +<img src="images/fig419.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 419. Seed of Ricinus or Castor oil plant, with +caruncle.</p> +</div> + +<p>385. The various terms which define the position or direction of the +ovule (erect, ascending, etc.) apply equally to the seed: so also the +terms anatropous, orthotropous, campylotropous, etc., as already defined +(<a href="#UNIT_320">320</a>, <a href="#UNIT_321">321</a>), and such terms as</p> + +<p><span class="smcap">Hilum</span>, or <i>Scar</i> left where the seed-stalk or funiculus falls away, or +where the seed was attached directly to the placenta when there is no +seed-stalk.</p> + +<p><span class="smcap">Rhaphe</span>, the line or ridge which runs from the hilum to the chalaza in +anatropous and amphitropous seeds.</p> + +<p><span class="smcap">Chalaza</span>, the place where the seed-coats and the kernel or nucleus are +organically connected,—at the hilum in orthotropous and campylotropous +seeds, at the extremity of the rhaphe or tip of the seed in other kinds.</p> + +<p><span class="smcap">Micropyle</span>, answering to the <i>Foramen</i> or orifice of the ovule. Compare +the accompanying figures and those of the ovules, Fig. <a href="#Fig341">341</a>-<a href="#Fig355">355</a>.</p> + +<p><span class="pagenum"><a name="Page_127" id="Page_127">[Pg 127]</a></span></p><div class="figcenter"> +<a name="Fig420" id="Fig420"></a> +<img src="images/fig420_423.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 420. Seed of a Violet (anatropous): <i>a</i>, hilum; +<i>b</i>, rhaphe; <i>c</i>, chalaza.</p> +<p><span class="smcap">Fig.</span> 421. Seed of a Larkspur (also anatropous); the parts +lettered as in the last.</p> +<p><span class="smcap">Fig.</span> 422. The same, cut through lengthwise: <i>a</i>, the +hilum; <i>c</i>, chalaza; <i>d</i>, outer seed coat; <i>e</i>, inner seed-coat; <i>f</i>, +the albumen; <i>g</i>, the minute embryo.</p> +<p><span class="smcap">Fig.</span> 423. Seed of a St. John's-wort, divided lengthwise; +here the whole kernel is embryo.</p> +</div> + +<p><a name="UNIT_386" id="UNIT_386">386.</a> <b>The Kernel, or Nucleus</b>, is the whole body of the seed within the +coats. In many seeds the kernel is all <i>Embryo</i>; in others a large part +of it is the <i>Albumen</i>. For example, in Fig. <a href="#Fig420">423</a>, it is wholly embryo; +in Fig. <a href="#Fig420">422</a>, all but the small speck (<i>g</i>) is albumen.</p> + +<p><a name="UNIT_387" id="UNIT_387">387.</a> <b>The Albumen or Endosperm</b> of the seed is sufficiently characterized +and its office explained in Sect. III., <a href="#UNIT_31">31-35</a>.</p> + +<p><a name="UNIT_388" id="UNIT_388">388.</a> <b>The Embryo</b> or <i>Germ</i>, which is the rudimentary plantlet and the +final result of blossoming, and its development in germination have been +extensively illustrated in Sections <a href="#Section_II">II.</a> and <a href="#Section_III">III.</a> Its essential parts are +the <i>Radicle</i> and the <i>Cotyledons</i>.</p> + +<p><a name="UNIT_389" id="UNIT_389">389.</a> <b>Its Radicle or Caulicle</b> (the former is the term long and generally +used in botanical descriptions, but the latter is the more correct one, +for it is the initial stem, which merely gives origin to the root), as +to its position in the seed, always points to and lies near the +micropyle. In relation to the pericarp it is</p> + +<p><i>Superior</i>, when it points to the apex of the fruit or cell, and</p> + +<p><i>Inferior</i>, when it points to its base, or downward.</p> + +<div class="figcenter"> +<a name="Fig424" id="Fig424"></a> +<img src="images/fig424.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 424. Embryo of Calycanthus; upper part cut away, to +show the convolute cotyledons.</p> +</div> + +<p><a name="UNIT_390" id="UNIT_390">390.</a> <b>The Cotyledons</b> have already been illustrated as respects their +number,—giving the important distinction of <i>Dicotyledonous</i>, +<i>Polycotyledonous</i> and <i>Monocotyledonous</i> embryos (<a href="#UNIT_36">36-43</a>),—also as +regards their thickness, whether <i>foliaceous</i> or <i>fleshy</i>; and some of +the very various shapes and adaptations to the seed have been figured. +They may be straight, or folded, or rolled up. In the latter case the +cotyledons may be rolled up as it were from one margin, as in +Calycanthus (Fig. <a href="#Fig424">424</a>), or from apex to base in a flat spiral, or they +may be both folded (<i>plicate</i>) and rolled up (<i>convolute</i>), as in Sugar +Maple (Fig. <a href="#Fig11">11</a>.) In one very natural family, the Cruciferæ, two +different modes prevail in the way the two cotyledons are brought round +against the radicle. In one series they are</p> + +<p><span class="pagenum"><a name="Page_128" id="Page_128">[Pg 128]</a></span></p><p><i>Accumbent</i>, that is, the edges of the flat cotyledons lie against the +radicle, as in Fig. <a href="#Fig425">425, 426</a>. In another they are</p> + +<div class="figcenter"> +<a name="Fig425" id="Fig425"></a> +<img src="images/fig425_426.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 425. Seed of Bitter Cress, Barbarea, cut across to +show the accumbent cotyledons. 426. Embryo of same, whole.</p> +</div> + +<p><i>Incumbent</i>, or with the plane of the cotyledons brought up in the +opposite direction, so that the back of one of them lies against the +radicle, as shown in Fig. <a href="#Fig427">427, 428</a>.</p> + +<div class="figcenter"> +<a name="Fig427" id="Fig427"></a> +<img src="images/fig427_428.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 427. Seed of a Sisymbrium, cut across to show the +incumbent cotyledons. 428. Embryo of the same, detached whole.</p> +</div> + +<p>391. As to the situation of the embryo with respect to the albumen of +the seed, when this is present in any quantity, the embryo may be +<i>Axile</i>, that is occupying the axis or centre, either for most of its +length, as in Violet (Fig. <a href="#Fig429">429</a>), Barberry (Fig. <a href="#Fig46">48</a>), and Pine (Fig. <a href="#Fig56">56</a>); +and in these it is straight. But it may be variously curved or coiled in +the albumen, as in Helianthemum (Fig. <a href="#Fig429">430</a>), in a Potato-seed (Fig. <a href="#Fig46">50</a>), +or Onion-seed (Fig. <a href="#Fig58">60</a>), and Linden (Fig. <a href="#Fig414">414</a>); or it may be coiled +around the outside of the albumen, partly or into a circle, as in +Chickweed (Fig. <a href="#Fig431">431, 432</a>) and in Mirabilis (Fig. <a href="#Fig46">52</a>). The latter mode +prevails in Campylotropous seeds. In the cereal grains, such as Indian +Corn (Fig. <a href="#Fig62">67</a>) and Rice (Fig. <a href="#Fig429">430<sup><i>a</i></sup></a>), and in all other Grasses, the +embryo is straight and applied to the outside of the abundant albumen.</p> + +<div class="figcenter"> +<a name="Fig429" id="Fig429"></a> +<img src="images/fig429_430.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 429. Section of seed of Violet; anatropous with +straight axile embryo in the albumen. 430. Section of seed of Rock Rose, +Helianthemum Canadense; orthotropous, with curved embryo in the albumen. +430<sup><i>a</i></sup>. Section of a grain of Rice, lengthwise, showing the embryo +outside the albumen, which forms the principal bulk.</p> +</div> + +<div class="figcenter"> +<a name="Fig431" id="Fig431"></a> +<img src="images/fig431_432.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 431. Seed of a Chickweed, campylotropous. 432. +Section of same, showing slender embryo coiled around the outside of the +albumen of the kernel.</p> +</div> + +<p>392. The matured seed, with embryo ready to germinate and reproduce the +kind, completes the cycle of the vegetable life in a phanerogamous +plant, the account of which began with the seed and seedling.</p> + + + +<hr class="chap" /> +<h2><a name="Section_XVI" id="Section_XVI"></a><span class="smcap">Section XVI.</span> VEGETABLE LIFE AND WORK.</h2> + + +<p>393. The following simple outlines of the anatomy and physiology of +plants (<a href="#UNIT_3">3</a>) are added to the preceding structural part for the better +preparation of students in descriptive and systematic botany; also to +give to all learners some general idea of the life, growth, intimate +structure, and action of the beings which compose so large a part of +organic nature. Those who would extend and verify the facts and +principles here outlined will use the Physiological Botany of the +"Botanical Text Book," by Professor Goodale, or some similar book.</p> + + +<p><span class="pagenum"><a name="Page_129" id="Page_129">[Pg 129]</a></span></p><h3><a name="Subsect_XVI_1" id="Subsect_XVI_1">§ 1.</a> ANATOMICAL STRUCTURE AND GROWTH.</h3> + +<p><a name="UNIT_394" id="UNIT_394">394.</a> <b>Growth</b> <i>is the increase of a living thing in size and substance</i>. +It appears so natural that plants and animals should grow, that one +rarely thinks of it as requiring explanation. It seems enough to say +that a thing is so because it grew so. Growth from the seed, the +germination and development of an embryo into a plantlet, and at length +into a mature plant (as illustrated in Sections <a href="#Section_II">II.</a> and <a href="#Section_III">III.</a>), can be +followed by ordinary observation. But the embryo is already a miniature +plantlet, sometimes with hardly any visible distinction of parts, but +often one which has already made very considerable growth in the seed. +To investigate the formation and growth of the embryo itself requires +well-trained eyes and hands, and the expert use of a good compound +microscope. So this is beyond the reach of a beginner.</p> + +<p>395. Moreover, although observation may show that a seedling, weighing +only two or three grains, may double its bulk and weight every week of +its early growth, and may in time produce a huge amount of vegetable +matter, it is still to be asked what this vegetable matter is, where it +came from, and by what means plants are able to increase and accumulate +it, and build it up into the fabric of herbs and shrubs and lofty trees.</p> + +<p><a name="UNIT_396" id="UNIT_396">396.</a> <b>Protoplasm.</b> All this fabric was built up under life, but only a +small portion of it is at any one time alive. As growth proceeds, life +is passed on from the old to the new parts, much as it has passed on +from parent to offspring, from generation to generation in unbroken +continuity. <i>Protoplasm</i> is the common name of that plant-stuff in which +life essentially resides. All growth depends upon it; for it has the +peculiar power of growing and multiplying and building up a living +structure,—the animal no less than the vegetable structure, for it is +essentially the same in both. Indeed, all the animal protoplasm comes +primarily from the vegetable, which has the prerogative of producing it; +and the protoplasm of plants furnishes all that portion of the food of +animals which forms their flesh and living fabric.</p> + +<p>397. The very simplest plants (if such may specifically be called plants +rather than animals, or one may say, the simplest living things) are +mere particles, or pellets, or threads, or even indefinite masses of +protoplasm of vague form, which possess powers of motion or of changing +their shape, of imbibing water, air, and even other matters, and of +assimilating these into plant-stuff for their own growth and +multiplication. Their growth is increase in substance by incorporation +of that which they take in and assimilate. Their multiplication is by +spontaneous division of their substance or body into two or more, each +capable of continuing the process.</p> + +<p>398. The embryo of a phanerogamous plant at its beginning (<a href="#UNIT_344">344</a>) is +essentially such a globule of protoplasm, which soon constricts itself +into two and more such globules, which hold together inseparably in a +row; then the last of the row divides without separation in the two +other planes, to <span class="pagenum"><a name="Page_130" id="Page_130">[Pg 130]</a></span>form a compound mass, each grain or globule of which +goes on to double itself as it grows; and the definite shaping of this +still increasing mass builds up the embryo into its form.</p> + +<div class="figcenter"> +<a name="Fig433" id="Fig433"></a> +<img src="images/fig433_436.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 433-436. Figures to illustrate the earlier stages in +the formation of an embryo; a single mass of protoplasm (Fig. 433) +dividing into two, three, and then into more incipient cells, which by +continued multiplication build up an embryo.</p> +</div> + +<p><a name="UNIT_399" id="UNIT_399">399.</a> <b>Cell-walls.</b> While this growth was going on, each grain of the +forming structure formed and clothed itself with a coat, thin and +transparent, of something different from protoplasm,—something which +hardly and only transiently, if at all, partakes of the life and action. +The protoplasm forms the living organism; the coat is a kind of +protective covering or shell. The protoplasm, like the flesh of animals +which it gives rise to, is composed of four chemical elements: Carbon, +Hydrogen, Oxygen, and Nitrogen. The coating is of the nature of wood +(is, indeed, that which makes wood), and has only the three elements, +Carbon, Hydrogen, and Oxygen, in its composition.</p> + +<div class="figcenter"> +<a name="Fig437" id="Fig437"></a> +<img src="images/fig437.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 437. Magnified view of some of a simple fresh water +Alga, the Tetraspora lubrica, each sphere of which may answer to an +individual plant.</p> +</div> + +<p>400. Although the forming structure of an embryo in the fertilized ovule +is very minute and difficult to see, there are many simple plants of +lowest grade, abounding in pools of water, which more readily show the +earlier stages or simplest states of plant-growth. One of these, which +is common in early spring, requires only moderate magnifying power to +bring to view what is shown in Fig. <a href="#Fig437">437</a>. In a slimy mass which holds all +loosely together, little spheres of green vegetable matter are seen, +assembled in fours, and these fours themselves in clusters of fours. A +transient inspection shows, what prolonged watching would confirm, that +each sphere divides first in one plane, then in the other, to make four, +soon acquiring the size of the original, and so on, producing successive +groups of fours. These pellets each form on their surface a transparent +wall, like that just described. The delicate wall is for some time +capable of expansive growth, but is from the first much firmer than the +protoplasm within; through it the latter imbibes surrounding moisture, +which becomes a watery sap, occupying vacuities in the protoplasmic mass +which enlarge or run together as the periphery increases and distends. +When full grown the protoplasm may become a mere lining to the wall, or +some of it central, as a nucleus, this usually connected with the +wall-lining by delicate threads of the same substance. So, when full +grown, the wall with its lining—a vesicle, containing liquid or some +<span class="pagenum"><a name="Page_131" id="Page_131">[Pg 131]</a></span>solid matters and in age mostly air—naturally came to be named a Cell. +But the name was suggested by, and first used only for, cells in +combination or built up into a fabric, much as a wall is built of +bricks, that is, into a</p> + +<p><a name="UNIT_401" id="UNIT_401">401.</a> <b>Cellular Structure or Tissue.</b> Suppose numerous cells like those of +Fig. <a href="#Fig437">437</a> to be heaped up like a pile of cannon-balls, and as they grew, +to be compacted together while soft and yielding; they would flatten +where they touched, and each sphere, being touched by twelve surrounding +ones would become twelve-sided. Fig. <a href="#Fig438">438</a> would represent one of them. +Suppose the contiguous faces to be united into one wall or partition +between adjacent cavities, and a <i>cellular structure</i> would be formed, +like that shown in Fig. <a href="#Fig438">439</a>. Roots, stems, leaves, and the whole of +phanerogamous plants are a fabric of countless numbers of such cells. No +such exact regularity in size and shape is ever actually found; but a +nearly truthful magnified view of a small portion of a slice of the +flower-stalk of a Calla Lily (Fig. <a href="#Fig440">440</a>) shows a fairly corresponding +structure; except that, owing to the great air-spaces of the interior, +the fabric may be likened rather to a stack of chimneys than to a solid +fabric. In young and partly transparent parts one may discern the +cellular structure by looking down directly on the surface, as of a +forming root. (Fig. <a href="#Fig81">82</a>, <a href="#Fig441">441, 442</a>).</p> + +<div class="figcenter"> +<a name="Fig438" id="Fig438"></a> +<img src="images/fig438_439.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 438. Diagram of a vegetable cell, such as it would +be if when spherical it were equally pressed by similar surrounding +cells in a heap.</p> +<p><span class="smcap">Fig.</span> 439. Ideal construction of cellular tissue so +formed, in section.</p> +</div> + +<div class="figcenter"> +<a name="Fig440" id="Fig440"></a> +<img src="images/fig440.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 440. Magnified view of a portion of a transverse +slice of stem of Calla Lily. The great spaces are tubular air-channels +built up by the cells.</p> +</div> + +<p><a name="UNIT_402" id="UNIT_402">402.</a> The substance of which cell-walls are mainly composed is called +<span class="smcap">Cellulose</span>. It is essentially the same in the stem of a delicate leaf or +petal and in the wood of an Oak, except that in the latter the walls are +<span class="pagenum"><a name="Page_132" id="Page_132">[Pg 132]</a></span>much thickened and the calibre small. The protoplasm of each living +cell appears to be completely shut up and isolated in its shell of +cellulose; but microscopic investigation has brought to view, in many +cases, minute threads of protoplasm which here and there traverse the +cell-wall through minute pores, thus connecting the living portion of +one cell with that of adjacent cells. (See Fig. <a href="#Fig444">447</a>, &c.)</p> + +<div class="figcenter"> +<a name="Fig441" id="Fig441"></a> +<img src="images/fig441_442.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 441. Much magnified small portion of young root of a +seedling Maple (such as of Fig. <a href="#Fig81">82</a>); and 442, a few cells of same more +magnified. The prolongations from the back of some of the cells are root +hairs.</p> +</div> + +<p>403. The hairs of plants are cells formed on the surface; either +elongated single cells (like the root-hairs of Fig. <a href="#Fig441">441, 442</a>), or a row +of shorter cells. Cotton fibres are long and simple cells growing from +the surface of the seed.</p> + +<p>404. The size of the cells of which common plants are made up varies +from about the thirtieth to the thousandth of an inch in diameter. An +ordinary size of short or roundish cells is from 1/300 to 1/500 of an +inch; so that there may generally be from 27 to 125 millions of cells in +the compass of a cubic inch!</p> + +<p>405. Some parts are built up as a compact structure; in others cells are +arranged so as to build up regular air-channels, as in the stems of +aquatic and other water-loving plants (Fig. <a href="#Fig440">440</a>), or to leave irregular +spaces, as in the lower part of most leaves, where the cells only here +and there come into close contact (Fig. <a href="#Fig443">443</a>).</p> + +<div class="figcenter"> +<a name="Fig443" id="Fig443"></a> +<img src="images/fig443.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 443. Magnified section through the thickness of a +leaf of Florida Star-Anise.</p> +</div> + +<p>406. All such soft cellular tissue, like this of leaves, that of pith, +and of the green bark, is called <span class="smcap">Parenchyma</span>, while fibrous and woody +parts are composed of <span class="smcap">Prosenchyma</span>, that is, of peculiarly transformed</p> + +<p><a name="UNIT_407" id="UNIT_407">407.</a> <b>Strengthening Cells.</b> Common cellular tissue, which makes up the +whole structure of all very young plants, and the whole of Mosses and +other vegetables of the lowest grade, even when full grown, is too +tender or too brittle to give needful strength and toughness for plants +which are to rise to any considerable height and support themselves. In +these needful strength is imparted, and the conveyance of sap through +the plant is facilitated, by the change, as they are formed, of some +cells into thicker-walled and tougher tubes, and by the running together +of some of <span class="pagenum"><a name="Page_133" id="Page_133">[Pg 133]</a></span>these, or the prolongation of others, into hollow fibres or +tubes of various size. Two sorts of such transformed cells go together, +and essentially form the</p> + +<p><a name="UNIT_408" id="UNIT_408">408.</a> <b>Wood.</b> This is found in all common herbs, as well as in shrubs and +trees, but the former have much less of it in proportion to the softer +cellular tissue. It is formed very early in the growth of the root, +stem, and leaves,—traces of it appearing in large embryos even while +yet in the seed. Those cells that lengthen, and at the same time thicken +their walls form the proper <span class="smcap">Woody Fibre</span> or <span class="smcap">Wood-cells</span>; those of larger +size and thinner walls, which are thickened only in certain parts so as +to have peculiar markings, and which often are seen to be made up of a +row of cylindrical cells, with the partitions between absorbed or broken +away, are called <span class="smcap">Ducts</span>, or sometimes <span class="smcap">Vessels</span>. There are all gradations +between wood-cells and ducts, and between both these and common cells. +But in most plants the three kinds are fairly distinct.</p> + +<div class="figcenter"> +<a name="Fig444" id="Fig444"></a> +<img src="images/fig444_447.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 444. Magnified wood-cells of the bark (bast-cells) +of Basswood, one and part of another. 445. Some wood cells from the wood +(and below part of a duct); and 446, a detached wood-cell of the same; +equally magnified.</p> +<p><span class="smcap">Fig.</span> 447. Some wood cells from Buttonwood, Platanus, +highly magnified, a whole cell and lower end of another on the left; a +cell cut half away lengthwise, and half of another on the right; some +pores or pits (<i>a</i>) seen on the left; while <i>b b</i> mark sections through +these on the cut surface. When living and young the protoplasm extends +into these and by minuter perforations connects across them. In age the +pits become open passages, facilitating the passage of sap and air.</p> +</div> + +<p>409. The proper cellular tissue, or <i>parenchyma</i>, is the ground-work of +root, stem, and leaves; this is traversed, chiefly lengthwise, by the +strengthening and conducting tissue, wood-cells and duct-cells, in the +form of bundles or threads, which, in the stems and stalks of herbs are +fewer and comparatively scattered, but in shrubs and trees so numerous +and crowded that in the stems and all permanent parts they make a solid +mass of wood. They extend into and ramify in the leaves, spreading out +in a horizontal plane, as the framework of ribs and veins, which +supports the softer cellular portion or parenchyma.</p> + +<p><a name="UNIT_410" id="UNIT_410">410.</a> <b>Wood-Cells, or Woody Fibres</b>, consist of tubes, commonly between one +and two thousandths, but in Pine-wood sometimes two or three hundredths, +of an inch in diameter. Those from the tough bark of the Basswood, +<span class="pagenum"><a name="Page_134" id="Page_134">[Pg 134]</a></span>shown in Fig. <a href="#Fig444">444</a>, are only the fifteen-hundredth of an inch wide. +Those of Buttonwood (Fig. <a href="#Fig444">447</a>) are larger, and are here highly magnified +besides. The figures show the way wood-cells are commonly put together, +namely, with their tapering ends overlapping each other,—spliced +together, as it were,—thus giving more strength and toughness. In hard +woods, such as Hickory and Oak, the walls of these tubes are very thick, +as well as dense; while in soft woods, such as White Pine and Basswood, +they are thinner.</p> + +<p><a name="UNIT_411" id="UNIT_411">411.</a> Wood-cells in the bark are generally longer, finer, and tougher +than those of the proper wood, and appear more like fibres. For example, +Fig. <a href="#Fig444">446</a> represents a cell of the wood of Basswood of average length, +and Fig. <a href="#Fig444">444</a> one (and part of another) of the fibrous bark, both drawn +to the same scale. As these long cells form the principal part of +fibrous bark, or <i>bast</i>, they are named <i>Bast-cells</i> or <i>Bast-fibres</i>. +These give the great toughness and flexibility to the inner bark of +Basswood (i. e. Bast-wood) and of Leatherwood; and they furnish the +invaluable fibres of flax and hemp; the proper wood of their stems being +tender, brittle, and destroyed by the processes which separate for use +the tough and slender bast-cells. In Leatherwood (Dirca) the bast-cells +are remarkably slender. A view of one, if magnified on the scale of Fig. +<a href="#Fig444">444</a>, would be a foot and a half long.</p> + +<div class="figcenter"> +<a name="Fig448" id="Fig448"></a> +<img src="images/fig448_450.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 448. Magnified bit of a pine shaving, taken parallel +with the silver grain. 449. Separate whole wood-cell, more magnified. +450. Same, still more magnified; both sections represented: <i>a</i>, disks +in section, <i>b</i>, in face.</p> +</div> + +<p>412. The wood-cells of Pines, and more or less of all other Coniferous +trees, have on two of their sides very peculiar disk-shaped markings +(Fig. <a href="#Fig448">448-450</a>) by which that kind of wood is recognizable.</p> + +<div class="figcenter"> +<a name="Fig451" id="Fig451"></a> +<img src="images/fig451_452.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 451, 452. A large and a smaller dotted duct from +Grape-Vine.</p> +</div> + +<p><a name="UNIT_413" id="UNIT_413">413.</a> <b>Ducts</b>, also called <span class="smcap">Vessels</span>, are mostly larger than wood-cells: +indeed, some of them, as in Red Oak, have calibre large enough to be +discerned on a cross section by the naked eye. They make the visible +porosity of such kinds of wood. This is particularly the case with</p> + +<p><i>Dotted</i> ducts (Fig. <a href="#Fig451">451, 452</a>), the surface of which appears as if +riddled with round or oval pores. Such ducts are commonly made up of a +row of large cells more or less confluent into a tube.</p> + +<p><i>Scalariform</i> ducts (Fig. <a href="#Fig453">458, 459</a>), common in Ferns, and generally +angled by mutual pressure in the bundles, <span class="pagenum"><a name="Page_135" id="Page_135">[Pg 135]</a></span>have transversely elongated +thin places, parallel with each other, giving a ladder-like appearance, +whence the name.</p> + +<p><i>Annular</i> ducts (Fig. <a href="#Fig453">457</a>) are marked with cross lines or rings, which +are thickened portions of the cell-wall.</p> + +<div class="figcenter"> +<a name="Fig453" id="Fig453"></a> +<img src="images/fig453_459.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 453, 454. Spiral ducts which uncoil into a single +thread. 455. Spiral duct which tears up as a band. 456. An annular duct, +with variations above. 457. Loose spiral duct passing into annular. 458. +Scalariform ducts of a Fern; part of a bundle, prismatic by pressure. +459. One torn into a band.</p> +</div> + +<p><i>Spiral</i> ducts or vessels (Fig. <a href="#Fig453">453-455</a>) have thin walls, strengthened +by a spiral fibre adherent within. This is as delicate and as strong as +spider-web: when uncoiled by pulling apart, it tears up and annihilates +the cell-wall. The uncoiled threads are seen by gently pulling apart +many leaves, such as those of Amaryllis, or the stalk of a Strawberry +leaflet.</p> + +<div class="figcenter"> +<a name="Fig460" id="Fig460"></a> +<img src="images/fig460_461.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 460. Milk Vessels of Dandelion, with cells of the +common cellular tissue. 461. Others from the same older and gorged with +milky juice. All highly magnified.</p> +</div> + +<p><i>Laticiferous ducts</i>, <i>Vessels of the Latex</i>, or <i>Milk-vessels</i> are +peculiar branching tubes which hold <i>latex</i> or milky juice in certain +plants. It is very difficult to see them, and more so to make out their +nature. They are peculiar in branching and inosculating, so as to make a +net-work of tubes, running in among the cellular tissue; and they are +very small, except when gorged and old (Fig. <a href="#Fig460">460, 461</a>).</p> + + +<p><span class="pagenum"><a name="Page_136" id="Page_136">[Pg 136]</a></span></p><h3><a name="Subsect_XVI_2" id="Subsect_XVI_2">§ 2.</a> CELL-CONTENTS.</h3> + +<p>414. The living contents of young and active cells are mainly protoplasm +with water or watery sap which this has imbibed. Old and effete cells +are often empty of solid matter, containing only water with whatever may +be dissolved in it, or air, according to the time and circumstances. All +the various products which plants in general elaborate, or which +particular plants specially elaborate, out of the common food which they +derive from the soil and the air, are contained in the cells, and in the +cells they are produced.</p> + +<p><a name="UNIT_415" id="UNIT_415">415.</a> <b>Sap</b> is a general name for the principal liquid contents,—<i>Crude +sap</i>, for that which the plant takes in, <i>Elaborated sap</i> for what it +has digested or assimilated. They must be undistinguishably mixed in the +cells.</p> + +<p>416. Among the solid matters into which cells convert some of their +elaborated sap two are general and most important. These are +<i>Chlorophyll</i> and <i>Starch</i>.</p> + +<p><a name="UNIT_417" id="UNIT_417">417.</a> <b>Chlorophyll</b> (meaning <i>leaf-green</i>) is what gives the green color to +herbage. It consists of soft grains of rather complex nature, partly +wax-like, partly protoplasmic. These abound in the cells of all common +leaves and the green rind of plants, wherever exposed to the light. The +green color is seen through the transparent skin of the leaf and the +walls of the containing cells. Chlorophyll is essential to ordinary +assimilation in plants: by its means, under the influence of sunlight, +the plant converts crude sap into vegetable matter.</p> + +<p>418. Far the largest part of all vegetable matter produced is that which +goes to build up the plant's fabric or cellular structure, either +directly or indirectly. There is no one good name for this most +important product of vegetation. In its final state of cell-walls, the +permanent fabric of herb and shrub and tree, it is called <i>Cellulose</i> +(<a href="#UNIT_408">408</a>): in its most soluble form it is <i>Sugar</i> of one or another kind; in +a less soluble form it is <i>Dextrine</i>, a kind of liquefied starch: in the +form of solid grains stored up in the cells it is <i>Starch</i>. By a series +of slight chemical changes (mainly a variation in the water entering +into the composition), one of these forms is converted into another.</p> + +<p><a name="UNIT_419" id="UNIT_419">419.</a> <b>Starch</b> (<i>Farina</i> or <i>Fecula</i>) is the form in which this common +plant material is, as it were, laid by for future use. It consists of +solid grains, somewhat different in form in different plants, in size +varying from 1/300 to 1/4000 of an inch, partly translucent when wet, +and of a pearly lustre. From the concentric lines, which commonly appear +under the microscope, the grains seem to be made up of layer over layer. +When loose they are commonly oval, as in potato-starch (Fig. <a href="#Fig462">462</a>): when +much compacted the grains may become angular (Fig. <a href="#Fig462">463</a>).</p> + +<div class="figcenter"> +<a name="Fig462" id="Fig462"></a> +<img src="images/fig462_463.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 462. Some magnified starch-grains, in two cells of a +potato. 463. Some cells of the albumen or floury part of Indian Corn, +filled with starch-grains.</p> +</div> + +<p>420. The starch in a potato was produced in the foliage. In the soluble +form of dextrine, or that of sugar, it was conveyed through the cells of +the herbage and stalks to a subterranean shoot, and there stored up in +the <span class="pagenum"><a name="Page_137" id="Page_137">[Pg 137]</a></span>tuber. When the potato sprouts, the starch in the vicinity of +developing buds or eyes is changed back again, first into mucilaginous +dextrine, then into sugar, dissolved in the sap, and in this form it is +made to flow to the growing parts, where it is laid down into cellulose +or cell-wall.</p> + +<div class="figcenter"> +<a name="Fig464" id="Fig464"></a> +<img src="images/fig464_470.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 464. Four cells from dried Onion-peel, each holding +a crystal of different shape, one of them twinned. 465. Some cells from +stalk of Rhubarb-plant, three containing chlorophyll; two (one torn +across) with rhaphides. 466. Rhaphides in a cell, from Arisæma, with +small cells surrounding. 467. Prismatic crystals from the bark of +Hickory. 468. Glomerate crystal in a cell, from Beet-root. 469. A few +cells of Locust-bark, a crystal in each. 470. A detached cell, with +rhaphides being forced out, as happens when put in water.</p> +</div> + +<p>421. Besides these cell-contents which are in obvious and essential +relation to nutrition, there are others the use of which is +problematical. Of such the commonest are</p> + +<p><a name="UNIT_422" id="UNIT_422">422.</a> <b>Crystals.</b> These when slender or needle-shaped are called <span class="smcap">Rhaphides</span>. +They are of inorganic matter, usually of oxalate or phosphate or +sulphate of lime. Some, at least of the latter, may be direct +crystallizations of what is taken in dissolved in the water absorbed, +but others must be the result of some elaboration in the plant. Some +plants have hardly any; others abound in them, especially in the foliage +and bark. In Locust-bark almost every cell holds a crystal; so that in a +square inch not thicker than writing-paper there may be over a million +and a half of them. When <span class="pagenum"><a name="Page_138" id="Page_138">[Pg 138]</a></span>needle-shaped (rhaphides), as in stalks of +Calla-Lily, Rhubarb, or Four-o'clock, they are usually packed in +sheaf-like bundles. (Fig. <a href="#Fig464">465, 466</a>.)</p> + + +<h3><a name="Subsect_XVI_3" id="Subsect_XVI_3">§ 3.</a> ANATOMY OF ROOTS AND STEMS.</h3> + +<p>423. This is so nearly the same that an account of the internal +structure of stems may serve for the root also.</p> + +<p>424. At the beginning, either in the embryo or in an incipient shoot +from a bud, the whole stem is of tender cellular tissue or parenchyma. +But wood (consisting of wood-cells and ducts or vessels) begins to be +formed in the earliest growth; and is from the first arranged in two +ways, making two general kinds of wood. The difference is obvious even +in herbs, but is more conspicuous in the enduring stems of shrubs and +trees.</p> + +<p>425. On one or the other of these two types the stems of all +phanerogamous plants are constructed. In one, the wood is made up of +separate threads, scattered here and there throughout the whole diameter +of the stem. In the other, the wood is all collected to form a layer (in +a slice across the stem appearing as a ring) between a central cellular +part which has none in it, the <i>Pith</i>, and an outer cellular part, the +<i>Bark</i>.</p> + +<div class="figcenter"> +<a name="Fig471" id="Fig471"></a> +<img src="images/fig471_473.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 471. Diagram of structure of Palm or Yucca. 472. +Structure of a Corn stalk, in transverse and longitudinal section. 473. +Same of a small Palm stem. The dots on the cross sections represent cut +ends of the woody bundles or threads.</p> +</div> + +<p>426. An Asparagus-shoot and a Corn-stalk for herbs, and a rattan for a +woody kind, represent the first kind. To it belong all plants with +monocotyledonous embryo (<a href="#UNIT_40">40</a>). A Bean-stalk and the stem of any common +shrub or tree represent the second; and to it belong all plants with +dicotyledonous or polycotyledonous embryo. The first has been called, +not very properly, <i>Endogenous</i>, which means inside-growing; the second, +properly enough, <i>Exogenous</i>, or outside-growing.</p> + +<p><a name="UNIT_427" id="UNIT_427">427.</a> <b>Endogenous Stems</b>, those of Monocotyls (<a href="#UNIT_40">40</a>), attain their greatest +size and most characteristic development in Palms and Dragon-trees, +therefore chiefly in warm climates, although the Palmetto and some +<span class="pagenum"><a name="Page_139" id="Page_139">[Pg 139]</a></span>Yuccas become trees along the southern borders of the United States. In +such stems the woody bundles are more numerous and crowded toward the +circumference, and so the harder wood is outside; while in an exogenous +stem the oldest and hardest wood is toward the centre. An endogenous +stem has no clear distinction of pith, bark, and wood, concentrically +arranged, no silver grain, no annual layers, no bark that peels off +clean from the wood. Yet old stems of Yuccas and the like, that continue +to increase in diameter, do form a sort of layers and a kind of scaly +bark when old. Yuccas show well the curving of the woody bundles (Fig. +<a href="#Fig471">471</a>) which below taper out and are lost at the rind.</p> + +<div class="figcenter"> +<a name="Fig474" id="Fig474"></a> +<img src="images/fig474.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 474. Short piece of stem of Flax, magnified, showing +the bark, wood, and pith in a cross section.</p> +</div> + +<p><a name="UNIT_428" id="UNIT_428">428.</a> <b>Exogenous Stems</b>, those of Dicotyls (<a href="#UNIT_37">37</a>), or of plants coming from +dicotyledonous and also polycotyledonous embryos, have a structure which +is familiar in the wood of our ordinary trees and shrubs. It is the same +in an herbaceous shoot (such as a Flax-stem, Fig. <a href="#Fig474">474</a>) as in a +Maple-stem of the first year's growth, except that the woody layer is +commonly thinner or perhaps reduced to a circle of bundles. It was so in +the tree-stem at the beginning. The wood all forms in a cylinder,—in +cross section a ring—around a central cellular part, dividing the +cellular core within, the pith, from a cellular bark without. As the +wood-bundles increase in number and in size, they press upon each other +and become wedge-shaped in the cross section; and they continue to grow +from the outside, next the bark, so that they become very thin wedges or +plates. Between the plates or wedges are very thin plates (in cross +section lines) of much compressed cellular tissue, which connect the +pith with the bark. The plan of a one-year-old woody stem of this kind +is exhibited in the figures, which are essentially diagrams.</p> + +<div class="figcenter"> +<a name="Fig475" id="Fig475"></a> +<img src="images/fig475_477.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 475. Diagram of a cross section of a very young +exogenous stem, showing six woody bundles or wedges. 476. Same later, +with wedges increased to twelve. 477. Still later, the wedges filling +the space, separated only by the thin lines, or medullary rays, running +from pith to bark.</p> +</div> + +<p>429. When such a stem grows on from year to year, it adds annually a +<span class="pagenum"><a name="Page_140" id="Page_140">[Pg 140]</a></span>layer of wood outside the preceding one, between that and the bark. +This is exogenous growth, or outside-growing, as the name denotes.</p> + +<div class="figcenter"> +<a name="Fig478" id="Fig478"></a> +<img src="images/fig478_480.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 478. Piece of a stem of Soft Maple, of a year old, +cut crosswise and lengthwise.</p> +<p><span class="smcap">Fig.</span> 479. A portion of the same, magnified.</p> +<p><span class="smcap">Fig.</span> 480. A small piece of the same, taken from one side, +reaching from the bark to the pith, and highly magnified: <i>a</i>, a small +bit of the pith; <i>b</i>, spiral ducts of what is called the <i>medullary +sheath</i>; <i>c</i>, the wood; <i>d</i>, <i>d</i>, dotted ducts in the wood; <i>e</i>, <i>e</i>, +annular ducts; <i>f</i>, the liber or inner bark; <i>g</i>, the green bark; <i>h</i>, +the corky layer; <i>i</i>, the skin, or epidermis; <i>j</i>, one of the medullary +rays, or plates of silver grain, seen on the cross-section.</p> +</div> + +<p>430. Some new bark is formed every year, as well as new wood, the former +inside, as the latter is outside of that of the year preceding. The ring +or zone of tender forming tissue between the bark and the wood has been +called the <i>Cambium Layer</i>. <i>Cambium</i> is an old name of the +physiologists for nutritive juice. And this thin layer is so gorged with +rich nutritive sap when spring growth is renewed, that the bark then +seems to be loose from the wood and a layer of viscid sap (or <i>cambium</i>) +to be poured out between the two. But there is all the while a +connection of the bark and the wood by delicate cells, rapidly +multiplying and growing.</p> + +<p><a name="UNIT_431" id="UNIT_431">431.</a> <b>The Bark</b> of a year-old stem consists of three parts, more or less +distinct, namely,—beginning next the wood,—</p> + +<p>1. The <span class="smcap">Liber</span> or <span class="smcap">Fibrous Bark</span>, the <i>Inner Bark</i>. This contains some +wood-cells, or their equivalent, commonly in the form of bast or +bast-cells (<a href="#UNIT_411">411</a>, Fig. <a href="#Fig444">444</a>), such as those of Basswood or Linden, and +among herbs those of flax and hemp, which are spun and woven or made +into cordage. It also contains cells which are named <i>sieve</i>-cells, on +account of numerous slits and pores in their walls, by which the +protoplasm of contiguous cells communicates. In woody stems, whenever a +new layer of wood is formed, some new liber or inner bark is also formed +outside of it.</p> + +<p><span class="pagenum"><a name="Page_141" id="Page_141">[Pg 141]</a></span></p><p>2. The <span class="smcap">Green Bark</span> or <i>Middle Bark</i>. This consists of cellular tissue +only, and contains the same green matter (<i>chlorophyll</i>, <a href="#UNIT_417">417</a>) as the +leaves. In woody stems, before the season's growth is completed, it +becomes covered by</p> + +<p>3. The <span class="smcap">Corky Layer</span> or <i>Outer Bark</i>, the cells of which contain no +chlorophyll, and are of the nature of <i>cork</i>. Common cork is the thick +corky layer of the bark of the Cork-Oak of Spain. It is this which gives +to the stems or twigs of shrubs and trees the aspect and the color +peculiar to each,—light gray in the Ash, purple in the Red Maple, red +in several Dogwoods, etc.</p> + +<p>4. The <span class="smcap">Epidermis</span>, or skin of the plant, consisting of a layer of +thick-sided empty cells, which may be considered to be the outermost +layer, or in most herbaceous stems the only layer, of cork-cells.</p> + +<div class="figcenter"> +<a name="Fig481" id="Fig481"></a> +<img src="images/fig481_482.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 481. Magnified view of surface of a bit of young +Maple wood from which the bark has been torn away, showing the +wood-cells and the bark-ends of medullary rays.</p> +<p><span class="smcap">Fig.</span> 482. Section in the opposite direction, from bark +(on the left) to beginning of pith (on the right), and a medullary ray +extending from one to the other.</p> +</div> + +<p>432. The green layer of bark seldom grows much after the first season. +Sometimes the corky layer grows and forms new layers, inside of the old, +for years, as in the Cork-Oak, the Sweet Gum-tree, and the White and the +Paper Birch. But it all dies after a while; and the continual +enlargement of the wood within finally stretches it more than it can +bear, and sooner or later cracks and rends it, while the weather acts +powerfully upon its surface; so the older bark perishes and falls away +piecemeal year by year.</p> + +<p>433. So on old trunks only the inner bark remains. This is renewed every +year from within and so kept alive, while the older and outer layers +die, are fissured and rent by the distending trunk, weathered and worn, +and thrown off in fragments,—in some trees slowly, so that the bark of +old trunks may acquire great thickness; in others, more rapidly. In +Honeysuckles and Grape-Vines, the layers of liber loosen and die when +only a year or two old. The annual layers of liber are sometimes as +distinct as those of the wood, but often not so.</p> + +<p><span class="pagenum"><a name="Page_142" id="Page_142">[Pg 142]</a></span></p><p><a name="UNIT_434" id="UNIT_434">434.</a> <b>The Wood</b> of an exogenous trunk, having the old growths covered by +the new, remains nearly unchanged in age, except from decay. Wherever +there is an annual suspension and renewal of growth, as in temperate +climates, the annual growths are more or less distinctly marked, in the +form of concentric rings on the cross section, so that the age of the +tree may be known by counting them. Over twelve hundred layers have been +counted on the stumps of Sequoias in California, and it is probable that +some trees now living antedate the Christian era.</p> + +<p>435. The reason why the annual growths are distinguishable is, that the +wood formed at the beginning of the season is more or less different in +the size or character of the cells from that of the close. In Oak, +Chestnut, etc., the first wood of the season abounds in dotted ducts, +the calibre of which is many times greater than that of the proper +wood-cells.</p> + +<p><a name="UNIT_436" id="UNIT_436">436.</a> <b>Sap-wood, or Alburnum.</b> This is the newer wood, living or recently +alive, and taking part in the conveyance of sap. Sooner or later, each +layer, as it becomes more and more deeply covered by the newer ones and +farther from the region of growth, is converted into</p> + +<p><a name="UNIT_437" id="UNIT_437">437.</a> <b>Heart-wood, or Duramen.</b> This is drier, harder, more solid, and much +more durable as timber, than sap-wood. It is generally of a different +color, and it exhibits in different species the hue peculiar to each, +such as reddish in Red-Cedar, brown in Black-Walnut, black in Ebony, +etc. The change of sap-wood into heart-wood results from the thickening +of the walls of the wood-cells by the deposition of hard matter, lining +the tubes and diminishing their calibre; and by the deposition of a +vegetable coloring-matter peculiar to each species. The heart-wood, +being no longer a living part, may decay, and often does so, without the +least injury to the tree, except by diminishing the strength of the +trunk, and so rendering it more liable to be overthrown.</p> + +<p><a name="UNIT_438" id="UNIT_438">438.</a> <b>The Living Parts of a Tree</b>, of the exogenous kind, are only these: +first, the rootlets at one extremity; second, the buds and leaves of the +season at the other; and third, a zone consisting of the newest wood and +the newest bark, connecting the rootlets with the buds or leaves, +however widely separated these may be,—in the tallest trees from two to +four hundred feet apart. And these parts of the tree are all renewed +every year. No wonder, therefore, that trees may live so long, since +they annually reproduce everything that is essential to their life and +growth, and since only a very small part of their bulk is alive at once. +The tree survives, but nothing now living has been so long. In it, as +elsewhere, life is a transitory thing, ever abandoning the <i>old</i>, and +renewed in the <i>young</i>.</p> + + +<h3><a name="Subsect_XVI_4" id="Subsect_XVI_4">§ 4.</a> ANATOMY OF LEAVES.</h3> + +<p>439. The wood in leaves is the framework of ribs, veins, and veinlets +(<a href="#UNIT_125">125</a>), serving not only to strengthen them, but also to bring in the +sap, and to distribute it throughout every part. The cellular portion is +the <span class="pagenum"><a name="Page_143" id="Page_143">[Pg 143]</a></span>green pulp, and is nearly the same as the green layer of the bark. +So that the leaf may properly enough be regarded as a sort of expansion +of the fibrous and green layers of the bark. It has no proper corky +layer; but the whole is covered by a transparent skin or <i>epidermis</i>, +resembling that of the stem.</p> + +<p>440. The cells of the leaf are of various forms, rarely so compact as to +form a close cellular tissue, usually loosely arranged, at least in the +lower part, so as to give copious intervening spaces or air passages, +communicating throughout the whole interior (Fig. <a href="#Fig443">443</a>, <a href="#Fig483">483</a>). The green +color is given by the chlorophyll (<a href="#UNIT_417">417</a>), seen through the very +transparent walls of the cells and through the translucent epidermis of +the leaf.</p> + +<div class="figcenter"> +<a name="Fig483" id="Fig483"></a> +<img src="images/fig483.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 483. Magnified section of a leaf of White Lily, to +exhibit the cellular structure, both of upper and lower stratum, the +air-passages of the lower, and the epidermis or skin, in section, also a +little of that of the lower face, with some of its stomates.</p> +</div> + +<p>441. In ordinary leaves, having an upper and under surface, the green +cells form two distinct strata, of different arrangement. Those of the +upper stratum are oblong or cylindrical, and stand endwise to the +surface of the leaf, usually close together, leaving hardly any vacant +spaces; those of the lower are commonly irregular in shape, most of them +with their longer diameter parallel to the face of the leaf, and are +very loosely arranged, leaving many and wide air-chambers. The green +color of the lower is therefore diluted, and paler than that of the +upper face of the leaf. The upper part of the leaf is so constructed as +to bear the direct action of the sunshine; the lower so as to afford +freer circulation of air, and to facilitate transpiration. It +communicates more directly than the upper with the external air by means +of <i>Stomates</i>.</p> + +<p><a name="UNIT_442" id="UNIT_442">442.</a> <b>The Epidermis</b> or skin of leaves and all young shoots is best seen +in the foliage. It may readily be stripped off from the surface of a +Lily-leaf, and still more so from more fleshy and soft leaves, such as +those <span class="pagenum"><a name="Page_144" id="Page_144">[Pg 144]</a></span>of Houseleek. The epidermis is usually composed of a single +layer, occasionally of two or three layers, of empty cells, mostly of +irregular outline. The sinuous lines which traverse it, and may be +discerned under low powers of the microscope (Fig. <a href="#Fig484">487</a>), are the +boundaries of the epidermal cells.</p> + +<div class="figcenter"> +<a name="Fig484" id="Fig484"></a> +<img src="images/fig484_487.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 484. Small portion of epidermis of the lower face of +a White-Lily leaf, with stomata.</p> +<p><span class="smcap">Fig.</span> 485. One of these, more magnified, in the closed +state. 486. Another stoma, open.</p> +<p><span class="smcap">Fig.</span> 487. Small portion of epidermis of the Garden +Balsam, highly magnified, showing very sinuous-walled cells, and three +stomata.</p> +</div> + +<p><a name="UNIT_443" id="UNIT_443">443.</a> <b>Breathing-pores, or Stomates, Stomata</b> (singular, a +<i>Stoma</i>,—literally, a mouth) are openings through the epidermis into +the air-chambers or intercellular passages, always between and guarded +by a pair of thin-walled guardian cells. Although most abundant in +leaves, especially on their lower face (that which is screened from +direct sunlight), they are found on most other green parts. They +establish a direct communication between the external air and that in +the loose interior of the leaf. Their guardian cells or lips, which are +soft and delicate, like those of the green pulp within, by their greater +or less turgidity open or close the orifice as the moisture or dryness +varies.</p> + +<p>444. In the White Lily the stomata are so remarkably large that they may +be seen by a simple microscope of moderate power, and may be discerned +even by a good hand lens. There are about 60,000 of them to the square +inch of the epidermis of the lower face of this Lily-leaf, and only +about 3000 to the same space on the upper face. It is computed that an +average leaf of an Apple-tree has on its lower face about 100,000 of +these mouths.</p> + + +<h3><a name="Subsect_XVI_5" id="Subsect_XVI_5">§ 5.</a> PLANT FOOD AND ASSIMILATION.</h3> + +<p>445. Only plants are capable of originating organizable matter, or the +materials which compose the structure of vegetables and animals. The +essential and peculiar work of plants is to take up portions of earth +and air (water belonging to both) upon which animals cannot live at all, +and to convert them into something organizable; that is, into something +that, under life, may be built up into vegetable and animal structures. +All the food of animals is produced by plants. Animals live upon +vegetables, <span class="pagenum"><a name="Page_145" id="Page_145">[Pg 145]</a></span>directly or at second hand, the carnivorous upon the +herbivorous; and vegetables live upon earth and air, immediately or at +second hand.</p> + +<p><a name="UNIT_446" id="UNIT_446">446.</a> <b>The Food of plants</b>, then, primarily, is earth and air. This is +evident enough from the way in which they live. Many plants will +flourish in pure sand or powdered chalk, or on the bare face of a rock +or wall, watered merely with rain. And almost any plant may be made to +grow from the seed in moist sand, and increase its weight many times, +even if it will not come to perfection. Many naturally live suspended +from the branches of trees high in the air, and nourished by it alone, +never having any connection with the soil; and some which naturally grow +on the ground, like the Live-forever of the gardens, when pulled up by +the roots and hung in the air will often flourish the whole summer long.</p> + +<p>447. It is true that fast-growing plants, or those which produce much +vegetable matter in one season (especially in such concentrated form as +to be useful as food for man or the higher animals) will come to +maturity only in an enriched soil. But what is a rich soil? One which +contains decomposing vegetable matter, or some decomposing animal +matter; that is, in either case, some decomposing organic matter +formerly produced by plants. Aided by this, grain-bearing and other +important vegetables will grow more rapidly and vigorously, and make a +greater amount of nourishing matter, than they could if left to do the +whole work at once from the beginning. So that in these cases also all +the organic or organizable matter was made by plants, and made out of +earth and air. Far the larger and most essential part was air and water.</p> + +<p>448. Two kinds of material are taken in and used by plants; of which the +first, although more or less essential to perfect plant-growth, are in a +certain sense subsidiary, if not accidental, viz.:—</p> + +<p><i>Earthy constituents</i>, those which are left in the form of ashes when a +leaf or a stick of wood is burned in the open air. These consist of some +<i>potash</i> (or <i>soda</i> in a marine plant), some <i>silex</i> (the same as +flint), and a little <i>lime</i>, <i>alumine</i>, or <i>magnesia</i>, <i>iron</i> or +<i>manganese</i>, <i>sulphur</i>, <i>phosphorus</i>, etc.,—some or all of these in +variable and usually minute proportions. They are such materials as +happen to be dissolved, in small quantity, in the water taken up by the +roots; and when that is consumed by the plant, or flies off pure (as it +largely does) by exhalation, the earthy matter is left behind in the +cells,—just as it is left incrusting the sides of a teakettle in which +much hard water has been boiled. Naturally, therefore, there is more +earthy matter (i. e. more ashes) in the leaves than in any other part +(sometimes as much as seven per cent, when the wood contains only two +per cent); because it is through the leaves that most of the water +escapes from the plant. Some of this earthy matter incrusts the +cell-walls, some goes to form crystals or rhaphides, which abound in +many plants (<a href="#UNIT_422">422</a>), some enters into certain special vegetable products, +and some appears to be necessary to the well-being of the higher orders +of plants, although forming no necessary part of the proper vegetable +structure.</p> + +<p><span class="pagenum"><a name="Page_146" id="Page_146">[Pg 146]</a></span></p><p><i>The essential constituents</i> of the organic fabric are those which are +dissipated into air and vapor in complete burning. They make up from 88 +to 99 per cent of the leaf or stem, and essentially the whole both of +the cellulose of the walls and the protoplasm of the contents. Burning +gives these materials of the plant's structure back to the air, mainly +in the same condition in which the plant took them, the same condition +which is reached more slowly in natural decay. The chemical elements of +the cell-walls (or cellulose, <a href="#UNIT_402">402</a>), as also of starch, sugar, and all +that class of organizable cell-material, are carbon, hydrogen, and +oxygen (<a href="#UNIT_399">399</a>). The same, with nitrogen, are the constituents of +protoplasm, or the truly vital part of vegetation.</p> + +<p>449. These chemical elements out of which organic matters are composed +are supplied to the plant by water, carbonic acid, and some combinations +of nitrogen.</p> + +<p><i>Water</i>, far more largely than anything else, is imbibed by the roots; +also more or less by the foliage in the form of vapor. Water consists of +oxygen and hydrogen; and cellulose or plant-wall, starch, sugar, etc., +however different in their qualities, agree in containing these two +elements in the same relative proportions as in water.</p> + +<p><i>Carbonic acid</i> gas (Carbon dioxide) is one of the components of the +atmosphere,—a small one, ordinarily only about 1/2500 of its +bulk,—sufficient for the supply of vegetation, but not enough to be +injurious to animals, as it would be if accumulated. Every current or +breeze of air brings to the leaves expanded in it a succession of fresh +atoms of carbonic acid, which it absorbs through its multitudinous +breathing-pores. This gas is also taken up by water. So it is brought to +the ground by rain, and is absorbed by the roots of plants, either as +dissolved in the water they imbibe, or in the form of gas in the +interstices of the soil. Manured ground, that is, soil containing +decomposing vegetable or animal matters, is constantly giving out this +gas into the interstices of the soil, whence the roots of the growing +crop absorb it. Carbonic acid thus supplied, primarily from the air, is +the source of the carbon which forms much the largest part of the +substance of every plant. The proportion of carbon may be roughly +estimated by charring some wood or foliage; that is, by heating it out +of contact with the air, so as to decompose and drive off all the other +constituents of the fabric, leaving the large bulk of charcoal or carbon +behind.</p> + +<p><i>Nitrogen</i>, the remaining plant-element, is a gas which makes up more +than two thirds of the atmosphere, is brought into the foliage and also +to the roots (being moderately soluble in water) in the same ways as is +carbonic acid. The nitrogen which, mixed with oxygen, a little carbonic +acid, and vapor of water, constitutes the air we breathe, is the source +of this fourth plant-element. But it is very doubtful if ordinary plants +can use any nitrogen gas directly as food; that is, if they can directly +cause it to combine with the other elements so as to form protoplasm. +But when combined with hydrogen (forming ammonia), or when combined with +oxygen <span class="pagenum"><a name="Page_147" id="Page_147">[Pg 147]</a></span>(nitric acid and nitrates) plants appropriate it with avidity. +And several natural processes are going on in which nitrogen of the air +is so combined and supplied to the soil in forms directly available to +the plant. The most efficient is <i>nitrification</i>, the formation of nitre +(nitrate of potash) in the soil, especially in all fertile soils, +through the action of a bacterial ferment.</p> + +<p><a name="UNIT_450" id="UNIT_450">450.</a> <b>Assimilation</b> in plants is the conversion of these inorganic +substances—essentially, water, carbonic acid, and some form of combined +or combinable nitrogen—into vegetable matter. This most dilute food the +living plant concentrates and assimilates to itself. Only plants are +capable of converting these mineral into organizable matters; and this +all-important work is done by them (so far as all ordinary vegetation is +concerned) only</p> + +<p>451. <i>Under the light of the sun, acting upon green parts or foliage</i>, +that is, upon the chlorophyll, or upon what answers to chlorophyll, +which these parts contain. The sun in some way supplies a power which +enables the living plant to originate these peculiar chemical +combinations,—to organize matter into forms which are alone capable of +being endowed with life. The proof of this proposition is simple; and it +shows at the same time, in the simplest way, what a plant does with the +water and carbonic acid it consumes. Namely, 1st, it is only in sunshine +or bright daylight that the green parts of plants give out oxygen +gas,—then they regularly do so; and 2d, the giving out of this oxygen +gas is required to render the chemical composition of water and carbonic +acid the same as that of <i>cellulose</i>, that is, of the plant's permanent +fabric. This shows why plants spread out so large a surface of foliage. +Leaves are so many workshops, full of machinery worked by sun-power. The +emission of oxygen gas from any sun-lit foliage is seen by placing some +of this under water, or by using an aquatic plant, by collecting the air +bubbles which rise, and by noting that a taper burns brighter in this +air. Or a leafy plant in a glass globe may be supplied with a certain +small percentage of carbonic acid gas, and after proper exposure to +sunshine, the air on being tested will be found to contain less carbonic +acid and just so much the more oxygen gas.</p> + +<p>452. Now if the plant is making cellulose or any equivalent +substance,—that is, is making the very materials of its fabric and +growth, as must generally be the case,—all this oxygen gas given off by +the leaves comes from the decomposition of carbonic acid taken in by the +plant. For cellulose, and also starch, dextrine, sugar, and the like are +composed of carbon along with oxygen and hydrogen in just the +proportions to form water. And the carbonic acid and water taken in, +less the oxygen which the carbon brought with it as carbonic acid, and +which is given off from the foliage in sunshine, just represents the +manufactured article, cellulose.</p> + +<p>453. It comes to the same if the first product of assimilation is sugar, +or dextrine which is a sort of soluble starch, or starch itself. And in +the plant all these forms are readily changed into one another. In the +tiny seedling, as fast as this assimilated matter is formed it is used +in growth, that is, in the formation of cell-walls. After a time some or +much of <span class="pagenum"><a name="Page_148" id="Page_148">[Pg 148]</a></span>the product may be accumulated in store for future growth, as +in the root of the turnip, or the tuber of the potato, or the seed of +corn or pulse. This store is mainly in the form of starch. When growth +begins anew, this starch is turned into dextrine or into sugar, in +liquid form, and used to nourish and build up the germinating embryo or +the new shoot, where it is at length converted into cellulose and used +to build up plant-structure.</p> + +<p>454. But that which builds plant-fabric is not the cellular structure +itself; the work is done by the living protoplasm which dwells within +the walls. This also has to take and to assimilate its proper food, for +its own maintenance and growth. Protoplasm assimilates, along with the +other three elements, the nitrogen of the plant's food. This comes +primarily from the vast stock in the atmosphere, but mainly through the +earth, where it is accumulated through various processes in a fertile +soil,—mainly, so far as concerns crops, from the decomposition of +former vegetables and animals. This protoplasm, which is formed at the +same time as the simpler cellulose, is essentially the same as the flesh +of animals, and the source of it. It is the common basis of vegetable +and of animal life.</p> + +<p>455. <i>So plant-assimilation produces all the food and fabric of +animals.</i> Starch, sugar, the oils (which are, as it were, these +farinaceous matters more deoxidated), chlorophyll, and the like, and +even cellulose itself, form the food of herbivorous animals and much of +the food of man. When digested they enter into the blood, undergo +various transformations, and are at length decomposed into carbonic acid +and water, and exhaled from the lungs in respiration,—in other words, +are given back to the air by the animal as the very same materials which +the plant took from the air as its food,—are given back to the air in +the same form that they would have taken if the vegetable matter had +been left to decay where it grew, or if it had been set on fire and +burned; and with the same result, too, as to the heat,—the heat in this +case producing and maintaining the proper temperature of the animal.</p> + +<p>456. The protoplasm and other products containing nitrogen (gluten, +legumine, etc.), and which are most accumulated in grains and seeds (for +the nourishment of their embryos when they germinate), compose the most +nutritious vegetable food consumed by animals; they form their proper +flesh and sinews, while the earthy constituents of the plant form the +earthy matter of the bones, etc. At length decomposed, in the secretions +and excretions, these nitrogenous constituents are through successive +changes finally resolved into mineral matter, into carbonic acid, water, +and ammonia or some nitrates,—into exactly or essentially the same +materials which the plants took up and assimilated. Animals depend upon +vegetables absolutely and directly for their subsistence; also +indirectly, because</p> + +<p>457. <i>Plants purify the air for animals.</i> In the very process by which +they create food they take from the air carbonic acid gas, injurious to +animal respiration, which is continually poured into it by the breathing +of all animals, by all decay, by the burning of fuel and all other +ordinary combustion; and <span class="pagenum"><a name="Page_149" id="Page_149">[Pg 149]</a></span>they restore an equal bulk of life-sustaining +oxygen needful for the respiration of animals,—needful, also, in a +certain measure, for plants in any work they do. For in plants, as well +as in animals, work is done at a certain cost.</p> + + +<h3><a name="Subsect_XVI_6" id="Subsect_XVI_6">§ 6.</a> PLANT WORK AND MOVEMENT.</h3> + +<p>458. As the organic basis and truly living material of plants is +identical with that of animals, so is the life at bottom essentially the +same; but in animals something is added at every rise from the lowest to +highest organisms. Action and work in living beings require movement.</p> + +<p>459. Living things move; those not living are only moved. Plants move as +truly as do animals. The latter, nourished as they are upon organized +food, which has been prepared for them by plants, and is found only here +and there, must needs have the power of going after it, of collecting +it, or at least of taking it in; which requires them to make spontaneous +movements. But ordinary plants, with their wide-spread surface, always +in contact with the earth and air on which they feed,—the latter +everywhere the same, and the former very much so,—might be thought to +have no need of movement. Ordinary plants, indeed, have no locomotion; +some float, but most are rooted to the spot where they grew. Yet +probably all of them execute various movements which must be as truly +self-caused as are those of the lower grades of animals,—movements +which are overlooked only because too slow to be directly observed. +Nevertheless, the motion of the hour-hand and of the minute-hand of a +watch is not less real than that of the second-hand.</p> + +<div class="figcenter"> +<a name="Fig488" id="Fig488"></a> +<img src="images/fig488.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 488. Two individuals of an Oscillaria, magnified.</p> +</div> + +<p><a name="UNIT_460" id="UNIT_460">460.</a> <b>Locomotion.</b> Moreover, many microscopic plants living in water are +seen to move freely, if not briskly, under the microscope; and so +likewise do more conspicuous aquatic plants in their embryo-like or +seedling state. Even at maturity, species of Oscillaria (such as in Fig. +<a href="#Fig488">488</a>, minute worm-shaped plants of fresh waters, taking this name from +their oscillating motions) freely execute three different kinds of +movement, the very delicate investing coat of cellulose not impeding the +action of the living protoplasm within. Even when this coat is firmer +and hardened with a siliceous deposit, such crescent-shaped or +boat-shaped one-celled plants as <i>Closterium</i> or <i>Naricula</i> are able in +some way to move along from place to place in the water.</p> + +<div class="figcenter"> +<a name="Fig489" id="Fig489"></a> +<img src="images/fig489.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 489. A few cells of a leaf of Naias flexilis, highly +magnified: the arrows indicate the courses of the circulating currents.</p> +</div> + +<p><a name="UNIT_461" id="UNIT_461">461.</a> <b>Movements in Cells</b>, <b>or Cell-circulation</b>, sometimes called +<i>Cyclosis</i>, has been detected in so many plants, especially in +comparatively <span class="pagenum"><a name="Page_150" id="Page_150">[Pg 150]</a></span>transparent aquatic plants and in hairs on the surface of +land plants (where it is easiest to observe), that it may be inferred to +take place in all cells during the most active part of their life. This +motion is commonly a streaming movement of threads of protoplasm, +carrying along solid granules by which the action may be observed and +the rate measured, or in some cases it is a rotation of the whole +protoplasmic contents of the cell. A comparatively low magnifying power +will show it in the cells of Nitella and Chara (which are cryptogamous +plants); and under a moderate power it is well seen in the Tape Grass of +fresh water, Vallisneria, and in Naias flexilis (Fig. <a href="#Fig489">489</a>). Minute +particles and larger greenish globules are seen to be carried along, as +if in a current, around the cell, passing up one side, across the end, +down the other and across the bottom, completing the circuit sometimes +within a minute or less when well warmed. To see it well in the cell, +which like a string of beads form the hairs on the stamens of +Spiderwort, a high magnifying power is needed.</p> + +<p><a name="UNIT_462" id="UNIT_462">462.</a> <b>Transference of Liquid from Cell to Cell</b>, and so from place to +place in the plant, the absorption of water by the rootlets, and the +exhalation of the greater part of it from the foliage,—these and +similar operations are governed by the physical laws which regulate the +diffusion of fluids, but are controlled by the action of living +protoplasm. Equally under vital control are the various chemical +transformations which attend assimilation and growth, and which involve +not only molecular movements but conveyance. Growth itself, which is the +formation and shaping of new parts, implies the direction of internal +activities to definite ends.</p> + +<p><a name="UNIT_463" id="UNIT_463">463.</a> <b>Movements of Organs.</b> The living protoplasm, in all but the lowest +grade of plants, is enclosed and to common appearance isolated in +separate cells, the walls of which can only in their earliest state be +said to be alive. Still plants are able to cause the protoplasm of +adjacent cells to act in concert, and by their combined action to effect +movements in roots, stems, or leaves, some of them very slow and +gradual, some manifest and striking. Such movements are brought about +through individually minute changes in the form or tension in the +protoplasm of the innumerable cells which make up the structure of the +organ. Some of the slower movements are effected during growth, and may +be explained by inequality of growth on the two sides of the bending +organ. But the more rapid changes of position, and some of the slow +ones, cannot be so explained.</p> + +<p><span class="pagenum"><a name="Page_151" id="Page_151">[Pg 151]</a></span></p><p><a name="UNIT_464" id="UNIT_464">464.</a> <b>Root-movements.</b> In its growth a root turns or bends away from the +light and toward the centre of the earth, so that in lengthening it +buries itself in the soil where it is to live and act. Every one must +have observed this in the germination of seeds. Careful observations +have shown that the tip of a growing root also makes little sweeps or +short movements from side to side. By this means it more readily +insinuates itself into yielding portions of the soil. The root-tips will +also turn toward moisture, and so secure the most favorable positions in +the soil.</p> + +<p><a name="UNIT_465" id="UNIT_465">465.</a> <b>Stem-movements.</b> The root end of the caulicle or first joint of stem +(that below the cotyledons) acts like the root, in turning downward in +germination (making a complete bend to do so if it happens to point +upward as the seed lies in the ground), while the other end turns or +points skyward. These opposite positions are taken in complete darkness +as readily as in the light, in dryness as much as in moisture: +therefore, so far as these movements are physical, the two portions of +the same internode appear to be oppositely affected by gravitation or +other influences.</p> + +<p>466. Rising into the air, the stem and green shoots generally, while +young and pliable, bend or direct themselves toward the light, or toward +the stronger light when unequally illuminated; while roots turn toward +the darkness.</p> + +<p>467. Many growing stems have also a movement of <i>Nutation</i>, that is, of +nodding successively in different directions. This is brought about by a +temporary increase of turgidity of the cells along one side, thus bowing +the stem over to the opposite side; and this line of turgescence travels +round the shoot continually, from right to left or from left to right +according to the species: thus the shoot bends to all points of the +compass in succession. Commonly this nutation is slight or hardly +observable. It is most marked in</p> + +<p><a name="UNIT_468" id="UNIT_468">468.</a> <b>Twining Stems</b> (Fig. <a href="#Fig90">90</a>). The growing upper end of such stems, as is +familiar in the Hop, Pole Beans, and Morning-Glory, turns over in an +inclined or horizontal direction, thus stretching out to reach a +neighboring support, and by the continual change in the direction of the +nodding, sweeps the whole circle, the sweeps being the longer as the +stem lengthens. When it strikes against a support, such as a stem or +branch of a neighboring plant, the motion is arrested at the contact, +but continues at the growing apex beyond, and this apex is thus made to +wind spirally around the supporting body.</p> + +<p><a name="UNIT_469" id="UNIT_469">469.</a> <b>Leaf-movements</b> are all but universal. The presentation by most +leaves of their upper surface to the light, from whatever direction that +may come, is an instance; for when turned upside down they twist or bend +round on the stalk to recover this normal position. Leaves, and the +leaflets of compound leaves, change this position at nightfall, or when +the light is withdrawn; they then take what is called their sleeping +posture, resuming the diurnal position when daylight returns. This is +very striking <span class="pagenum"><a name="Page_152" id="Page_152">[Pg 152]</a></span>in Locust-trees, in the Sensitive Plant (Fig. <a href="#Fig490">490</a>), and +in Woodsorrel. Young seedlings droop or close their leaves at night in +plants which are not thus affected in the adult foliage. All this is +thought to be a protection against the cold by nocturnal radiation.</p> + +<p>470. Various plants climb by a coiling movement of their leaves or their +leaf-stalks. Familiar examples are seen in Clematis, Maurandia, +Tropæolum, and in a Solanum which is much cultivated in greenhouses +(Fig. <a href="#Fig172">172</a>). In the latter, and in other woody plants which climb in this +way, the petioles thicken and harden after they have grasped their +support, thus securing a very firm hold.</p> + +<p><a name="UNIT_471" id="UNIT_471">471.</a> <b>Tendril movements.</b> Tendrils are either leaves or stems (<a href="#UNIT_98">98</a>, <a href="#UNIT_168">168</a>), +specially developed for climbing purposes. Cobæa is a good example of +partial transformation; some of the leaflets are normal, some of the +same leaf are little tendrils, and some intermediate in character. The +Passion-flowers give good examples of simple stem-tendrils (Fig. <a href="#Fig92">92</a>); +Grape-Vines, of branched ones. Most tendrils make revolving sweeps, like +those of twining stems. Those of some Passion-flowers, in sultry +weather, are apt to move fast enough for the movement actually to be +seen for a part of the circuit, as plainly as that of the second-hand of +a watch. Two herbaceous species, Passiflora gracilis and P. sicyoides +(the first an annual, the second a strong-rooted perennial of the +easiest cultivation), are admirable for illustration both of revolving +movements and of sensitive coiling.</p> + +<div class="figcenter"> +<a name="Fig490" id="Fig490"></a> +<img src="images/fig490.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 490. Piece of stem of Sensitive Plant (Mimosa +pudica), with two leaves, the lower open, the upper in the closed +state.</p> +</div> + +<p><a name="UNIT_472" id="UNIT_472">472.</a> <b>Movements under Irritation.</b> The most familiar case is that of the +Sensitive Plant (Fig. <a href="#Fig490">490</a>). The leaves suddenly take their nocturnal +position when roughly touched or when shocked by a jar. The leaflets +close in pairs, the four outspread partial petioles come closer +together, and the common petiole is depressed. The seat of the movements +is at the base of the leaf-stalk and stalklets. Schrankia, a near +relative of the Sensitive Plant, acts in the same way, but is slower. +These are not anomalous actions, but only extreme manifestations of a +faculty more or less common in foliage. In Locust and Honey-Locusts for +example, repeated jars will slowly produce similar effects.</p> + +<p><span class="pagenum"><a name="Page_153" id="Page_153">[Pg 153]</a></span></p><p>473. Leaf-stalks and tendrils are adapted to their uses in climbing by +a similar sensitiveness. The coiling of the leaf-stalk is in response to +a kind of irritation produced by contact with the supporting body. This +may be shown by gentle rubbing or prolonged pressure upon the upper face +of the leaf-stalk, which is soon followed by a curvature. Tendrils are +still more sensitive to contact or light friction. This causes the free +end of the tendril to coil round the support, and the sensitiveness, +propagated downward along the tendril, causes that side of it to become +less turgescent or the opposite side more so, thus throwing the tendril +into coils. This shortening draws the plant up to the support. Tendrils +which have not laid hold will at length commonly coil spontaneously, in +a simple coil, from the free apex downward. In Sicyos, Echinocystis, and +the above mentioned Passion-flowers (<a href="#UNIT_471">471</a>), the tendril is so sensitive, +under a high summer temperature, that it will curve and coil promptly +after one or two light strokes by the hand.</p> + +<div class="figcenter"> +<a name="Fig491" id="Fig491"></a> +<img src="images/fig491.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 491. Portion of stem and leaves of Telegraph-plant +(Desmodium gyrans), almost of natural size.</p> +</div> + +<p>474. Among spontaneous movements the most singular are those of +Desmodium gyrans of India, sometimes called Telegraph-plant, which is +cultivated on account of this action. Of its three leaflets, the larger +(terminal) one moves only by drooping at nightfall and rising with the +dawn. But its two small lateral leaflets, when in a congenial high +temperature, by day and by night move upward and downward in a +succession of jerks, stopping occasionally, as if to recover from +exhaustion. In most plant-movements some obviously useful purpose is +subserved: this of Desmodium gyrans is a riddle.</p> + +<p><a name="UNIT_475" id="UNIT_475">475.</a> <b>Movements in Flowers</b> are very various. The most remarkable are in +some way connected with fertilization (Sect. <a href="#Section_XIII">XIII.</a>). Some occur under +irritation: the stamens of Barberry start forward when touched at the +base inside: those of many polyandrous flowers (of Sparmannia very +strikingly) spread outwardly when lightly brushed: the two lips or lobes +<span class="pagenum"><a name="Page_154" id="Page_154">[Pg 154]</a></span>of the stigma in Mimulus close after a touch. Some are automatic and +are connected with dichogamy (<a href="#UNIT_339">339</a>): the style of Sabbatia and of +large-flowered species of Epilobium bends over strongly to one side or +turns downward when the blossom opens, but slowly erects itself a day or +two later.</p> + +<p><a name="UNIT_476" id="UNIT_476">476.</a> <b>Extraordinary Movements connected with Capture of Insects.</b> The most +striking cases are those of Drosera and Dionæa; for an account of which +see "How Plants Behave," and Goodale's "Physiological Botany."</p> + +<p>477. The upper face of the leaves of the common species of Drosera, or +Sundew, is beset with stout bristles, having a glandular tip. This tip +secretes a drop of a clear but very viscid liquid, which glistens like a +dew-drop in the sun; whence the popular name. When a fly or other small +insect, attracted by the liquid, alights upon the leaf, the viscid drops +are so tenacious that they hold it fast. In struggling it only becomes +more completely entangled. Now the neighboring bristles, which have not +been touched, slowly bend inward from all sides toward the captured +insect, and bring their sticky apex against its body, thus increasing +the number of bonds. Moreover, the blade of the leaf commonly aids in +the capture by becoming concave, its sides or edges turning inward, +which brings still more of the gland-tipped bristles into contact with +the captive's body. The insect perishes; the clear liquid disappears, +apparently by absorption into the tissue of the leaf. It is thought that +the absorbed secretion takes with it some of the juices of the insect or +the products of its decomposition.</p> + +<div class="figcenter"> +<a name="Fig492" id="Fig492"></a> +<img src="images/fig492.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 492. Plant of Dionæa muscipula, or Venus's Fly-trap, +reduced in size.</p> +</div> + +<p>478. Dionæa muscipula, the most remarkable vegetable fly-trap (Fig. <a href="#Fig176">176</a>, +<a href="#Fig492">492</a>), is related to the Sundews, and has a more special and active +apparatus for fly-catching, formed of the summit of the leaf. The two +halves of this rounded body move as if they were hinged upon the midrib; +their edges are fringed with spiny but not glandular bristles, which +interlock when the organ closes. Upon the face are two or three short +and delicate bristles, which are sensitive. They do not themselves move +when touched, but they propagate the sensitiveness to the organ itself, +causing it to close with a quick movement. In a fresh <span class="pagenum"><a name="Page_155" id="Page_155">[Pg 155]</a></span>and vigorous +leaf, under a high summer temperature, and when the trap lies widely +open, a touch of any one of the minute bristles on the face, by the +finger or any extraneous body, springs the trap (so to say), and it +closes suddenly; but after an hour or so it opens again. When a fly or +other small insect alights on the trap, it closes in the same manner, +and so quickly that the intercrossing marginal bristles obstruct the +egress of the insect, unless it be a small one and not worth taking. +Afterwards and more slowly it completely closes, and presses down upon +the prey; then some hidden glands pour out a glairy liquid, which +dissolves out the juices of the insect's body; next all is re-absorbed +into the plant, and the trap opens to repeat the operation. But the same +leaf perhaps never captures more than two or three insects. It ages +instead, becomes more rigid and motionless, or decays away.</p> + +<p>479. That some few plants should thus take animal food will appear less +surprising when it is considered that hosts of plants of the lower +grade, known as Fungi, moulds, rusts, ferments, Bacteria, etc., live +upon animal or other organized matter, either decaying or living. That +plants should execute movements in order to accomplish the ends of their +existence is less surprising now when it is known that the living +substance of plants and animals is essentially the same; that the beings +of both kingdoms partake of a common life, to which, as they rise in the +scale, other and higher endowments are successively superadded.</p> + +<p><a name="UNIT_480" id="UNIT_480">480.</a> <b>Work uses up material and energy</b> in plants as well as in animals. +The latter live and work by the consumption and decomposition of that +which plants have assimilated into organizable matter through an energy +derived from the sun, and which is, so to say, stored up in the +assimilated products. In every internal action, as well as in every +movement and exertion, some portion of this assimilated matter is +transformed and of its stored energy expended. The steam-engine is an +organism for converting the sun's radiant energy, stored up by plants in +the fuel, into mechanical work. An animal is an engine fed by vegetable +fuel in the same or other forms, from the same source, by the +decomposition of which it also does mechanical work. The plant is the +producer of food and accumulator of solar energy or force. But the +plant, like the animal, is a consumer whenever and by so much as it does +any work except its great work of assimilation. Every internal change +and movement, every transformation, such as that of starch into sugar +and of sugar into cell-walls, as well as every movement of parts which +becomes externally visible, is done at the expense of a certain amount +of its assimilated matter and of its stored energy; that is, by the +decomposition or combustion of sugar or some such product into carbonic +acid and water, which is given back to the air, just as in the animal it +is given back to the air in respiration. So the respiration of plants is +as real and as essential as that of animals. But what plants consume or +decompose in their life and action is of insignificant amount in +comparison with what they compose.</p> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_156" id="Page_156">[Pg 156]</a></span></p> +<h2><a name="Section_XVII" id="Section_XVII"></a><span class="smcap">Section XVII.</span> CRYPTOGAMOUS OR FLOWERLESS PLANTS.</h2> + + +<p>481. Even the beginner in botany should have some general idea of what +cryptogamous plants are, and what are the obvious distinctions of the +principal families. Although the lower grades are difficult, and need +special books and good microscopes for their study, the higher orders, +such as Ferns, may be determined almost as readily as phanerogamous +plants.</p> + +<p>482. Linnæus gave to this lower grade of plants the name of +<i>Cryptogamia</i>, thereby indicating that their organs answering to stamens +and pistils, if they had any, were recondite and unknown. There is no +valid reason why this long-familiar name should not be kept up, along +with the counterpart one of <i>Phanerogamia</i> (<a href="#UNIT_6">6</a>), although organs +analogous to stamens and pistil, or rather to pollen and ovule, have +been discovered in all the higher and most of the lower grades of this +series of plants. So also the English synonymous name of <i>Flowerless +Plants</i> is both good and convenient: for they have not flowers in the +proper sense. The essentials of flowers are stamens and pistils, giving +rise to seeds, and the essential of a seed is an embryo (<a href="#UNIT_8">8</a>). +Cryptogamous or Flowerless plants are propagated by <span class="smcap">Spores</span>; and a spore +is not an embryo-plantlet, but mostly a single plant-cell (<a href="#UNIT_399">399</a>).</p> + +<p><a name="UNIT_483" id="UNIT_483">483.</a> <b>Vascular Cryptogams</b>, which compose the higher orders of this series +of plants, have stems and (usually) leaves, constructed upon the general +plan of ordinary plants; that is, they have wood (wood-cells and +vessels, <a href="#UNIT_408">408</a>) in the stem and leaves, in the latter as a frame work of +veins. But the lower grades, having only the more elementary cellular +structure, are called <i>Cellular Cryptogams</i>. Far the larger number of +the former are Ferns: wherefore that class has been called</p> + +<p><a name="UNIT_484" id="UNIT_484">484.</a> <b>Pteridophyta, Pteridophytes</b> in English form, meaning +<i>Fern-plants</i>,—that is, Ferns and their relatives. They are mainly +Horsetails, Ferns, Club-Mosses, and various aquatics which have been +called <i>Hydropterides</i>, i. e. Water-Ferns.</p> + +<p><a name="UNIT_485" id="UNIT_485">485.</a> <b>Horsetails</b>, <i>Equisetaceæ</i>, is the name of a family which consists +only (among now-living plants) of <i>Equisetum</i>, the botanical name of +Horsetail and Scouring Rush. They have hollow stems, with partitions at +the nodes; the leaves consist only of a whorl of scales at each node, +these coalescent into a sheath: from the axils of these leaf-scales, in +many species, branches grow out, which are similar to the stem but on a +much smaller scale, close-jointed, and with the tips of the leaves more +apparent. At the apex of the stem appears the <i>fructification</i>, as it is +called for lack of a better term, in the form of a short spike or head. +This consists of a good number of stalked shields, bearing on their +inner or under face several wedge-shaped spore-cases. The spore-cases +when they ripen open down the inner <span class="pagenum"><a name="Page_157" id="Page_157">[Pg 157]</a></span>side and discharge a great number +of green spores of a size large enough to be well seen by a hand-glass. +The spores are aided in their discharge and dissemination by four +club-shaped threads attached to one part of them. These are hygrometric: +when moist they are rolled up over the spore; when dry they straighten, +and exhibit lively movements, closing over the spore when breathed upon, +and unrolling promptly a moment after as they dry. (See Fig. <a href="#Fig493">493-498</a>.)</p> + +<div class="figcenter"> +<a name="Fig493" id="Fig493"></a> +<img src="images/fig493_498.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 493. Upper part of a stem of a Horsetail, Equisetum +sylvaticum. 494. Part of the head or spike of spore-cases, with some of +the latter taken off. 495. View (more enlarged) of under side of the +shield-shaped body, bearing a circle of spore-cases. 496. One of the +latter detached and more magnified. 497. A spore with the attached arms +moistened. 498. Same when dry, the arms extended.</p> +</div> + +<div class="figcenter"> +<a name="Fig499" id="Fig499"></a> +<img src="images/fig499.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 499. A Tree-Fern, Dicksonia arborescens, with a +young one near its base. In front a common herbaceous Fern (Polypodium +vulgare) with its creeping stem or rootstock.</p> +</div> + +<div class="figcenter"> +<a name="Fig500" id="Fig500"></a> +<img src="images/fig500.jpg" alt="" /><br /> +<span class="smcap">Fig.</span> 500. A section of the trunk of a Tree-Fern. +</div> + +<p><a name="UNIT_486" id="UNIT_486">486.</a> <b>Ferns, or Filices</b>, a most attractive family of plants, are very +numerous and varied. In warm and equable climates some rise into +forest-trees, with habit of Palms; but most of them are perennial herbs. +The wood of a Fern-trunk is very different, however, from that of a +palm, or of any exogenous stem either. A section is represented in Fig. +<a href="#Fig500">500</a>. The curved plates of wood each terminate <span class="pagenum"><a name="Page_158" id="Page_158">[Pg 158]</a></span>upward in a leaf-stalk. +The subterranean trunk or stem of any strong-growing herbaceous Fern +shows a similar structure. Most Ferns are circinate in the bud; that is, +are rolled up in the manner shown in Fig. <a href="#Fig193">197</a>. Uncoiling as they grow, +they have some likeness to a crosier.</p> + +<div class="figcenter"> +<a name="Fig501" id="Fig501"></a> +<img src="images/fig501_508.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 501. The Walking-Fern, Camptosorus, reduced in size, +showing its fruit-dots on the veins approximated in pairs. 502. A small +piece (pinnule) of a Shield-Fern: a row of fruit-dots on each side of +the midrib, each covered by its kidney-shaped indusium. 503. A +spore-case from the latter, just bursting by the partial straightening +of the incomplete ring; well magnified. 504. Three of the spores of <a href="#Fig509">509</a>, +more magnified. 505. Schizæa pusilla, a very small and simple-leaved +Fern, drawn nearly of natural size. 506. One of the lobes of its +fruit-bearing portion, magnified, bearing two rows of spore-cases. 507. +Spore-case of the latter, detached, opening lengthwise. 508. +Adder-tongue, Ophioglossum; spore-cases in a kind of spike: <i>a</i>, a +portion of the fruiting part, about natural size; showing two rows of +the firm spore-cases, which open transversely into two valves.</p> +</div> + +<p>487. The fructification of Ferns is borne on the back or under side of +the leaves. The early botanists thought this such a peculiarity that +they always called a Fern-leaf a <span class="smcap">Frond</span>, and its petiole a <span class="smcap">Stipe</span>. Usage +continues these terms, although they are superfluous. The fruit of Ferns +consists of <span class="smcap">Spore-cases</span>, technically <span class="smcap">Sporangia</span>, which grow out of the +veins of the leaf. Sometimes these are distributed over the whole lower +<span class="pagenum"><a name="Page_159" id="Page_159">[Pg 159]</a></span>surface of the leaf or frond, or over the whole surface when there are +no proper leaf-blades to the frond, but all is reduced to stalks. +Commonly the spore-cases occupy only detached spots or lines, each of +which is called a <span class="smcap">Sorus</span>, or in English merely a Fruit-dot. In many Ferns +these fruit-dots are naked; in others they are produced under a +scale-like bit of membrane, called an <span class="smcap">Indusium</span>. In Maidenhair-Ferns a +little lobe of the leaf is folded back over each fruit-dot, to serve as +its shield or indusium. In the true Brake or Bracken (Pteris) the whole +edge of the fruit-bearing part of the leaf is folded back over it like a +hem.</p> + +<p>488. The form and structure of the spore-cases can be made out with a +common hand magnifying glass. The commonest kind (shown in Fig. <a href="#Fig501">503</a>) has +a stalk formed of a row of jointed cells, and is itself composed of a +layer of thin-walled cells, but is incompletely surrounded by a border +of thicker-walled cells, forming the <span class="smcap">Ring</span>. This extends from the stalk +up one side of the spore-case, round its summit, descends on the other +side, but there gradually vanishes. In ripening and drying the shrinking +of the cells of the ring on the outer side causes it to straighten; in +doing so it tears the spore-case open on the weaker side and discharges +the minute spores that fill it, commonly with a jerk which scatters them +to the wind. Another kind of spore-case (Fig. <a href="#Fig501">507</a>) is stalkless, and has +its ring-cells forming a kind of cap at the top: at maturity it splits +from top to bottom by a regular dehiscence. A third kind is of firm +texture and opens across into two valves, like a clam-shell (Fig. +<a href="#Fig501">508<sup><i>a</i></sup></a>): this kind makes an approach to the next family.</p> + +<div class="figcenter"> +<a name="Fig509" id="Fig509"></a> +<img src="images/fig509_510.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 509. A young prothallus of a Maiden-hair, moderately +enlarged, and an older one with the first fern-leaf developed from near +the notch. 510. Middle portion of the young one, much magnified, showing +below, partly among the rootlets, the <i>antheridia</i> or fertilizing +organs, and above, near the notch, three <i>pistillidia</i> to be +fertilized.</p> +</div> + +<p>489. The spores germinate on moistened ground. In a conservatory they +may be found germinating on a damp wall or on the edges of a +well-watered flower-pot. Instead of directly forming a fern-plantlet, +the spore grows first into a body which <span class="pagenum"><a name="Page_160" id="Page_160">[Pg 160]</a></span>closely resembles a small +Liverwort. This is named a <span class="smcap">Prothallus</span> (Fig. <a href="#Fig509">509</a>): from some point of +this a bud appears to originate, which produces the first fern-leaf, +soon followed by a second and third, and so the stem and leaves of the +plant are set up.</p> + +<div class="figcenter"> +<a name="Fig511" id="Fig511"></a> +<img src="images/fig511_519.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 511. Lycopodium Carolinianum, of nearly natural +size. 512. Inside view of one of the bracts and spore-case, magnified.</p> +<p><span class="smcap">Fig.</span> 513. Open 4-valved spore-case of a Selaginella, and +its four large spores (macrospores), magnified. 514. Macrospores of +another Selaginella. 515. Same separated.</p> +<p><span class="smcap">Fig.</span> 516. Plant of Isoetes. 517. Base of a leaf and +contained sporocarp filled with microspores cut across, magnified. 518. +Same divided lengthwise, equally magnified; some microspores seen at the +left. 519. Section of a spore-case containing macrospores, equally +magnified; at the right three macrospores more magnified.</p> +</div> + +<p>490. Investigation of this prothallus under the microscope resulted in +the discovery of a wholly unsuspected kind of fertilization, taking +place at this germinating stage of the plant. On the under side of the +prothallus two kinds of organs appear (Fig. <a href="#Fig509">510</a>). One may be likened to +an open and depressed ovule, with a single cell at bottom answering to +nucleus; the other, to an anther; but instead of pollen, it discharges +corkscrew-shaped microscopic filaments, which bear some cilia of extreme +tenuity, by the rapid vibration of which the filaments move freely over +a wet surface. These filaments travel over the surface of the +prothallus, and even to other prothalli (for there are natural hybrid +Ferns), reach and enter the ovule-like <span class="pagenum"><a name="Page_161" id="Page_161">[Pg 161]</a></span>cavities, and fertilize the +cell. This thereupon sets up a growth, forms a vegetable bud, and so +develops the new plant.</p> + +<p>491. An essentially similar process of fertilization has been discovered +in the preceding and the following families of Pteridophytes; but it is +mostly subterranean and very difficult to observe.</p> + +<p><a name="UNIT_492" id="UNIT_492">492.</a> <b>Club-Mosses or Lycopodiums.</b> Some of the common kinds, called Ground +Pine, are familiar, being largely used for Christmas wreaths and other +decoration. They are low evergreens, some creeping, all with +considerable wood in their stems: this thickly beset with small leaves. +In the axils of some of these leaves, or more commonly, in the axils of +peculiar leaves changed into bracts (as in Fig. <a href="#Fig511">511, 512</a>) spore-cases +appear, as roundish or kidney-shaped bodies, of firm texture, opening +round the top into two valves, and discharging a great quantity of a +very fine yellow powder, the spores.</p> + +<p>493. The Selaginellas have been separated from Lycopodium, which they +much resemble, because they produce two kinds of spores, in separate +spore-cases. One kind (<span class="smcap">Microspores</span>) is just that of Lycopodium; the +other consists of only four large spores (<span class="smcap">Macrospores</span>), in a spore-case +which usually breaks in pieces at maturity (Fig. <a href="#Fig511">513-515</a>).</p> + +<p><a name="UNIT_494" id="UNIT_494">494.</a> <b>The Quillworts, Isoetes</b> (Fig. <a href="#Fig511">516-519</a>), are very unlike Club Mosses +in aspect, but have been associated with them. They look more like +Rushes, and live in water, or partly out of it. A very short stem, like +a corm, bears a cluster of roots underneath; above it is covered by the +broad bases of a cluster of awl-shaped or thread-shaped leaves. The +spore-cases are immersed in the bases of the leaves. The outer +leaf-bases contain numerous macrospores; the inner are filled with +innumerable microspores.</p> + +<div class="figcenter"> +<a name="Fig520" id="Fig520"></a> +<img src="images/fig520.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 520. Plant of Marsilia quadrifoliata, reduced in +size; at the right a pair of sporocarps of about natural size.</p> +</div> + +<p><a name="UNIT_495" id="UNIT_495">495.</a> <b>The Pillworts</b> (<i>Marsilia</i> and <i>Pilularia</i>) are low aquatics, which +<span class="pagenum"><a name="Page_162" id="Page_162">[Pg 162]</a></span>bear globular or pill-shaped fruit (<span class="smcap">Sporocarps</span>) on the lower part of +their leaf-stalks or on their slender creeping stems. The leaves of the +commoner species of Marsilia might be taken for four-leaved Clover. (See +Fig. <a href="#Fig520">520</a>.) The sporocarps are usually raised on a short stalk. Within +they are divided lengthwise by a partition, and then crosswise by +several partitions. These partitions bear numerous delicate sacs or +spore-cases of two kinds, intermixed. The larger ones contain each a +large spore, or macrospore; the smaller contain numerous microspores, +immersed in mucilage. At maturity the fruit bursts or splits open at +top, and the two kinds of spores are discharged. The large ones in +germination produce a small prothallus; upon which the contents of the +microspores act in the same way as in Ferns, and with a similar result.</p> + +<p><a name="UNIT_496" id="UNIT_496">496.</a> <b>Azolla</b> is a little floating plant, looking like a small Liverwort +or Moss. Its branches are covered with minute and scale-shaped leaves. +On the under side of the branches are found egg-shaped thin-walled +sporocarps of two kinds. The small ones open across and discharge +microspores; the larger burst irregularly, and bring to view globose +spore-cases, attached to the bottom of the sporocarp by a slender stalk. +These delicate spore-cases burst and set free about four macrospores, +which are fertilized at germination, in the manner of the Pillworts and +Quillworts. (See Fig. <a href="#Fig521">521-526</a>.)</p> + +<div class="figcenter"> +<a name="Fig521" id="Fig521"></a> +<img src="images/fig521_526.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 521. Small plant of Azolla Caroliniana. 522. Portion +magnified, showing the two kinds of sporocarp; the small ones contain +microspores. 523 represents one more magnified. 524. The larger +sporocarp more magnified. 525. Same more magnified and burst open, +showing stalked spore-cases. 526. Two of the latter highly magnified; +one of them bursting shows four contained macrospores; between the two, +three of these spores highly magnified.</p> +</div> + +<p><a name="UNIT_497" id="UNIT_497">497.</a> <b>Cellular Cryptogams</b> (<a href="#UNIT_483">483</a>) are so called because composed, even in +their higher forms, of cellular tissue only, without proper wood-cells +or vessels. Many of the lower kinds are mere plates, or ribbons, or +simple rows of cells, or even single cells. But their highest orders +follow the plan of Ferns and phanerogamous plants in having stem and +leaves for their upward growth, and commonly roots, or at least +rootlets, <span class="pagenum"><a name="Page_163" id="Page_163">[Pg 163]</a></span>to attach them to the soil, or to trunks, or to other bodies +on which they grow. Plants of this grade are chiefly Mosses. So as a +whole they take the name of</p> + +<p><a name="UNIT_498" id="UNIT_498">498.</a> <b>Bryophyta, Bryophytes</b> in English form, Bryum being the Greek name +of a Moss. These plants are of two principal kinds: true Mosses +(<i>Musci</i>, which is their Latin name in the plural); and Hepatic Mosses, +or Liverworts (<i>Hepaticæ</i>).</p> + +<div class="figcenter"> +<a name="Fig527" id="Fig527"></a> +<img src="images/fig527_528.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 527. Single plant of Physcomitrium pyriforme, +magnified. 528. Top of a leaf, cut across; it consists of a single layer +of cells.</p> +</div> + +<p><a name="UNIT_499" id="UNIT_499">499.</a> <b>Mosses or Musci.</b> The pale Peat-mosses (species of Sphagnum, the +principal component of sphagnous bogs) and the strong-growing Hair-cap +Moss (Polytrichum) are among the larger and commoner representatives of +this numerous family; while Fountain Moss (Fontinalis) in running water +sometimes attains the length of a yard or more. On the other hand, some +are barely individually distinguishable to the naked eye. Fig. <a href="#Fig527">527</a> +represents a common little Moss, enlarged to about twelve times its +natural size; and by its side is part of a leaf, much magnified, showing +that it is composed of cellular tissue (parenchyma-cells) only. The +leaves of Mosses are always simple, distinct, and sessile on the stem. +The fructification is an urn-shaped spore-case, in this as in most cases +raised on a slender stalk. The spore-case loosely bears on its summit a +thin and pointed cap, like a candle-extinguisher, called a <i>Calyptra</i>. +Detaching this, it is found that the spore-case is like a pyxis (<a href="#UNIT_376">376</a>), +that is, the top at maturity comes off as a lid (<i>Operculum</i>); and that +the interior is filled with a green powder, the spores, which are +discharged through the open mouth. In most Mosses there is a fringe of +one or two rows of teeth or membrane around this mouth or orifice, the +<i>Peristome</i>. When moist the peristome closes hygrometrically over the +orifice more or less; when drier the teeth or processes commonly bend +outward or recurve; and then the spores more readily escape. In Hair-cap +Moss a membrane is stretched quite across the mouth, like a drum-head, +retaining the spores until this wears away. See Figures <a href="#Fig527">527</a>-<a href="#Fig529">541</a> for +details.</p> + +<p>500. Fertilization in Mosses is by the analogues of stamens and pistils, +which are hidden in the axils of leaves, or in the cluster of leaves at the +<span class="pagenum"><a name="Page_164" id="Page_164">[Pg 164]</a></span>end of the stem. The analogue of the anther (<i>Antheridium</i>) is a +cellular sac, which in bursting discharges innumerable delicate cells +floating in a mucilaginous liquid; each of these bursts and sets free a +vibratile self-moving thread. These threads, one or more, reach the +orifice of the pistil-shaped body, the <i>Pistillidium</i>, and act upon a +particular cell at its base within. This cell in its growth develops +into the spore-case and its stalk (when there is any), carrying on its +summit the wall of the pistillidium, which becomes the calyptra.</p> + +<div class="figcenter"> +<a name="Fig529" id="Fig529"></a> +<img src="images/fig529_541.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 529. Mnium cuspidatum, smaller than nature. 530. Its +calyptra, detached, enlarged. 531. Its spore-case, with top of stalk, +magnified, the lid (532) being detached, the outer peristome appears. +533. Part of a cellular ring (<i>annulus</i>) which was under the lid, +outside of the peristome, more magnified. 534. Some of the outer and of +the inner peristome (consisting of jointed teeth) much magnified. 535. +Antheridia and a pistillidium (the so-called flower) at end of a stem of +same plant, the leaves torn away (♂, antheridia, +♀, pistillidium), magnified. 536. A bursting antheridium, and +some of the accompanying jointed threads, highly magnified. 537. Summit +of an open spore-case of a Moss, which has a peristome of 16 pairs of +teeth. 538. The double peristome of a Hypnum. 539-541. Spore-case, +detached calyptra, and top of more enlarged spore-case and detached lid, +of Physcomitrium pyriforme (Fig. <a href="#Fig527">527</a>): orifice shows that there is no +peristome.</p> +</div> + +<p><a name="UNIT_501" id="UNIT_501">501.</a> <b>Liverworts or Hepatic Mosses</b> (<i>Hepaticæ</i>) in some kinds resemble +true Mosses, having distinct stem and leaves, although their leaves +occasionally run together; while in others there is no distinction of +stem and leaf, but the whole plant is a leaf-like body, which produces +rootlets on the lower face and its fructification on the upper. Those of +the moss-like kind (sometimes called Scale-Mosses) have their tender +spore-cases splitting into four valves; and with their spores are +intermixed some slender spiral <span class="pagenum"><a name="Page_165" id="Page_165">[Pg 165]</a></span>and very hygrometric threads (called +<i>Elaters</i>) which are thought to aid in the dispersion of the spores. +(Fig. <a href="#Fig542">542-544</a>.)</p> + +<div class="figcenter"> +<a name="Fig542" id="Fig542"></a> +<img src="images/fig542_544.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 542. Fructification of a Jungermannia, magnified; +its cellular spore-stalk, surrounded at base by some of the leaves, at +summit the 4-valved spore-case opening, discharging spores and elaters. +543. Two elaters and some spores from the same, highly magnified.</p> +<p><span class="smcap">Fig.</span> 544. One of the frondose Liverworts, Steetzia, +otherwise like a Jungermannia; the spore-case not yet protruded from its +sheath.</p> +</div> + +<p>502. Marchantia, the commonest and largest of the true Liverworts, forms +large green plates or fronds on damp and shady ground, and sends up from +some part of the upper face a stout stalk, ending in a several-lobed +umbrella-shaped body, under the lobes of which hang several thin-walled +spore-cases, which burst open and discharge spores and elaters. Riccia +natans (Fig. <a href="#Fig545">545</a>) consists of wedge-shaped or heart-shaped fronds, which +float free in pools of still water. The under face bears copious +rootlets; in the substance of the upper face are the spore-cases, their +pointed tips merely projecting: there they burst open, and discharge +their spores. These are comparatively few and large, and are in fours; +so they are very like the macrospores of Pillworts or Quillworts.</p> + +<p><a name="UNIT_503" id="UNIT_503">503.</a> <b>Thallophyta, or Thallophytes</b> in English form. This is the name for +the lower class of Cellular Cryptogams,—plants in which there is no +marked distinction into root, stem, and leaves. Roots in any proper +sense they never have, as organs for absorbing, although some of the +larger Seaweeds (such as the Sea Colander, Fig. <a href="#Fig553">553</a>) have them as +holdfasts. Instead of axis and foliage, there is a stratum of frond, in +such plants commonly called a <span class="smcap">Thallus</span> (by a strained use of a Greek and +Latin word which means a green shoot or bough), which may have any kind +of form, leaf-like, stem-like, branchy, extended to a flat plate, or +gathered into a sphere, or drawn out into threads, or reduced to a +single row of cells, or even reduced to single cells. Indeed, +Thallophytes are so multifarious, so numerous in kinds, so protean in +their stages and transformations, so recondite in their fructification, +and many so microscopic in size, either of <span class="pagenum"><a name="Page_166" id="Page_166">[Pg 166]</a></span>the plant itself or its +essential organs, that they have to be elaborately described in separate +books and made subjects of special study.</p> + +<div class="figcenter"> +<a name="Fig545" id="Fig545"></a> +<img src="images/fig545_549.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 545, 546. Two plants of Riccia natans, about natural +size. 547. Magnified section of a part of the frond, showing two +immersed spore-cases, and one emptied space. 548. Magnified section of a +spore-case with some spores. 549. Magnified spore-case torn out, and +spores; one figure of the spores united; the other of the four +separated.</p> +</div> + +<p>504. Nevertheless, it may be well to try to give some general idea of +what Algæ and Lichens and Fungi are. Linnæus had them all under the +orders of Algæ and Fungi. Afterwards the Lichens were separated; but of +late it has been made most probable that a Lichen consists of an Alga +and a Fungus conjoined. At least it must be so in some of the ambiguous +forms. Botanists are in the way of bringing out new classifications of +the Thallophytes, as they come to understand their structure and +relations better. Here, it need only be said that</p> + +<p>505. Lichens live in the air, that is, on the ground, or on rocks, +trunks, walls, and the like, and grow when moistened by rains. They +assimilate air, water, and some earthy matter, just as do ordinary +plants. Algæ, or Seaweeds, live in water, and live the same kind of life +as do ordinary plants. Fungi, whatever medium they inhabit, live as +animals do, upon organic matter,—upon what other plants have +assimilated, or upon the products of <span class="pagenum"><a name="Page_167" id="Page_167">[Pg 167]</a></span>their decay. True as these general +distinctions are, it is no less true that these orders run together in +their lowest forms; and that Algæ and Fungi may be traced down into +forms so low and simple that no clear line can be drawn between them; +and even into forms of which it is uncertain whether they should be +called plants or animals. It is as well to say that they are not high +enough in rank to be distinctively either the one or the other. On the +other hand there is a peculiar group of plants, which in simplicity of +composition resemble the simpler Algæ, while in fructification and in +the arrangements of their simple cells into stem and branches they seem +to be of a higher order, viz.:—</p> + +<div class="figcenter"> +<a name="Fig550" id="Fig550"></a> +<img src="images/fig550_552.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 550. Branch of a Chara, about natural size. 551. A +fruiting portion, magnified, showing the structure; a sporocarp, and an +antheridium. 552. Outlines of a portion of the stem in section, showing +the central cell and the outer or cortical cells.</p> +</div> + +<p><a name="UNIT_506" id="UNIT_506">506.</a> <b>Characeæ.</b> These are aquatic herbs, of considerable size, abounding +in ponds. The simpler kinds (Nitella) have the stem formed of a single +row of tubular cells, and at the nodes, or junction of the cells, a +whorl of similar branches. Chara (Fig. <a href="#Fig550">550-552</a>) is the same, except that +the cells which make up the stem and the principal branches are +strengthened by a coating of many smaller tubular cells, applied to the +surface of the main or central cell. The fructification consists of a +globular sporocarp of considerable size, which is spirally enwrapped by +tubular cells twisted around it: by the side of this is a smaller and +globular antheridium. The latter breaks up into eight shield-shaped +<span class="pagenum"><a name="Page_168" id="Page_168">[Pg 168]</a></span>pieces, with an internal stalk, and bearing long and ribbon shaped +filaments, which consist of a row of delicate cells, each of which +discharges a free-moving microscopic thread (the analogue of the pollen +or pollen-tube), nearly in the manner of Ferns and Mosses. One of these +threads reaches and fertilizes a cell at the apex of the nucleus or +solid body of the sporocarp. This subsequently germinates and forms a +new individual.</p> + +<p><a name="UNIT_507" id="UNIT_507">507.</a> <b>Algæ or Seaweeds.</b> The proper Seaweeds may be studied by the aid of +Professor Farlow's "Marine Algæ of New England;" the fresh-water +species, by Prof. H. C. Woods's "Fresh-water Algæ of North America," a +larger and less accessible volume. A few common forms are here very +briefly mentioned and illustrated, to give an idea of the family. But +they are of almost endless diversity.</p> + +<div class="figcenter"> +<a name="Fig553" id="Fig553"></a> +<img src="images/fig553_554.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 553. Agarum Turneri, Sea Colander (so called from +the perforations with which the frond, as it grows, becomes riddled); +very much reduced in size.</p> +<p><span class="smcap">Fig.</span> 554. Upper end of a Rockweed, Fucus vesiculosus, +reduced half or more, <i>b</i>, the fructification.</p> +</div> + +<p>508. The common Rockweed (Fucus vesiculosus, Fig. <a href="#Fig553">554</a>, abounding between +high and low water mark on the coast), the rarer Sea Colander (Agarum +Turneri, Fig. <a href="#Fig553">553</a>), and Laminaria, of which the larger forms are called +Devil's Aprons, are good representatives of the olive green or brownish +Seaweeds. They are attached either by a disk-like base or by root-like +holdfasts to the rocks or stones on which they grow.</p> + +<div class="figcenter"> +<a name="Fig555" id="Fig555"></a> +<img src="images/fig555_556.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 555. Magnified section through a fertile conceptacle +of Rockweed, showing the large spores in the midst of threads of cells. +556. Similar section of a sterile conceptacle, containing slender +antheridia. From Farlow's "Marine Algæ of New England."</p> +</div> + +<p>509. The hollow and inflated places in the Fucus vesiculosus or Rockweed +(Fig. <a href="#Fig553">554</a>) are air-bladders for buoyancy. The fructification forms in +the substance of the tips of the frond: the rough dots mark the places +where the conceptacles open. The spores and the fertilizing cells are in +different plants. Sections of the two kinds of conceptacles are given in +Fig. <a href="#Fig555">555 and 556</a>. The contents of the conceptacles are discharged +through <span class="pagenum"><a name="Page_169" id="Page_169">[Pg 169]</a></span>a small orifice which in each figure is at the margin of the +page. The large spores are formed eight together in a mother-cell. The +minute motile filaments of the antheridia fertilize the large spores +after injection into the water: and then the latter promptly acquire a +cell-wall and germinate.</p> + +<p>510. The Florideæ or Rose-red series of marine Algæ (which, however, are +sometimes green or brownish) are the most attractive to amateurs. The +delicate Porphyra or Laver is in some countries eaten as a delicacy, and +the cartilaginous Chondrus crispus has been largely used for jelly. +Besides their conceptacles, which contain true spores (Fig. <a href="#Fig557">560</a>), they +mostly have a fructification in <i>Tetraspores</i>, that is, of spores +originating in fours (Fig. <a href="#Fig557">559</a>).</p> + +<div class="figcenter"> +<a name="Fig557" id="Fig557"></a> +<img src="images/fig557_560.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 557. Small plant of Chondrus crispus, or Carrageen +Moss, reduced in size, in fruit; the spots represent the fructification, +consisting of numerous tetraspores in bunches in the substance of the +plant. 558. Section through the thickness of one of the lobes, +magnified, passing through two of the imbedded fruit-clusters. 559. Two +of its tetraspores (spores in fours), highly magnified.</p> +<p><span class="smcap">Fig.</span> 560. Section through a conceptacle of Delesseria +Leprieurei, much magnified, showing the spores, which are single +specialized cells, two or three in a row.</p> +</div> + +<div class="figcenter"> +<a name="Fig561" id="Fig561"></a> +<img src="images/fig561_563.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 561. A piece of the rose-red Delesseria Leprieurei, +double natural size. 562. A piece cut out and much magnified, showing +that it is composed of a layer of cells. 563. A few of the cells more +highly magnified: the cells are gelatinous and thick-walled.</p> +</div> + +<p>511. The Grass-green Algæ sometimes form broad membranous fronds, such +as those of the common Ulva of the sea-shore, but most of them form mere +threads, either simple or branched. To this division belong almost <span class="pagenum"><a name="Page_170" id="Page_170">[Pg 170]</a></span>all +the Fresh-water Algæ, such as those which constitute the silky threads +or green slime of running streams or standing pools, and which were all +called Confervas before their immense diversity was known. Some are +formed of a single row of cells, developed each from the end of another. +Others branch, the top of one cell producing more than one new one (Fig. +<a href="#Fig564">564</a>). Others, of a kind which is very common in fresh water, simple +threads made of a line of cells, have the chlorophyll and protoplasm of +each cell arranged in spiral lines or bands. They form spores in a +peculiar way, which gives to this family the designation of conjugating +Algæ.</p> + +<div class="figcenter"> +<a name="Fig564" id="Fig564"></a> +<img src="images/fig564_565.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 564. The growing end of a branching Conferva +(Cladophora glomerata), much magnified; showing how, by a kind of +budding growth, a new cell is formed by a cross partition separating the +newer tip from the older part below; also, how the branches arise.</p> +<p><span class="smcap">Fig.</span> 565. Two magnified individuals of a Spirogyra, +forming spores by conjugation; a completed spore at base: above, +successive stages of the conjugation are represented.</p> +</div> + +<p>512. At a certain time two parallel threads approach each other more +closely; contiguous parts of a cell of each thread bulge or grow out, +and unite when they meet; the cell-wall partitions between them are +absorbed so as to open a free communication; the spiral band of green +matter in both cells breaks up; the whole of that of one cell passes +over into the other; and of the united contents a large green spore is +formed. Soon the old cells decay, and the spore <span class="pagenum"><a name="Page_171" id="Page_171">[Pg 171]</a></span>set free is ready to +germinate. Fig. <a href="#Fig564">565</a> represents several stages of the conjugating +process, which, however, would never be found all together like this in +one pair of threads.</p> + +<div class="figcenter"> +<a name="Fig566" id="Fig566"></a> +<img src="images/fig566_567.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 566. Closterium acutum, a common Desmid, moderately +magnified. It is a single firm-walled cell, filled with green +protoplasmic matter.</p> +<p><span class="smcap">Fig.</span> 567. More magnified view of three stages of the +conjugation of a pair of the same.</p> +</div> + +<p>513. Desmids and Diatomes, which are microscopic one-celled plants of +the same class, conjugate in the same way, as is shown in a Closterium +by Fig. <a href="#Fig566">566, 567</a>. Here the whole living contents of two individuals are +incorporated into one spore, for a fresh start. A reproduction which +costs the life of two individuals to make a single new one would be +fatal to the species if there were not a provision for multiplication by +the prompt division of the new-formed individual into two, and these +again into two, and so on in geometrical ratio. And the costly process +would be meaningless if there were not some real advantage in such a +fresh start, that is, in sexes.</p> + +<div class="figcenter"> +<a name="Fig568" id="Fig568"></a> +<img src="images/fig568_574.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 568. Early stage of a species of Botrydium, a +globose cell. 569, 570. Stages of growth. 571. Full-grown plant, +extended and ramified below in a root-like way. 572. A Vaucheria; single +cell grown on into a much-branched thread; the end of some branches +enlarging, and the green contents in one (<i>a</i>) there condensed into a +spore. 573. More magnified view of <i>a</i>, and the mature spore escaping. +574. Bryopsis plumosa; apex of a stem with its branchlets; all the +extension of one cell. Variously magnified.</p> +</div> + +<p>514. There are other Algæ of the grass-green series which consist of +single cells, but which by continued growth form plants of considerable +size. Three kinds of these are represented in Fig. <a href="#Fig568">568-574</a>.</p> + +<p><a name="UNIT_515" id="UNIT_515">515.</a> <b>Lichens</b>, Latin <i>Lichenes</i>, are to be studied in the works of the +late Professor Tuckerman, but a popular exposition is greatly needed. +The subjoined illustrations (Fig. <a href="#Fig575">575-580</a>) may simply indicate what some +of the commoner forms are like. The cup, or shield-shaped spot, or knob, +which bears the fructification is named the <i>Apothecium</i>. This is mainly +<span class="pagenum"><a name="Page_172" id="Page_172">[Pg 172]</a></span>composed of slender sacs (<i>Asci</i>), having thread-shaped cells +intermixed; and each ascus contains few or several spores, which are +commonly double or treble. Most Lichens are flat expansions of grayish +hue; some of them foliaceous in texture, but never of bright green +color; more are crustaceous; some are wholly pulverulent and nearly +formless. But in several the vegetation lengthens into an axis (as in +Fig. <a href="#Fig575">580</a>), or imitates stem and branches or threads, as in the +Reindeer-Moss on the ground in our northern woods, and the Usnea hanging +from the boughs of old trees overhead.</p> + +<div class="figcenter"> +<a name="Fig575" id="Fig575"></a> +<img src="images/fig575_580.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 575. A stone on which various Lichens are growing, +such as (passing from left to right) a Parmelia, a Sticta, and on the +right, Lecidia geographica, so called from its patches resembling the +outline of islands or continents as depicted upon maps. 576. Piece of +thallus of Parmelia conspersa, with section through an apothecium. 577. +Section of a smaller apothecium, enlarged. 578. Two asci of same, and +contained spores, and accompanying filaments; more magnified. 579. Piece +of thallus of a Sticta, with section, showing the immersed apothecia; +the small openings of these dot the surface. 580. Cladonia coccinea; the +fructification is in the scarlet knobs, which surround the cups.</p> +</div> + +<p><a name="UNIT_516" id="UNIT_516">516.</a> <b>Fungi.</b> For this immense and greatly diversified class, it must here +suffice to indicate the parts of a Mushroom, a Sphæria, and of one or +two common Moulds. The true vegetation of common Fungi consists of +slender cells which form what is called a <i>Mycelium</i>. These filamentous +<span class="pagenum"><a name="Page_173" id="Page_173">[Pg 173]</a></span>cells lengthen and branch, growing by the absorption through their +whole surface of the decaying, or organizable, or living matter which +they feed upon. In a Mushroom (Agaricus), a knobby mass is at length +formed, which develops into a stout stalk (<i>Stipe</i>), bearing the cap +(<i>Pileus</i>): the under side of the cap is covered by the <i>Hymenium</i>, in +this genus consisting of radiating plates, the gills or <i>Lamellæ</i>; and +these bear the powdery spores in immense numbers. Under the microscope, +the gills are found to be studded with projecting cells, each of which, +at the top, produces four stalked spores. These form the powder which +collects on a sheet of paper upon which a mature Mushroom is allowed to +rest for a day or two. (Fig. <a href="#Fig581">581-586</a>.)</p> + +<p>517. The esculent Morel, also Sphæria (Fig. <a href="#Fig581">585, 586</a>), and many other +Fungi bear their spores in sacs (asci) exactly in the manner of Lichens +(<a href="#UNIT_515">515</a>).</p> + +<div class="figcenter"> +<a name="Fig581" id="Fig581"></a> +<img src="images/fig581_586.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 581. Agaricus campestris, the common edible +Mushroom. 582. Section of cap and stalk. 583. Minute portion of a +section of a gill, showing some spore-bearing cells, much magnified. +584. One of these, with its four spores, more magnified.</p> +<p><span class="smcap">Fig.</span> 585. Sphæria rosella. 586. Two of the asci and +contained double spores, quite like those of a Lichen; much magnified.</p> +</div> + +<p>518. Of the Moulds, one of the commoner is the Bread-Mould (Fig. <a href="#Fig587">587</a>). +In fruiting it sends up a slender stalk, which bears a globular sac; +<span class="pagenum"><a name="Page_174" id="Page_174">[Pg 174]</a></span>this bursts at maturity and discharges innumerable spores. The blue +Cheese-Mould (Fig. <a href="#Fig587">588</a>) bears a cluster of branches at top, each of +which is a row of naked spores, like a string of beads, all breaking +apart at maturity. Botrytis (Fig. <a href="#Fig587">589</a>), the fruiting stalk of which +branches, and each branch is tipped with a spore, is one of the many +moulds which live and feed upon the juices of other plants or of +animals, and are often very destructive. The extremely numerous kinds of +smut, rust, mildew, the ferments, bacteria, and the like, many of them +very destructive to other vegetable and to animal life, are also low +forms of the class of Fungi.<a name="FNanchor_1_3" id="FNanchor_1_3"></a><a href="#Footnote_1_3" class="fnanchor">[1]</a></p> + +<div class="figcenter"> +<a name="Fig587" id="Fig587"></a> +<img src="images/fig587_589.jpg" alt="" /><br /> +<p><span class="smcap">Fig.</span> 587. Ascophora, the Bread-Mould. 588. Aspergillus +glaucus, the mould of cheese, but common on mouldy vegetables. 589. A +species of Botrytis. All magnified.</p> +</div> + + +<div class="footnotes"><h3>FOOTNOTES:</h3> + +<div class="footnote"><p><a name="Footnote_1_3" id="Footnote_1_3"></a><a href="#FNanchor_1_3"><span class="label">[1]</span></a> The "Introduction to Cryptogamous Botany," or third volume +of "The Botanical Text Book," now in preparation by the author's +colleague, Professor Farlow, will be the proper guide in the study of +the Flowerless Plants, especially of the Algæ and Fungi.</p></div> +</div> + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_175" id="Page_175">[Pg 175]</a></span></p> +<h2><a name="Section_XVIII" id="Section_XVIII"></a>SECTION XVIII. CLASSIFICATION AND NOMENCLATURE.</h2> + + +<p>519. Classification, in botany, is the consideration of plants in +respect to their kinds and relationships. Some system of Nomenclature, +or naming, is necessary for fixing and expressing botanical knowledge so +as to make it available. The vast multiplicity of plants and the various +degrees of their relationship imperatively require order and system, not +only as to <i>names</i> for designating the kinds of plants, but also as to +<i>terms</i> for defining their differences. Nomenclature is concerned with +the names of plants. Terminology supplies names of organs or parts, and +terms to designate their differences.</p> + + +<h3><a name="Subsect_XVIII_1" id="Subsect_XVIII_1">§ 1.</a> KINDS AND RELATIONSHIP.</h3> + +<p>520. Plants and animals have two great peculiarities: 1st, they form +themselves; and 2d, they multiply themselves. They reproduce their kind +in a continued succession of</p> + +<p><a name="UNIT_521" id="UNIT_521">521.</a> <b>Individuals.</b> Mineral things occur as <i>masses</i>, which are divisible +into smaller and still smaller ones without alteration of properties. +But organic things (vegetables and animals) exist as <i>individual +beings</i>. Each owes its existence to a parent, and produces similar +individuals in its turn. So each individual is a link of a chain; and to +this chain the natural-historian applies the name of</p> + +<p><a name="UNIT_522" id="UNIT_522">522.</a> <b>Species.</b> All the descendants from the same stock therefore compose +one species. And it was from our observing that the several sorts of +plants or animals steadily reproduce themselves, or, in other words, +keep up a succession of similar individuals, that the idea of species +originated. There are few species, however, in which man has actually +observed the succession for many generations. It could seldom be proved +that all the White Pine trees or White Oaks of any forest came from the +same stock. But observation having familiarized us with the general fact +that individuals proceeding from the same stock are essentially alike, +we infer from their close resemblance that these similar individuals +belong to the same species. That is, we infer it when the individuals +are as much like each other as those are which we know, or confidently +suppose, to have sprung from the same stock.</p> + +<p>523. Identity in species is inferred from close similarity in all +essential respects, or whenever the differences, however considerable, +are not known or reasonably supposed to have been originated in the +course of time under changed conditions. No two individuals are exactly +alike; a tendency to variation pervades all living things. In +cultivation, where variations are looked after and cared for, very +striking differences come to light; and if in wild nature they are less +common or less conspicuous, it is partly because they are uncared for. +When such variant forms are pretty well marked they are called</p> + +<p><span class="pagenum"><a name="Page_176" id="Page_176">[Pg 176]</a></span></p><p><a name="UNIT_524" id="UNIT_524">524.</a> <b>Varieties.</b> The White Oak, for example, presents two or three +varieties in the shape of the leaves, although they may be all alike +upon each particular tree. The question often arises, and it is often +hard to answer, whether the difference in a particular case is that of a +variety, or is specific. If the former, it may commonly be proved by +finding such intermediate degrees of difference in various individuals +as to show that no clear distinction can be drawn between them; or else +by observing the variety to vary back again in some of its offspring. +The sorts of Apples, Pears, Potatoes, and the like, show that +differences which are permanent in the individual, and continue +unchanged through a long series of generations when propagated by +division (as by offsets, cuttings, grafts, bulbs, tubers, etc.), are not +likely to be reproduced by seed. Still they sometimes are so, and +perhaps always tend in that direction. For the fundamental law in +organic nature is that offspring shall be like parent.</p> + +<p><span class="smcap">Races</span> are such strongly marked varieties, capable of coming true to +seed. The different sorts of Wheat, Maize, Peas, Radishes, etc., are +familiar examples. By selecting those individuals of a species which +have developed or inherited any desirable peculiarity, keeping them from +mingling with their less promising brethren, and selecting again the +most promising plants raised from their seeds, the cultivator may in a +few generations render almost any variety transmissible by seed, so long +as it is cared for and kept apart. In fact, this is the way the +cultivated domesticated races, so useful to man, have been fixed and +preserved. Races, in fact, can hardly, if at all, be said to exist +independently of man. But man does not really produce them. Such +peculiarities—often surprising enough—now and then originate, we know +not how (the plant <i>sports</i>, as the gardeners say); they are only +preserved, propagated, and generally further developed, by the +cultivator's skilful care. If left alone, they are likely to dwindle and +perish, or else revert to the original form of the species. Vegetable +races are commonly annuals, which can be kept up only by seed, or herbs +of which a succession of generations can be had every year or two, and +so the education by selection be completed without great lapse of time. +But all fruit-trees could probably be fixed into races in an equal +number of generations.</p> + +<p><span class="smcap">Bud-varieties</span> are those which spring from buds instead of seed. They are +uncommon to any marked extent. They are sometimes called <i>Sports</i>, but +this name is equally applied to variations among seedlings.</p> + +<p><span class="smcap">Cross-breeds</span>, strictly so-called, are the variations which come from +cross-fertilizing one variety of a species with another.</p> + +<p><span class="smcap">Hybrids</span> are the varieties, if they may be so called,—which come from +the crossing of species (<a href="#UNIT_331">331</a>). Only nearly related species can be +hybridized; and the resulting progeny is usually self-sterile, but not +always. Hybrid plants, however, may often be fertilized and made +prolific by the pollen of one or the other parent. This produces another +kind of cross-breeds.</p> + +<p>525. Species are the units in classification. Varieties, although of +<span class="pagenum"><a name="Page_177" id="Page_177">[Pg 177]</a></span>utmost importance in cultivation and of considerable consequence in the +flora of any country, are of less botanical significance. For they are +apt to be indefinite and to shade off one form into another. But +species, the botanist <i>expects</i> to be distinct. Indeed, the practical +difference to the botanist between species and varieties is the definite +limitation of the one and the indefiniteness of the other. The +botanist's determination is partly a matter of observation, partly of +judgment.</p> + +<p>526. In an enlarged view, varieties may be incipient species; and nearly +related species probably came from a common stock in earlier times. For +there is every reason to believe that existing vegetation came from the +more or less changed vegetation of a preceding geological era. However +that may be, species are regarded as permanent and essentially unchanged +in their succession of individuals through the actual ages.</p> + +<p>527. There are, at nearly the lowest computation, as many as one hundred +thousand species of phanerogamous plants, and the cryptogamous species +are thought to be still more numerous. They are all connected by +resemblances or relationships, near and remote, which show that they are +all parts of one system, realizations in nature, as we may affirm, of +the conception of One Mind. As we survey them, they do not form a single +and connected chain, stretching from the lowest to the highest organized +species, although there obviously are lower and higher grades. But the +species throughout group themselves, as it were, into clusters or +constellations, and these into still more comprehensive clusters, and so +on, with gaps between. It is this clustering which is the ground of the +recognition of <i>kinds</i> of species, that is, of groups of species of +successive grades or degree of generality; such as that of similar +species into <i>Genera</i>, of genera into <i>Families</i> or <i>Orders</i>, of orders +into <i>Classes</i>. In classification the sequence, proceeding from higher +or more general to lower or special, is always <span class="smcap">Class</span>, <span class="smcap">Order</span>, <span class="smcap">Genus</span>, +<span class="smcap">Species</span>, <span class="smcap">Variety</span> (if need be).</p> + +<p><a name="UNIT_528" id="UNIT_528">528.</a> <b>Genera</b> (in the singular, <i>Genus</i>) are assemblages of closely +related species, in which the essential parts are all constructed on the +same particular type or plan. White Oak, Red Oak, Scarlet Oak, Live Oak, +etc., are so many species of the Oak genus (Latin, <i>Quercus</i>). The +Chestnuts compose another genus; the Beeches another. The Apple, Pear, +and Crab are species of one genus, the Quince represents another, the +various species of Hawthorn a third. In the animal kingdom the common +cat, the wild-cat, the panther, the tiger, the leopard, and the lion are +species of the cat kind or genus; while the dog, the jackal, the +different species of wolf, and the foxes, compose another genus. Some +genera are represented by a vast number of species, others by few, very +many by only one known species. For the genus may be as perfectly +represented in one species as in several, although, if this were the +case throughout, genera and species would of course be identical. The +Beech genus and the Chestnut genus would be just as distinct from the +Oak genus even if but one Beech and Chestnut were known; as indeed was +once the case.</p> + +<p><span class="pagenum"><a name="Page_178" id="Page_178">[Pg 178]</a></span></p><p><a name="UNIT_529" id="UNIT_529">529.</a> <b>Orders</b> are groups of genera that resemble each other; that is, +they are to genera what genera are to species. As familiar +illustrations, the Oak, Chestnut, and Beech genera, along with the Hazel +genus and the Hornbeams, all belong to one order. The Birches and the +Alders make another; the Poplars and Willows, another; the Walnuts (with +the Butternut) and the Hickories, still another. The Apple genus, the +Quince and the Hawthorns, along with the Plums and Cherries and the +Peach, the Raspberry with the Blackberry, the Strawberry, the Rose, +belong to a large order, which takes its name from the Rose. Most +botanists use the names "Order" and "Family" synonymously; the latter +more popularly, as "the Rose Family," the former more technically, as +"Order <i>Rosaceæ</i>."</p> + +<p>530. But when the two are distinguished, as is common in zoölogy, Family +is of lower grade than Order.</p> + +<p><a name="UNIT_531" id="UNIT_531">531.</a> <b>Classes</b> are still more comprehensive assemblages, or great groups. +Thus, in modern botany, the Dicotyledonous plants compose one class, the +Monocotyledonous plants another (<a href="#UNIT_36">36-40</a>).</p> + +<p>532. These four grades, Class, Order, Genus, Species, are of universal +use. Variety comes in upon occasion. For, although a species may have no +recognized varieties, a genus implies at least one species belonging to +it; every genus is of some order, and every order of some class.</p> + +<p>533. But these grades by no means exhaust the resources of +classification, nor suffice for the elucidation of all the distinctions +which botanists recognize. In the first place, a higher grade than that +of class is needful for the most comprehensive of divisions, that of all +plants into the two <i>Series</i> of Phanerogamous and Cryptogamous (<a href="#UNIT_6">6</a>); and +in natural history there are the two <i>Kingdoms</i> or <i>Realms</i>, the +Vegetable and the Animal.</p> + +<p>534. Moreover, the stages of the scaffolding have been variously +extended, as required, by the recognition of assemblages lower than +class but higher than order, viz. <i>Subclass</i> and <i>Cohort</i>; or lower than +order, a <i>Suborder</i>; or between this and genus, a <i>Tribe</i>; or between +this and tribe, a <i>Subtribe</i>; or between genus and species, a +<i>Subgenus</i>; and by some a species has been divided into <i>Subspecies</i>, +and a variety into <i>Subvarieties</i>. Last of all are <i>Individuals</i>. +Suffice it to remember that the following are the principal grades in +classification, with the proper sequence; also that only those here +printed in small capitals are fundamental and universal in botany:—</p> + +<ul class="lsoff"><li> +<span class="smcap">Series</span>, +<ul class="lsoff"><li> +<span class="smcap">Class</span>, Subclass, Cohort, +<ul class="lsoff"><li> +<span class="smcap">Order</span>, or <span class="smcap">Family</span>, Suborder, Tribe, Subtribe, +<ul class="lsoff"><li> +<span class="smcap">Genus</span>, Subgenus or Section, +<ul class="lsoff"><li> +<span class="smcap">Species</span>, Variety. +</li></ul></li></ul></li></ul></li></ul></li></ul> + + +<p><span class="pagenum"><a name="Page_179" id="Page_179">[Pg 179]</a></span></p><h3><a name="Subsect_XVIII_2" id="Subsect_XVIII_2">§ 2.</a> NAMES, TERMS, AND CHARACTERS.</h3> + +<p>535. The name of a plant is the name of its genus followed by that of +the species. The name of the genus answers to the surname (or family +name); that of the species to the baptismal name of a person. Thus +<i>Quercus</i> is the name of the Oak genus; <i>Quercus alba</i>, that of the +White Oak, <i>Q. rubra</i>, that of Red Oak, <i>Q. nigra</i>, that of the +Black-Jack, etc. Botanical names being Latin or Latinized, the adjective +name of the species comes after that of the genus.</p> + +<p><a name="UNIT_536" id="UNIT_536">536.</a> <b>Names of Genera</b> are of one word, a substantive. The older ones are +mostly classical Latin, or Greek adopted into Latin; such as <i>Quercus</i> +for the Oak genus, <i>Fagus</i> for the Beech, <i>Corylus</i>, the Hazel, and the +like. But as more genera became known, botanists had new names to make +or borrow. Many are named from some appearance or property of the +flowers, leaves, or other parts of the plant. To take a few examples +from the early pages of the "Manual of the Botany of the Northern United +States,"—the genus <i>Hepatica</i> comes from the shape of the leaf, +resembling that of the liver. <i>Myosurus</i> means mouse-tail. <i>Delphinium</i> +is from delphin, a dolphin, and alludes to the shape of the flower, +which was thought to resemble the classical figures of the dolphin. +<i>Xanthorrhiza</i> is from two Greek words meaning yellow-root, the common +name of the plant. <i>Cimicifuga</i> is formed of two Latin words meaning to +drive away bugs, i. e. Bugbane, the Siberian species being used to keep +away such vermin. <i>Sanguinaria</i>, the Bloodroot, is named from the +blood-like color of its juice. Other genera are dedicated to +distinguished botanists or promoters of science, and bear their names: +such are <i>Magnolia</i>, which commemorates the early French botanist, +Magnol; and <i>Jeffersonia</i>, named after President Jefferson, who sent the +first exploring expedition over the Rocky Mountains. Others bear the +name of the discoverer of the plant; as, <i>Sarracenia</i>, dedicated to Dr. +Sarrazin, of Quebec, who was one of the first to send the common +Pitcher-plant to the botanists of Europe; and <i>Claytonia</i>, first made +known by the early Virginian botanist Clayton.</p> + +<p><a name="UNIT_537" id="UNIT_537">537.</a> <b>Names of Species.</b> The name of a species is also a single word, +appended to that of the genus. It is commonly an adjective, and +therefore agrees with the generic name in case, gender, etc. Sometimes +it relates to the country the species inhabits; as, Claytonia +<i>Virginica</i>, first made known from Virginia; Sanguinaria <i>Canadensis</i>, +from Canada, etc. More commonly it denotes some obvious or +characteristic trait of the species; as, for example, in Sarracenia, our +northern species is named <i>purpurea</i>, from the purple blossoms, while a +more southern one is named <i>flava</i>, because its petals are yellow; the +species of Jeffersonia is called <i>diphylla</i>, meaning two-leaved, because +its leaf is divided into two leaflets. Some species are named after the +discoverer, or in compliment to a botanist who has made them known; as, +Magnolia <i>Fraseri</i>, named after the botanist Fraser, one <span class="pagenum"><a name="Page_180" id="Page_180">[Pg 180]</a></span>of the first +to find this species; and Sarracenia <i>Drummondii</i>, for a Pitcher-plant +found by Mr. Drummond in Florida. Such personal specific names are of +course written with a capital initial letter. Occasionally some old +substantive name is used for the species; as Magnolia <i>Umbrella</i>, the +Umbrella tree, and Ranunculus <i>Flammula</i>. These are also written with a +capital initial, and need not accord with the generic name in gender. +Geographical specific names, such as <i>Canadensis</i>, <i>Caroliniana</i>, +<i>Americana</i>, in the later usage are by some written without a capital +initial, but the older usage is better, or at least more accordant with +English orthography.</p> + +<p><a name="UNIT_538" id="UNIT_538">538.</a> <b>Varietal Names</b>, when any are required, are made on the plan of +specific names, and follow these, with the prefix <i>var</i>. Ranunculus +Flammula, var. <i>reptans</i>, the creeping variety: R. abortivus, var. +<i>micranthus</i>, the small-flowered variety of the species.</p> + +<p>539. In recording the name of a plant it is usual to append the name, or +an abbreviation of the name, of the botanist who first published it; and +in a flora or other systematic work, this reference to the source of the +name is completed by a further citation of the name of the book, the +volume and page where it was first published. So "<i>Ranunculus acris</i>, +L.," means that this Buttercup was first so named and described by +Linnæus; "<i>R. multifidus</i>, Pursh," that this species was so named and +published by Pursh. The suffix is no part of the name, but is an +abbreviated reference, to be added or omitted as convenience or +definiteness may require. The authority for a generic name is similarly +recorded. Thus, "<i>Ranunculus</i>, L.," means that the genus was so named by +Linnæus; "<i>Myosurus</i>, Dill.," that the Mouse-tail was established as a +genus under this name by Dillenius; <i>Caulophyllum</i>, Michx., that the +Blue Cohosh was published under this name by Michaux. The full reference +in the last-named instance would be, "in Flora Boreali-Americana, first +volume, 205th page,"—in the customary abbreviation, "Michx. Fl. i. +205."</p> + +<p><a name="UNIT_540" id="UNIT_540">540.</a> <b>Names of Orders</b> are given in the plural number, and are commonly +formed by prolonging the name of a genus of the group taken as a +representative of it. For example, the order of which the Buttercup or +Crowfoot genus, <i>Ranunculus</i>, is the representative, takes from it the +name of <i>Ranunculaceæ</i>; meaning <i>Plantæ Ranunculaceæ</i> when written out +in full, that is, Ranunculaceous Plants. Some old descriptive names of +orders are kept up, such as <i>Cruciferæ</i> for the order to which Cress and +Mustard belong, from the cruciform appearance of their expanded corolla, +and <i>Umbelliferæ</i>, from the flowers being in umbels.</p> + +<p><a name="UNIT_541" id="UNIT_541">541.</a> <b>Names of Tribes</b>, also of suborders, subtribes, and the like, are +plurals of the name of the typical genus, less prolonged, usually in +<i>eæ</i>, <i>ineæ</i>, <i>ideæ</i>, etc. Thus the proper Buttercup tribe is +<i>Ranunculeæ</i>, of the Clematis tribe, <i>Clematideæ</i>. While the Rose family +is <i>Rosaceæ</i>, the special Rose tribe is <i>Roseæ</i>.</p> + +<p><a name="UNIT_542" id="UNIT_542">542.</a> <b>Names of Classes, etc.</b> For these see the following synopsis of the +actual classification adopted, p. <a href="#Page_183">183</a>.</p> + +<p><span class="pagenum"><a name="Page_181" id="Page_181">[Pg 181]</a></span></p><p>543. So a plant is named in two words, the generic and the specific +names, to which may be added a third, that of the variety, upon +occasion. The generic name is peculiar: obviously it must not be used +twice over in botany. The specific name must not be used twice over in +the same genus, but is free for any other genus. A <i>Quercus alba</i>, or +White Oak, is no hindrance to <i>Betula alba</i>, or White Birch; and so of +other names.</p> + +<p><a name="UNIT_544" id="UNIT_544">544.</a> <b>Characters and Descriptions.</b> Plants are <i>characterized</i> by a terse +statement, in botanical terms, of their peculiarities or distinguishing +marks. The character of the order should include nothing which is common +to the whole class it belongs to; that of the genus, nothing which is +common to the order; that of the species nothing which is shared with +all other species of the genus; and so of other divisions. +<i>Descriptions</i> may enter into complete details of the whole structure.</p> + +<p><a name="UNIT_545" id="UNIT_545">545.</a> <b>Terminology</b>, also called <i>Glossology</i>, is nomenclature applied to +organs or parts, and their forms or modifications. Each organ or special +part has a substantive name of its own: shapes and other modifications +of an organ or part are designated by adjective terms, or, when the +forms are peculiar, substantive names are given to them. By the correct +use of such botanical terms, and by proper subordination of the +characters under the order, genus, species, etc., plants may be +described and determined with much precision. The classical language of +botany is Latin. While modern languages have their own names and terms, +these usually lack the precision of the Latin or Latinized botanical +terminology. Fortunately, this Latinized terminology has been largely +adopted and incorporated into the English technical language of botany, +thus securing precision. And these terms are largely the basis of +specific names of plants.</p> + +<p>546. A glossary or vocabulary of the principal botanical terms used in +phanerogamous and vascular cryptogamous botany is appended to this +volume, to which the student may refer, as occasion arises.</p> + + +<h3><a name="Subsect_XVIII_3" id="Subsect_XVIII_3">§ 3.</a> SYSTEM.</h3> + +<p>547. Two systems of classification used to be recognized in botany,—the +artificial and the natural; but only the latter is now thought to +deserve the name of a system.</p> + +<p>548. Artificial classifications have for object merely the ascertaining +of the name and place of a plant. They do not attempt to express +relationships, but serve as a kind of dictionary. They distribute the +genera and species according to some one peculiarity or set of +peculiarities (just as a dictionary distributes words according to their +first letters), disregarding all other considerations. At present an +artificial classification in botany is needed only as a key to the +natural orders,—as an aid in referring an unknown plant to its proper +family; and such keys are still very needful, at least for the beginner. +Formerly, when the orders themselves were not clearly made out, an +artificial classification was required to lead the <span class="pagenum"><a name="Page_182" id="Page_182">[Pg 182]</a></span>student down to the +genus. Two such classifications were long in vogue: First, that of +Tournefort, founded mainly on the leaves of the flower, the calyx and +corolla: this was the prevalent system throughout the first half of the +eighteenth century; but it has long since gone by. It was succeeded by +the well-known</p> + +<p><a name="UNIT_549" id="UNIT_549">549.</a> <b>Artificial System of Linnæus</b>, which was founded on the stamens and +pistils. It consists of twenty-four classes, and of a variable number of +orders; the classes founded mainly on the number and disposition of the +stamens; the orders partly upon the number of styles or stigmas, partly +upon other considerations. Useful and popular as this system was down to +a time within the memory of still surviving botanists, it is now +completely obsolete. But the tradition of it survives in the names of +its classes, Monandria, Diandria, Triandria, etc., which are familiar in +terminology in the adjective terms monandrous, diandrous, triandrous, +etc. (<a href="#UNIT_284">284</a>); also of the orders, Monogynia, Digynia, Trigynia, etc., +preserved in the form of monogynous, digynous, trigynous, etc. (<a href="#UNIT_301">301</a>); +and in the name Cryptogamia, that of the 24th class, which is continued +for the lower series in the natural classification.</p> + +<p><a name="UNIT_550" id="UNIT_550">550.</a> <b>Natural System.</b> A genuine system of botany consists of the orders +or families, duly arranged under their classes, and having the tribes, +the genera, and the species arranged in them according to their +relationships. This, when properly carried out, is the <i>Natural System</i>; +because it is intended to express, as well as possible, the various +degrees of relationship among plants, as presented in nature; that is, +to rank those species and those genera, etc., next to each other in the +classification which are really most alike in all respects, or, in other +words, which are constructed most nearly on the same particular plan.</p> + +<p>551. There can be only <i>one</i> natural system of botany, if by this term +is meant the plan according to which the vegetable creation was called +into being, with all its grades and diversities among the species, as +well of past as of the present time. But there may be many natural +systems, if we mean the attempts of men to interpret and express that +plan,—systems which will vary with advancing knowledge, and with the +judgment and skill of different botanists. These must all be very +imperfect, bear the impress of individual minds, and be shaped by the +current philosophy of the age. But the endeavor always is to make the +classification answer to Nature, as far as any system can which has to +be expressed in a definite and serial arrangement.</p> + +<p>552. So, although the classes, orders, genera, etc., are natural, or as +natural as the systematist can make them, their grouping or order of +arrangement in a book, must necessarily be in great measure artificial. +Indeed, it is quite impossible to arrange the orders, or even the few +classes, in a single series, and yet have each group stand next to its +nearest relatives on both sides.</p> + +<p>553. Especially it should be understood that, although phanerogamous +<span class="pagenum"><a name="Page_183" id="Page_183">[Pg 183]</a></span>plants are of higher grade than cryptogamous, and angiospermous or +ordinary phanerogamous higher than the gymnospermous, yet there is no +culmination in the vegetable kingdom, nor any highest or lowest order of +phanerogamous plants.</p> + +<p><a name="UNIT_554" id="UNIT_554">554.</a> The particular system most largely used at present in the +classification of the orders is essentially the following:—</p> +<ul class="lsoff" style="margin-left:1em;text-indent:-1em;"> +<li><span class="smcap">Series</span> I. PHANEROGAMIA: <span class="smcap">Phanerogamous or Flowering Plants</span>. + +<ul class="lsoff"> +<li><span class="smcap">Class</span> I. DICOTYLEDONES ANGIOSPERMEÆ, called for shortness in +English, <span class="smcap">Dicotyledons</span> or <span class="smcap">Dicotyls</span>. Ovules in a closed ovary. +Embryo dicotyledonous. Stem with exogenous plan of growth. Leaves +reticulate-veined, + +<ul class="lsoff"> +<li><i>Artificial Division I.</i> <span class="smcap">Polypetalæ</span>, with petals mostly present +and distinct. Orders about 80 in number, <i>Ranunculaceæ</i> to +<i>Cornaceæ</i>. +</li> + +<li><i>Artificial Division II.</i> <span class="smcap">Gamopetalæ</span>, with gamopetalous corolla. +Orders about 45, <i>Caprifoliaceæ</i> to <i>Plantaginaceæ</i>. +</li> + +<li><i>Artificial Division III.</i> <span class="smcap">Apetalæ</span> or <span class="smcap">Incompletæ</span>, with perianth, +when present, of calyx only. Orders about 35 in number, from +<i>Nyctaginaceæ</i> to <i>Salicaceæ</i>. +</li></ul></li> + +<li><span class="smcap">Class</span> II. DICOTYLEDONES GYMNOSPERMEÆ, in English <span class="smcap">Gymnosperms</span>. No +ovary or pericarp, but ovules and seeds naked, and no proper calyx +nor corolla. Embryo dicotyledonous or polycotyledonous. Stem with +exogenous plan of growth. Leaves mostly parallel-veined. Consists +of order <i>Gnetaceæ</i>, which strictly connects with Angiospermous +Dicotyls, of <i>Coniferæ</i>, and of <i>Cycadaceæ</i>. +</li> + +<li><span class="smcap">Class</span> III. MONOCOTYLEDONES, in English <span class="smcap">Monocotyledons</span> or +<span class="smcap">Monocotyls</span>. Angiospermous. Embryo monocotyledonous. Stem with +endogenous plan of growth. Leaves mostly parallel-veined. + +<ul class="lsoff"> +<li><i>Division I.</i> <span class="smcap">Petaloideæ</span>. Perianth complete, having the +equivalent of both calyx and corolla, and all the inner series +corolline. About 18 orders. +</li> + +<li><i>Division II.</i> <span class="smcap">Calycinæ</span>. Perianth complete (in two series) but +not corolline, mostly thickish or glumaceous. Chiefly two +orders, <i>Juncaceæ</i>, the true Rushes, and <i>Palmæ</i>, Palms. +</li> + +<li><i>Division III.</i> <span class="smcap">Spadicifloræ</span> or <span class="smcap">Nudifloræ</span>. Perianth none, or +rudimentary and incomplete: inflorescence spadiceous. Of five +orders, <i>Typhaceæ</i> and <i>Aroideæ</i> the principal. +</li> + +<li><i>Division IV.</i> <span class="smcap">Glumaceæ</span>. Perianth none, or very rudimentary: +glumaceous bracts to the flowers. Orders mainly <i>Cyperaceæ</i> and +<i>Gramineæ</i>. +</li></ul></li></ul></li> + +<li><span class="smcap">Series</span> II. CRYPTOGAMIA: <span class="smcap">Cryptogamous or Flowerless Plants</span>. + +<ul class="lsoff"> +<li>CLASS I. PTERIDOPHYTA, <span class="smcap">Pteridophytes</span> (<a href="#UNIT_484">484</a>). +</li> + +<li>CLASS II. BRYOPHYTA, <span class="smcap">Bryophytes</span> (<a href="#UNIT_498">498</a>). +</li> + +<li>CLASS III. THALLOPHYTA, <span class="smcap">Thallophytes</span> (<a href="#UNIT_503">503</a>). +</li></ul></li></ul> + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_184" id="Page_184">[Pg 184]</a></span></p> +<h2><a name="Section_XIX" id="Section_XIX"></a>SECTION XIX. BOTANICAL WORK.</h2> + + +<p>555. Some hints and brief instructions for the collection, examination, +and preservation of specimens are added. They are especially intended +for the assistance of those who have not the advantage of a teacher. +They apply to phanerogamous plants and Ferns only, and to systematic +botany.<a name="FNanchor_1_4" id="FNanchor_1_4"></a><a href="#Footnote_1_4" class="fnanchor">[1]</a></p> + + +<h3><a name="Subsect_XIX_1" id="Subsect_XIX_1">§ 1.</a> COLLECTION, OR HERBORIZATION.</h3> + +<p>556. As much as possible, plants should be examined in the living state, +or when freshly gathered. But dried specimens should be prepared for +more leisurely examination and for comparison. To the working botanist +good dried specimens are indispensable.</p> + +<p><a name="UNIT_557" id="UNIT_557">557.</a> <b>Botanical Specimens</b>, to be complete, should have root or rootstock, +stem, leaves, flowers, both open and in bud, and fruit. Sometimes these +may all be obtained at one gathering; more commonly two or three +gatherings at different times are requisite, especially for trees and +shrubs.</p> + +<p><a name="UNIT_558" id="UNIT_558">558.</a> <b>In Herborizing</b>, a good knife and a narrow and strong trowel are +needed; but a very strong knife will serve instead of a trowel or small +pick for digging out bulbs, tubers, and the like. To carry the +specimens, either the tin box (<i>vasculum</i>) or a portfolio, or both are +required. The tin box is best for the collection of specimens to be used +fresh, as in the class-room; also for very thick or fleshy plants. The +portfolio is indispensable for long expeditions, and is best for +specimens which are to be preserved in the herbarium.</p> + +<p>559. The <i>Vasculum</i>, or <i>Botanical Collecting-box</i>, is made of tin, in +shape like a candle-box, only flatter, or the smaller sizes like an +English sandwich-case; the lid opening for nearly the whole length of +one side of the box. Any portable tin box of convenient size, and +capable of holding specimens a foot or fifteen inches long, will answer +the purpose. The box should shut close, so that the specimens may not +wilt: then it will keep leafy branches and most flowers perfectly fresh +for a day or two, especially if slightly moistened. They should not be +wet.</p> + +<p>560. <i>The Portfolio</i> is best made of two pieces of solid binder's-board, +covered with enamel cloth, which also forms the back, and fastened by +straps and buckles. It may be from a foot to twenty inches long, from +nine to eleven or twelve inches wide. It should contain a needful +quantity of smooth but strong and pliable paper (thin so-called Manilla +paper is best), either fastened at the back as in a book, or loose in +folded sheets when not very many specimens are required. As soon as +gathered, the specimens should be separately laid between the leaves or +in the folded sheets, and kept under moderate pressure in the closed +portfolio.</p> + +<p><span class="pagenum"><a name="Page_185" id="Page_185">[Pg 185]</a></span></p><p>561. Of small herbs, especially annuals, the whole plant, root and all, +should be taken for a specimen. Of larger ones branches will suffice, +with some leaves from near the root. Enough of the root or subterranean +part of the plant should be collected to show whether it is an annual, a +biennial, or a perennial. Thick roots, bulbs, tubers, or branches of +specimens intended to be pressed should be thinned with a knife, or cut +into slices. Keep the specimens within the length of fifteen or sixteen +inches, by folding, or when that cannot be done, by cutting into +lengths.</p> + +<p><a name="UNIT_562" id="UNIT_562">562.</a> <b>For Drying Specimens</b> a good supply of soft and unsized paper is +wanted; and some convenient means of applying considerable pressure. To +make good dried botanical specimens, dry them as rapidly as possible +between many thicknesses of sun-dried paper to absorb their moisture, +under as much pressure as can be given without crushing the more +delicate parts. This pressure may be had by a botanical press, of which +various forms have been contrived; or by weights placed upon a +board,—from forty to eighty or a hundred pounds, according to the +quantity of specimens drying at the time. For use while travelling, a +good portable press may be made of thick binders' boards for the sides, +and the pressure may be applied by strong straps with buckles. Still +better, on some accounts, are portable presses made of wire network, +which allow the dampness to escape by evaporation between the meshes. +For herborization in a small way, a light wire-press may be taken into +the field and made to serve also as a portfolio.</p> + +<p>563. It is well to have two kinds of paper, namely, <i>driers</i> of bibulous +paper, stitched into pads (or the pads may be of thick carpet-paper, cut +to size) and thin smooth paper, folded once; the specimens to be laid +into the fold, either when gathered or on returning from the excursion. +These sheets are to hold the specimens until they are quite dry. Every +day, or at first even twice a day, the specimens, left undisturbed in +their sheets, are to be shifted into fire-dried or sun-dried fresh +driers, and the pressure renewed, while the moist sheets are spread out +to dry, so as to take their turn again at the next shifting. This course +must be continued until the specimens are no longer moist to the touch. +Good and comely specimens are either made or spoiled within the first +twenty-four or thirty-six hours. After that, when plenty of driers are +used, it may not be necessary to change them so frequently.</p> + +<p>564. Succulent plants, which long refuse to part with life and moisture, +and Spruces and some other evergreens which are apt to cast off their +leaves, may be plunged for a moment into boiling water, all but the +flowers. Delicate flowers may be encased in thin tissue paper when put +into the press. Thick parts, like the heads of Sunflowers and Thistles, +may be cut in two or into slices.</p> + +<p>565. Dried specimens may be packed in bundles, either in folded paper or +upon single half-sheets. It is better that such paper should not be +bibulous. The packages should be well wrapped or kept in close cases.</p> + +<p><span class="pagenum"><a name="Page_186" id="Page_186">[Pg 186]</a></span></p><p><a name="UNIT_566" id="UNIT_566">566.</a> <b>Poisoning</b> is necessary if specimens are to be permanently +preserved from the depredation of insects. The usual application is an +almost saturated solution of corrosive sublimate in 95 per cent alcohol, +freely applied with a large and soft brush, or the specimens dipped into +some of the solution poured into a large and flat dish; the wetted +specimens to be transferred for a short time to driers.</p> + + +<h3><a name="Subsect_XIX_2" id="Subsect_XIX_2">§ 2.</a> HERBARIUM.</h3> + +<p>567. The botanist's collection of dried specimens, ticketed with their +names, place, and time of collection, and systematically arranged under +their genera, orders, etc., forms a <i>Hortus Siccus</i> or <i>Herbarium</i>. It +comprises not only the specimens which the proprietor has himself +collected, but those which he acquires through friendly exchanges, or in +other ways. The specimens of an herbarium may be kept in folded sheets +of paper; or they may be fastened on half-sheets of thick and white +paper, either by gummed slips, or by glue applied to the specimens +themselves. The former is best for private and small herbaria; the +latter for large ones which are much turned over. Each sheet should be +appropriated to one species; two or more different plants should never +be attached to the same sheet. The generic and specific name of the +plant should be added to the lower right-hand corner, either written on +the sheet, or on a ticket pasted down; and the time of collection, the +locality, the color of the flowers, and any other information which the +specimens themselves do not afford, should be duly recorded upon the +sheet or the ticket. The sheets of the herbarium should all be of +exactly the same dimensions. The herbarium of Linnæus is on paper of the +common foolscap size, about eleven inches long and seven wide. This is +too small. Sixteen and three eighths inches by eleven and a half inches +is an approved size.</p> + +<p>568. The sheets containing the species of each genus are to be placed in +<i>genus-covers</i>, made of a full sheet of thick paper (such as the +strongest Manilla-hemp paper), to be when folded of the same dimensions +as the species-sheet but slightly wider: the name of the genus is to be +written on one of the lower corners. These are to be arranged under the +orders to which they belong, and the whole kept in closed cases or +cabinets, either laid flat in compartments, like "pigeon-holes," or else +placed in thick portfolios, arranged like folio volumes. All should be +kept, as much as practicable, in dust-proof and insect-proof cases or +boxes.</p> + +<p>569. Fruits, tubers, and other hard parts, too thick for the herbarium, +may be kept in pasteboard or light wooden boxes, in a collection apart. +Small loose fruits, seeds, detached flowers, and the like may be +conveniently preserved in paper capsules or envelopes, attached to the +herbarium-sheets.</p> + + +<p><span class="pagenum"><a name="Page_187" id="Page_187">[Pg 187]</a></span></p><h3><a name="Subsect_XIX_3" id="Subsect_XIX_3">§ 3.</a> INVESTIGATION AND DETERMINATION OF PLANTS.</h3> + +<p><a name="UNIT_570" id="UNIT_570">570.</a> <b>The Implements</b> required are a hand magnifying glass, a pocket lens +of an inch or two focus, or a glass of two lenses, one of the lower and +the other of the higher power; and a sharp penknife for dissection. With +these and reasonable perseverance the structure of the flowers and +fructification of most phanerogamous plants and Ferns can be made out. +But for ease and comfort, as well as for certainty and right training, +the student should have some kind of simple stage microscope, and under +this make all dissections of small parts. Without it the student will be +apt to fall into the bad habit of guessing where he ought to ascertain.</p> + +<p>571. The simple microscope may be reduced to a good lens or doublet, of +an inch focus, mounted over a glass stage, so that it can be moved up +and down and also sidewise, and with (or without) a little mirror +underneath. A better one would have one or two additional lenses (say of +half and of a quarter inch focus), a pretty large stage, on the glass of +which several small objects can be placed and conveniently brought under +the lens; and its height or that of the lens should be adjustable by a +rack-work; also a swivel-mounted little mirror beneath, which is needed +for minute objects to be viewed by transmitted light.</p> + +<p>572. For dissecting and displaying small parts on the stage of the +microscope, besides a thin-bladed knife, the only tools needed are a +good stock of common needles of various sizes, mounted in handles, and +one or more saddler's-needles, which, being triangular, may be ground to +sharp edges convenient for dissection. Also a pair of delicate-pointed +forceps; those with curved points used by the dentist are most +convenient. A cup of clean water is indispensable, with which to moisten +or wet, or in which occasionally to float delicate parts. Small flowers, +buds, fruits, and seeds of dried specimens can be dissected quite as +well as fresh ones. They have only to be soaked in warm or boiling +water.</p> + +<p>573. The compound microscope is rarely necessary except in cryptogamic +botany and vegetable anatomy; but it is very useful and convenient, +especially for the examination of pollen. To the advanced botanist it is +a necessity, to all students of botany an aid and delight.</p> + +<p><a name="UNIT_574" id="UNIT_574">574.</a> <b>Analysis.</b> A few directions and hints may be given. The most +important is this: In studying an unknown plant, make a complete +examination of all its parts, and form a clear idea of its floral +structure and that of its fruit, from pericarp down to the embryo, or as +far as the materials in hand allow, before taking a step toward finding +out its name and relationship by means of the keys or other helps which +the Manuals and Floras provide. If it is the name merely that is wanted, +the shorter way is to ask some one who already knows it. To verify the +points of structure one by one as they happen to occur in an artificial +key, without any preparatory investigation, is a usual but is not the +best nor the surest <span class="pagenum"><a name="Page_188" id="Page_188">[Pg 188]</a></span>way. It is well to make drawings or outline +sketches of the smaller parts, and especially diagrams of the plan of +the flower, such as those of Fig. <a href="#Fig224">225</a>, <a href="#Fig226">227</a>, <a href="#Fig239">241, 244</a>, <a href="#Fig275">275</a>-<a href="#Fig277">277</a>. For +these, cross sections of the flower-bud or flower are to be made: and +longitudinal sections, such as Fig. <a href="#Fig270">270</a>-<a href="#Fig273">274</a>, are equally important. The +dissection even of small seeds is not difficult after some practice. +Commonly they need to be soaked or boiled.</p> + +<p>575. The right appreciation of characters and terms used in description +needs practice and calls for judgment. Plants do not grow exactly by +rule and plummet, and measurements must be taken loosely. Difference of +soil and situation are responded to by considerable variations, and +other divergences occur which cannot be accounted for by the +surroundings, nor be anticipated in general descriptions. Annuals may be +very depauperate in dry soils or seasons, or very large when +particularly well nourished. Warm and arid situations promote, and wet +ones are apt to diminish pubescence. Salt water causes increased +succulence. The color of flowers is apt to be lighter in shade, and +brighter in open and elevated situations. A color or hue not normal to +the species now and then occurs, which nothing in the conditions will +account for. <i>A white-flowered variation of any other colored blossom +may always be expected</i>; this, though it may be notable, no more +indicates a distinct variety of the species than an albino would a +variety of the human species. The numerical plan is subject to variation +in some flowers; those on the plan of five may now and then vary to four +or to six. Variations of the outline or lobing of leaves are so familiar +that they do not much mislead. Only wider and longer observation +suffices to prevent or correct mistakes in botanical study. But the +weighing of evidence and the balancing of probabilities, no less than +the use of the well-ordered and logical system of classification, give +as excellent training to the judgment as the search for the facts +themselves does to the observing powers.</p> + + +<h3><a name="Subsect_XIX_4" id="Subsect_XIX_4">§ 4.</a> SIGNS AND ABBREVIATIONS.</h3> + +<p>576. For a full account of these, whether of former or actual use, see +"Structural Botany" of the "Botanical Text Book," pp. 367, 392, as also +for the principles which govern the accentuation of names. It is needful +here to explain only those used in the Manuals and Floras of this +country, for which the present volume is an introduction and companion. +They are not numerous.</p> + +<p>577. In arranging the species, at least those of a large genus, the +divisions are denoted and graduated as follows: The sign § is prefixed +to sections of the highest rank: these sections when they have names +affixed to them (as <span class="smcap">Prunus</span> § <span class="smcap">Cerasus</span>) may be called subgenera. When the +divisions of a genus are not of such importance, or when divisions are +made under the subgenus itself, the most comprehensive ones are marked +by asterisks, * for the first, * * for the second, and so on. +Subdivisions are <span class="pagenum"><a name="Page_189" id="Page_189">[Pg 189]</a></span>marked with a prefixed +; those under this head with +++; and those under this with =, if there be so many grades. A similar +notation is followed in the synopsis of the genera of an order.</p> + +<p>578. The interrogation point is used in botany to indicate doubt. Thus +<i>Clematis crispa</i>, L.? expresses a doubt whether the plant in question +is really the <i>Clematis crispa</i> of Linnæus. <i>Clematis? polypetala</i> +expresses a doubt whether the plant so named is really a Clematis. On +the other hand the exclamation point (!) is used to denote certainty +whenever there is special need to affirm this.</p> + +<p>579. For size or height, the common signs of degrees, minutes, and +seconds, have been used, thus, 1°, 2', 3", stand respectively for a +foot, two inches, and three lines or twelfths of an inch. A better way, +when such brevity is needed, is to write 1<sup>ft</sup>. 2<sup>in</sup>. 3<sup>l</sup>.</p> + +<p>580. Signs for duration used by Linnæus were ☉ for an annual, +♂ for a biennial, ♃ for a perennial herb, +<a href="images/figsymbols.jpg">[Symbol like numeral 5 without top bar]</a> for a shrub or tree. DeCandolle brought in ☉ for a +plant that died after once flowering, ① if annual, ② if biennial.</p> + +<p>581. To indicate sexes, ♂ means staminate or male plant or +blossom; ♀, pistillate or female; <a href="images/figsymbols.jpg">[Symbol like ☿ with two inverted breves]</a>, perfect or +hermaphrodite.</p> + +<p>582. To save room it is not uncommon to use ∞ in place of +"many;" thus, "Stamens ∞," for stamens indefinitely numerous: +"∞ flora" for pluriflora or many-flowered. Still more common +is the form "Stamens 5-20," or "Calyx 4-5-parted," for stamens from five +to twenty, calyx four-parted or five-parted, and the like. Such +abbreviations hardly need explanation.</p> + +<p>583. The same may be said of such abbreviations as <i>Cal.</i> for calyx, +<i>Cor.</i> for corolla, <i>Pet.</i> for petals, <i>St.</i> for stamens, <i>Pist.</i> for +pistil, <i>Hab.</i> for habitat, meaning place of growth, <i>Herb.</i> for +herbarium, <i>Hort.</i> for garden. Also <i>l. c.</i>, loco citato, which avoids +repetition of volume and page.</p> + +<p>584. "Structural Botany" has six pages of abbreviations of the names of +botanists, mostly of botanical authors. As they are not of much +consequence to the beginner, while the more advanced botanist will know +the names in full, or know where to find them, only a selection is here +appended.</p> + + +<div class="footnotes"><h3>FOOTNOTES:</h3> + +<div class="footnote"><p><a name="Footnote_1_4" id="Footnote_1_4"></a><a href="#FNanchor_1_4"><span class="label">[1]</span></a> For fuller directions in many particulars, see "Structural +Botany," pp. 370-374.</p></div> +</div> + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_190" id="Page_190">[Pg 190]</a></span></p> +<h2><a name="ABBREVIATIONS" id="ABBREVIATIONS"></a>ABBREVIATIONS OF THE NAMES OF BOTANISTS.</h2> + + + + +<div class="center"> +<table border="0" cellpadding="1" cellspacing="0" summary=""> +<tr><td align="left"><i>Adans.</i></td><td align="center" style="width:2em;">=</td><td align="left">Adanson.</td></tr> +<tr><td align="left"><i>Ait.</i></td><td align="center"> </td><td align="left">Aiton.</td></tr> +<tr><td align="left"><i>All.</i></td><td align="center"> </td><td align="left">Allioni.</td></tr> +<tr><td align="left"><i>Andr.</i></td><td align="center"> </td><td align="left">Andrews.</td></tr> +<tr><td align="left"><i>Arn.</i></td><td align="center"> </td><td align="left">Arnott.</td></tr> +<tr><td align="left"><i>Aub.</i></td><td align="center"> </td><td align="left">Aublet.</td></tr> +<tr><td align="left"><i>Bartr.</i></td><td align="center"> </td><td align="left">Bartram.</td></tr> +<tr><td align="left"><i>Beauv.</i></td><td align="center"> </td><td align="left">Palisot de Beauvois.</td></tr> +<tr><td align="left"><i>Benth.</i></td><td align="center"> </td><td align="left">Bentham.</td></tr> +<tr><td align="left"><i>Bernh.</i></td><td align="center"> </td><td align="left">Bernhardi.</td></tr> +<tr><td align="left"><i>Bigel.</i></td><td align="center"> </td><td align="left">Jacob Bigelow.</td></tr> +<tr><td align="left"><i>Bong.</i></td><td align="center"> </td><td align="left">Bongard.</td></tr> +<tr><td align="left"><i>Bonpl.</i></td><td align="center"> </td><td align="left">Bonpland.</td></tr> +<tr><td align="left"><i>Br.</i> or <i>R. Br.</i></td><td align="center"> </td><td align="left">Robert Brown.</td></tr> +<tr><td align="left"><i>Cass.</i></td><td align="center"> </td><td align="left">Cassini.</td></tr> +<tr><td align="left"><i>Cav.</i></td><td align="center"> </td><td align="left">Cavanilles.</td></tr> +<tr><td align="left"><i>Cham.</i></td><td align="center"> </td><td align="left">Chamisso.</td></tr> +<tr><td align="left"><i>Chapm.</i></td><td align="center"> </td><td align="left">Chapman.</td></tr> +<tr><td align="left"><i>Chois.</i></td><td align="center"> </td><td align="left">Choisy.</td></tr> +<tr><td align="left"><i>Clayt.</i></td><td align="center"> </td><td align="left">Clayton.</td></tr> +<tr><td align="left"><i>Curt.</i></td><td align="center"> </td><td align="left">Curtis.</td></tr> +<tr><td align="left"><i>Curt. (M. A.)</i></td><td align="center"> </td><td align="left">M. A. Curtis.</td></tr> +<tr><td align="left"><i>Darl.</i></td><td align="center"> </td><td align="left">Darlington.</td></tr> +<tr><td align="left"><i>DC.</i></td><td align="center" rowspan="2" valign="middle"><span style="font-size:200%;">}</span></td><td align="left" rowspan="2" valign="middle">DeCandolle.</td></tr> +<tr><td align="left"><i>DeCand.</i></td></tr> +<tr><td align="left"><i>A. DC.</i></td><td align="center"> </td><td align="left">Alphonse DeCandolle.</td></tr> +<tr><td align="left"><i>Desc.</i></td><td align="center"> </td><td align="left">Descourtilz.</td></tr> +<tr><td align="left"><i>Desf.</i></td><td align="center"> </td><td align="left">Desfontaines.</td></tr> +<tr><td align="left"><i>Desv.</i></td><td align="center"> </td><td align="left">Desvaux.</td></tr> +<tr><td align="left"><i>Dill.</i></td><td align="center"> </td><td align="left">Dillenius.</td></tr> +<tr><td align="left"><i>Dougl.</i></td><td align="center"> </td><td align="left">Douglas.</td></tr> +<tr><td align="left"><i>Duham.</i></td><td align="center"> </td><td align="left">Duhamel.</td></tr> +<tr><td align="left"><i>Dun.</i></td><td align="center"> </td><td align="left">Dunal.</td></tr> +<tr><td align="left"><i>Eat.</i></td><td align="center"> </td><td align="left">Eaton (Amos) or D. C.</td></tr> +<tr><td align="left"><i>Ehrh.</i></td><td align="center"> </td><td align="left">Ehrhart.</td></tr> +<tr><td align="left"><i>Ell.</i></td><td align="center"> </td><td align="left">Elliott.</td></tr> +<tr><td align="left"><i>Endl.</i></td><td align="center"> </td><td align="left">Endlicher.</td></tr> +<tr><td align="left"><i>Engelm.</i></td><td align="center"> </td><td align="left">Engelmann.</td></tr> +<tr><td align="left"><i>Engl.</i></td><td align="center"> </td><td align="left">Engler.</td></tr> +<tr><td align="left"><i>Fisch.</i></td><td align="center"> </td><td align="left">Fischer.</td></tr> +<tr><td align="left"><i>Frœl.</i></td><td align="center"> </td><td align="left">Frœlich.</td></tr> +<tr><td align="left"><i>Gærtn.</i></td><td align="center"> </td><td align="left">Gærtner.</td></tr> +<tr><td align="left"><i>Gaud.</i></td><td align="center"> </td><td align="left">Gaudin.</td></tr> +<tr><td align="left"><i>Gaudich.</i></td><td align="center"> </td><td align="left">Gaudichaud.</td></tr> +<tr><td align="left"><i>Ging.</i></td><td align="center"> </td><td align="left">Gingins.</td></tr> +<tr><td align="left"><i>Gmel.</i></td><td align="center"> </td><td align="left">Gmelin.</td></tr> +<tr><td align="left"><i>Good.</i></td><td align="center"> </td><td align="left">Goodenough.</td></tr> +<tr><td align="left"><i>Grev.</i></td><td align="center"> </td><td align="left">Greville.</td></tr> +<tr><td align="left"><i>Griseb.</i></td><td align="center"> </td><td align="left">Grisebach.</td></tr> +<tr><td align="left"><i>Gron.</i></td><td align="center" rowspan="2" valign="middle"><span style="font-size:200%;">}</span></td><td align="left" rowspan="2" valign="middle">Gronovius.</td></tr> +<tr><td align="left"><i>Gronov.</i></td></tr> +<tr><td align="left"><i>Hall.</i></td><td align="center"> </td><td align="left">Haller.</td></tr> +<tr><td align="left"><i>Hartm.</i></td><td align="center"> </td><td align="left">Hartmann.</td></tr> +<tr><td align="left"><i>Hartw.</i></td><td align="center"> </td><td align="left">Hartweg.</td></tr> +<tr><td align="left"><i>Harv.</i></td><td align="center"> </td><td align="left">Harvey.</td></tr> +<tr><td align="left"><i>Haw.</i></td><td align="center"> </td><td align="left">Haworth.</td></tr> +<tr><td align="left"><i>Hegelm.</i></td><td align="center"> </td><td align="left">Hegelmaier.</td></tr> +<tr><td align="left"><i>Hemsl.</i></td><td align="center"> </td><td align="left">Hemsley.</td></tr> +<tr><td align="left"><i>Herb.</i></td><td align="center"> </td><td align="left">Herbert.</td></tr> +<tr><td align="left"><i>Hoffm.</i></td><td align="center"> </td><td align="left">Hoffmann.</td></tr> +<tr><td align="left"><i>Hoffmans.</i></td><td align="center"> </td><td align="left">Hoffmansegg.</td></tr> +<tr><td align="left"><i>Hook.</i></td><td align="center"> </td><td align="left">Hooker.</td></tr> +<tr><td align="left"><i>Hook. f.</i></td><td align="center"> </td><td align="left">J. D. Hooker.</td></tr> +<tr><td align="left"><i>Hornem.</i></td><td align="center"> </td><td align="left">Hornemann.</td></tr> +<tr><td align="left"><i>Huds.</i></td><td align="center"> </td><td align="left">Hudson.</td></tr> +<tr><td align="left"><i>Humb.</i></td><td align="center"> </td><td align="left">Humboldt.</td></tr> +<tr><td align="left"><i>HBK.</i></td><td align="center"> </td><td align="left">Humboldt, Bonpland, and Kunth.</td></tr> +<tr><td align="left"><i>Jacq.</i></td><td align="center"> </td><td align="left">Jacquin.</td></tr> +<tr><td align="left"><i>Jacq. f.</i></td><td align="center"> </td><td align="left">J. F. Jacquin.</td></tr> +<tr><td align="left"><i>Juss.</i></td><td align="center"> </td><td align="left">Jussieu.</td></tr> +<tr><td align="left"><i>A. Juss.</i></td><td align="center"> </td><td align="left">Adrien de Jussieu.</td></tr> +<tr><td align="left"><i>Kit.</i></td><td align="center"> </td><td align="left">Kitaibel.</td></tr> +<tr><td align="left"><i>L.</i> or <i>Linn.</i></td><td align="center"> </td><td align="left">Linnæus.</td></tr> +<tr><td align="left"><i>Labill.</i></td><td align="center"> </td><td align="left">Labillardiere.</td></tr> +<tr><td align="left"><i>Lag.</i></td><td align="center"> </td><td align="left">Lagasca.</td></tr> +<tr><td align="left"><i>Lam.</i></td><td align="center"> </td><td align="left">Lamarck.</td></tr> +<tr><td align="left"><i>Ledeb.</i></td><td align="center"> </td><td align="left">Ledebour.</td></tr> +<tr><td align="left"><i>Lehm.</i></td><td align="center"> </td><td align="left">Lehmann.</td></tr> +<tr><td align="left"><i>Lesq.</i></td><td align="center"> </td><td align="left">Lesquereux.</td></tr> +<tr><td align="left"><i>Less.</i></td><td align="center"> </td><td align="left">Lessing.</td></tr> +<tr><td align="left"><i>Lestib.</i></td><td align="center"> </td><td align="left">Lestibudois.</td></tr> +<tr><td align="left"><i>L'Her.</i></td><td align="center"> </td><td align="left">L'Heritier.</td></tr> +<tr><td align="left"><i>Lindb.</i></td><td align="center"> </td><td align="left">Lindberg.</td></tr> +<tr><td align="left"><i>Lindh.</i></td><td align="center"> </td><td align="left">Lindheimer.</td></tr> +<tr><td align="left"><i>Lindl.</i></td><td align="center"> </td><td align="left">Lindley.</td></tr> +<tr><td align="left"><i>Lodd.</i></td><td align="center"> </td><td align="left">Loddiges.</td></tr> +<tr><td align="left"><i>Loud.</i></td><td align="center"> </td><td align="left">Loudon.</td></tr> +<tr><td align="left"><i>M. Bieb.</i></td><td align="center"> </td><td align="left">Marschall von Bieberstein.</td></tr> +<tr><td align="left"><i>Marsh.</i></td><td align="center"> </td><td align="left">Marshall (Humphrey).</td></tr> +<tr><td align="left"><i>Mart.</i></td><td align="center"> </td><td align="left">Martius.</td></tr> +<tr><td align="left"><span class="pagenum"><a name="Page_191" id="Page_191">[Pg 191]</a></span><i>Mast.</i></td><td align="center">=</td><td align="left">Masters.</td></tr> +<tr><td align="left"><i>Maxim.</i></td><td align="center"> </td><td align="left">Maximowicz.</td></tr> +<tr><td align="left"><i>Meisn.</i></td><td align="center" rowspan="2" valign="middle"><span style="font-size:200%;">}</span></td><td align="left">Meisner or</td></tr> +<tr><td align="left"><i>Meissn.</i></td><td align="left">Meissner.</td></tr> +<tr><td align="left"><i>Michx.</i> or <i>Mx.</i></td><td align="center"> </td><td align="left">Michaux.</td></tr> +<tr><td align="left"><i>Michx. f.</i></td><td align="center"> </td><td align="left">F. A. Michaux.</td></tr> +<tr><td align="left"><i>Mill.</i></td><td align="center"> </td><td align="left">Miller.</td></tr> +<tr><td align="left"><i>Miq.</i></td><td align="center"> </td><td align="left">Miquel.</td></tr> +<tr><td align="left"><i>Mitch.</i></td><td align="center"> </td><td align="left">Mitchell.</td></tr> +<tr><td align="left"><i>Moç.</i></td><td align="center"> </td><td align="left">Moçino.</td></tr> +<tr><td align="left"><i>Moq.</i></td><td align="center"> </td><td align="left">Moquin-Tandon.</td></tr> +<tr><td align="left"><i>Moric.</i></td><td align="center"> </td><td align="left">Moricand.</td></tr> +<tr><td align="left"><i>Moris.</i></td><td align="center"> </td><td align="left">Morison.</td></tr> +<tr><td align="left"><i>Muell. Arg.</i></td><td align="center"> </td><td align="left">J. Mueller.</td></tr> +<tr><td align="left"><i>Muell. (F.)</i></td><td align="center"> </td><td align="left">Ferdinand Mueller.</td></tr> +<tr><td align="left"><i>Muhl.</i></td><td align="center"> </td><td align="left">Muhlenberg.</td></tr> +<tr><td align="left"><i>Murr.</i></td><td align="center"> </td><td align="left">Murray.</td></tr> +<tr><td align="left"><i>Naud.</i></td><td align="center"> </td><td align="left">Naudin.</td></tr> +<tr><td align="left"><i>Neck.</i></td><td align="center"> </td><td align="left">Necker.</td></tr> +<tr><td align="left"><i>Nees</i></td><td align="center" rowspan="2" valign="middle"><span style="font-size:200%;">}</span></td><td align="left" rowspan="2" valign="middle">Nees von Esenbeck.</td></tr> +<tr><td align="left"><i>N. ab E.</i></td></tr> +<tr><td align="left"><i>Nutt.</i></td><td align="center"> </td><td align="left">Nuttall.</td></tr> +<tr><td align="left"><i>Œd.</i></td><td align="center"> </td><td align="left">Œder.</td></tr> +<tr><td align="left"><i>Ort.</i></td><td align="center"> </td><td align="left">Ortega.</td></tr> +<tr><td align="left"><i>P. de Beauv.</i></td><td align="center"> </td><td align="left">Palisot de Beauvois.</td></tr> +<tr><td align="left"><i>Pall.</i></td><td align="center"> </td><td align="left">Pallas.</td></tr> +<tr><td align="left"><i>Parl.</i></td><td align="center"> </td><td align="left">Parlatore.</td></tr> +<tr><td align="left"><i>Pav.</i></td><td align="center"> </td><td align="left">Pavon.</td></tr> +<tr><td align="left"><i>Pers.</i></td><td align="center"> </td><td align="left">Persoon.</td></tr> +<tr><td align="left"><i>Planch.</i></td><td align="center"> </td><td align="left">Planchon.</td></tr> +<tr><td align="left"><i>Pluk.</i></td><td align="center"> </td><td align="left">Plukenet.</td></tr> +<tr><td align="left"><i>Plum.</i></td><td align="center"> </td><td align="left">Plumier.</td></tr> +<tr><td align="left"><i>Poir.</i></td><td align="center"> </td><td align="left">Poiret.</td></tr> +<tr><td align="left"><i>Radlk.</i></td><td align="center"> </td><td align="left">Radlkofer.</td></tr> +<tr><td align="left"><i>Raf.</i></td><td align="center"> </td><td align="left">Rafinesque.</td></tr> +<tr><td align="left"><i>Red.</i></td><td align="center"> </td><td align="left">Redouté.</td></tr> +<tr><td align="left"><i>Reichenb.</i></td><td align="center"> </td><td align="left">Reichenbach.</td></tr> +<tr><td align="left"><i>Rich.</i></td><td align="center"> </td><td align="left">L. C. Richard.</td></tr> +<tr><td align="left"><i>Rich. f.</i> or <i>A.</i></td><td align="center"> </td><td align="left">Achille Richard.</td></tr> +<tr><td align="left"><i>Richards.</i></td><td align="center"> </td><td align="left">Richardson.</td></tr> +<tr><td align="left"><i>Ridd.</i></td><td align="center"> </td><td align="left">Riddell.</td></tr> +<tr><td align="left"><i>Rœm. & Schult.</i></td><td align="center"> </td><td align="left">Rœmer & Schultes.</td></tr> +<tr><td align="left"><i>Rottb.</i></td><td align="center"> </td><td align="left">Rottbœll.</td></tr> +<tr><td align="left"><i>Rupr.</i></td><td align="center"> </td><td align="left">Ruprecht.</td></tr> +<tr><td align="left"><i>St. Hil.</i></td><td align="center"> </td><td align="left">Saint-Hilaire.</td></tr> +<tr><td align="left"><i>Salisb.</i></td><td align="center"> </td><td align="left">Salisbury.</td></tr> +<tr><td align="left"><i>Schk.</i></td><td align="center"> </td><td align="left">Schkuhr.</td></tr> +<tr><td align="left"><i>Schlecht.</i></td><td align="center"> </td><td align="left">Schlechtendal.</td></tr> +<tr><td align="left"><i>Schrad.</i></td><td align="center"> </td><td align="left">Schrader.</td></tr> +<tr><td align="left"><i>Schreb.</i></td><td align="center"> </td><td align="left">Schreber.</td></tr> +<tr><td align="left"><i>Schwein.</i></td><td align="center"> </td><td align="left">Schweinitz.</td></tr> +<tr><td align="left"><i>Scop.</i></td><td align="center"> </td><td align="left">Scopoli.</td></tr> +<tr><td align="left"><i>Spreng.</i></td><td align="center"> </td><td align="left">Sprengel.</td></tr> +<tr><td align="left"><i>Sternb.</i></td><td align="center"> </td><td align="left">Sternberg.</td></tr> +<tr><td align="left"><i>Steud.</i></td><td align="center"> </td><td align="left">Steudel.</td></tr> +<tr><td align="left"><i>Sull.</i></td><td align="center"> </td><td align="left">Sullivant.</td></tr> +<tr><td align="left"><i>Thunb.</i></td><td align="center"> </td><td align="left">Thunberg.</td></tr> +<tr><td align="left"><i>Torr.</i></td><td align="center"> </td><td align="left">Torrey.</td></tr> +<tr><td align="left"><i>Tourn.</i></td><td align="center"> </td><td align="left">Tournefort.</td></tr> +<tr><td align="left"><i>Trautv.</i></td><td align="center"> </td><td align="left">Trautvetter.</td></tr> +<tr><td align="left"><i>Trin.</i></td><td align="center"> </td><td align="left">Trinius.</td></tr> +<tr><td align="left"><i>Tuck.</i></td><td align="center"> </td><td align="left">Tuckerman.</td></tr> +<tr><td align="left"><i>Vaill.</i></td><td align="center"> </td><td align="left">Vaillant.</td></tr> +<tr><td align="left"><i>Vent.</i></td><td align="center"> </td><td align="left">Ventenat.</td></tr> +<tr><td align="left"><i>Vill.</i></td><td align="center"> </td><td align="left">Villars.</td></tr> +<tr><td align="left"><i>Wahl.</i></td><td align="center"> </td><td align="left">Wahlenberg.</td></tr> +<tr><td align="left"><i>Walds.</i></td><td align="center"> </td><td align="left">Waldstein.</td></tr> +<tr><td align="left"><i>Wall.</i></td><td align="center"> </td><td align="left">Wallich.</td></tr> +<tr><td align="left"><i>Wallr.</i></td><td align="center"> </td><td align="left">Wallroth.</td></tr> +<tr><td align="left"><i>Walp.</i></td><td align="center"> </td><td align="left">Walpers.</td></tr> +<tr><td align="left"><i>Walt.</i></td><td align="center"> </td><td align="left">Walter.</td></tr> +<tr><td align="left"><i>Wang.</i></td><td align="center"> </td><td align="left">Wangenheim.</td></tr> +<tr><td align="left"><i>Wats.</i></td><td align="center"> </td><td align="left">Sereno Watson, unless other initials are given.</td></tr> +<tr><td align="left"><i>Wedd.</i></td><td align="center"> </td><td align="left">Weddell.</td></tr> +<tr><td align="left"><i>Wendl.</i></td><td align="center"> </td><td align="left">Wendland.</td></tr> +<tr><td align="left"><i>Wiks.</i></td><td align="center"> </td><td align="left">Wikstrom.</td></tr> +<tr><td align="left"><i>Willd.</i></td><td align="center"> </td><td align="left">Willdenow.</td></tr> +<tr><td align="left"><i>Wulf.</i></td><td align="center"> </td><td align="left">Wulfen.</td></tr> +<tr><td align="left"><i>Zucc.</i></td><td align="center"> </td><td align="left">Zuccarini.</td></tr> +<tr><td align="left"><i>Zuccag.</i></td><td align="center"> </td><td align="left">Zuccagini.</td></tr> +</table></div> + + + +<hr class="chap" /><p><span class="pagenum"><a name="Page_193" id="Page_193">[Pg 193]</a></span></p> +<h2><a name="INDEX" id="INDEX"></a>GLOSSARY AND INDEX,</h2> + +<div class="likeheading3">OR<br /> + +DICTIONARY OF THE PRINCIPAL TERMS IN DESCRIPTIVE BOTANY, COMBINED WITH +AN INDEX.</div> + + +<blockquote><p>For the convenience of unclassical students, the commoner Latin +and Greek words (or their equivalents in English form) which +enter into the composition of botanical names, as well as of +technical terms, are added to this Glossary. The numbers refer +to pages. </p></blockquote> + +<div class="index"> +<p><i>A</i>, at the beginning of words of Greek derivation, commonly signifies a +negative, or the absence of something; as <i>a</i>petalous, without petals; +<i>a</i>phyllous, leafless, &c. In words beginning with a vowel, the prefix +is <i>an</i>; as <i>an</i>antherous, destitute of anther.</p> + +<p><i>Abnormal</i>, contrary to the usual or the natural structure.</p> + +<p><i>Aboriginal</i>, original in the strictest sense; same as indigenous.</p> + +<p><i>Abortive</i>, imperfectly formed, or rudimentary.</p> + +<p><i>Abortion</i>, the imperfect formation or the non-formation of some part.</p> + +<p><i>Abrupt</i>, suddenly terminating; as, for instance,</p> + +<p><i>Abruptly pinnate</i>, pinnate without an odd leaflet at the end, <a href="#Page_58">58</a>.</p> + +<p><i>Acantho-</i>, spiny.</p> + +<p><i>Acaulescent</i> (<i>acaulis</i>), apparently stemless; the proper stem, bearing +the leaves and flowers, being very short or subterranean.</p> + +<p><i>Accessory</i>, something additional; as <i>Accessory buds</i>, <a href="#Page_30">30</a>, <a href="#Page_31">31</a>; +<i>Accessory fruits</i>, <a href="#Page_118">118</a>.</p> + +<p><i>Accrescent</i>, growing larger after flowering.</p> + +<p><i>Accrete</i>, grown to.</p> + +<p><i>Accumbent</i>, lying against a thing. The cotyledons are accumbent when +they lie with their edges against the radicle, <a href="#Page_128">128</a>.</p> + +<p><i>Acephalous</i>, headless.</p> + +<p><i>Acerose</i>, needle-shaped, as the leaves of Pines.</p> + +<p><i>Acetabuliform</i>, saucer-shaped.</p> + +<p><i>Achænium</i>, or <i>Achenium</i> (plural <i>achenia</i>), a one-seeded, seed-like +fruit, <a href="#Page_120">120</a>.</p> + +<p><i>Achlamydeous</i> (flower), without floral envelopes, <a href="#Page_86">86</a>.</p> + +<p><i>Acicular</i>, needle-shaped; more slender than acerose.</p> + +<p><i>Acinaciform</i>, scimitar-shaped, like some bean-pods.</p> + +<p><i>Acines</i>, the separate grains of a fruit, such as the raspberry.</p> + +<p><i>Acorn</i>, the nut of the Oak, <a href="#Page_122">122</a>.</p> + +<p><i>Acotyledonous</i>, destitute of cotyledons or seed-leaves.</p> + +<p><i>Acrogenous</i>, growing from the apex, as the stems of Ferns and Mosses. +<i>Acrogens</i>, or <i>Acrogenous Plants</i>, a name for the vascular cryptogamous +plants, <a href="#Page_156">156</a>.</p> + +<p><i>Aculeate</i>, armed with prickles, i. e. <i>aculei</i>; as the Rose and Brier.</p> + +<p><i>Aculeolate</i>, armed with small prickles, or slightly prickly.</p> + +<p><i>Acuminate</i>, taper-pointed, <a href="#Page_54">54</a>.</p> + +<p><i>Acute</i>, merely sharp-pointed, or ending in a point less than a right +angle, <a href="#Page_54">54</a>.</p> + +<span class="pagenum"><a name="Page_194" id="Page_194">[Pg 194]</a></span><p><i>Adelphous</i> (stamens), joined in a fraternity (<i>adelphia</i>); see +<a href="#Monadelphous"><i>monadelphous</i></a>, &c.</p> + +<p><i>Aden</i>, Greek for gland. So <i>Adenophorous</i>, gland-bearing.</p> + +<p><i>Adherent</i>, sticking to, or more commonly, growing fast to another body.</p> + +<p><i>Adnate</i>, literally, growing fast to, born adherent, <a href="#Page_95">95</a>. The anther is +adnate when fixed by its whole length to the filament or its +prolongation, <a href="#Page_101">101</a>.</p> + +<p><i>Adnation</i>, the state of being adnate, <a href="#Page_94">94</a>.</p> + +<p><i>Adpressed</i> or <i>appressed</i>, brought into contact with, but not united.</p> + +<p><i>Adscendent</i>, <i>ascendent</i>, or <i>ascending</i>, rising gradually upwards, <a href="#Page_39">39</a>.</p> + +<p><i>Adsurgent</i>, or <i>assurgent</i>, same as ascending, <a href="#Page_39">39</a>.</p> + +<p><i>Adventitious</i>, out of the proper or usual place; e. g. <i>Adventitious +buds</i>, <a href="#Page_30">30</a>.</p> + +<p><i>Adventive</i>, applied to foreign plants accidentally or sparingly +introduced into a country, but hardly to be called naturalized.</p> + +<p><i>Æquilateral</i>, equal-sided; opposed to oblique.</p> + +<p><i>Aerial roots</i>, &c., <a href="#Page_36">36</a>.</p> + +<p><i>Æruginous</i>, verdigris-colored.</p> + +<p><i>Aestival</i>, produced in summer.</p> + +<p><a name="aestivation" id="aestivation"></a><i>Aestivation</i>, the arrangement of parts in a flower-bud, <a href="#Page_97">97</a>.</p> + +<p><i>Agamous</i>, sexless.</p> + +<p><i>Aggregate fruits</i>, <a href="#Page_118">118</a>.</p> + +<p><i>Agrestis</i>, growing in fields.</p> + +<p><i>Air-cells</i> or <i>Air-passages</i>, spaces in the tissue of leaves and some +stems, <a href="#Page_131">131</a>.</p> + +<p><i>Air-Plants</i>, <a href="#Page_36">36</a>.</p> + +<p><i>Akene</i> or <i>Akenium</i>, <a href="#Page_120">120</a>.</p> + +<p><i>Ala</i> (plural, <i>alæ</i>), a wing; the side-petals of a papilionaceous +corolla, <a href="#Page_92">92</a>.</p> + +<p><i>Alabastrum</i>, a flower-bud.</p> + +<p><i>Alar</i>, situated in the forks of a stem.</p> + +<p><i>Alate</i>, winged.</p> + +<p><i>Albescent</i>, whitish, or turning white.</p> + +<p><i>Albus</i>, Latin for white.</p> + +<p><i>Albumen</i> of the seed, nourishing matter stored up with the embryo, <a href="#Page_21">21</a>, +<a href="#Page_127">127</a>.</p> + +<p><i>Albumen</i>, a vegetable product, of four elements.</p> + +<p><i>Albuminous</i> (seeds), furnished with albumen, <a href="#Page_21">21</a>.</p> + +<p><i>Alburnum</i>, young wood, sap-wood, <a href="#Page_142">142</a>.</p> + +<p><i>Alliaceous</i>, with odor of garlic.</p> + +<p><i>Allogamous</i>, close fertilization.</p> + +<p><i>Alpestrine</i>, subalpine.</p> + +<p><i>Alpine</i>, belonging to high mountains above the limit of forests.</p> + +<p><i>Alternate</i> (leaves), one after another, <a href="#Page_29">29</a>, <a href="#Page_67">67</a>. Petals are <i>alternate</i> +with the sepals, or stamens with the petals, when they stand over the +intervals between them, <a href="#Page_82">82</a>.</p> + +<p><i>Alveolate</i>, honeycomb-like.</p> + +<p><a name="Ament" id="Ament"></a><i>Ament</i>, the scaly spike of trees like the Birch and Willow, <a href="#Page_75">75</a>.</p> + +<p><i>Amentaceous</i>, catkin-like, or catkin-bearing.</p> + +<p><i>Amorphous</i>, shapeless, without any definite form.</p> + +<p><i>Amphicarpous</i>, producing two kinds of fruit.</p> + +<p><i>Amphigastrium</i> (plural, <i>amphigastria</i>), a peculiar stipule-like leaf +of Liverworts.</p> + +<p><i>Amphitropous</i> ovules or seeds, <a href="#Page_111">111</a>.</p> + +<p><i>Amphora</i>, a pitcher-shaped organ.</p> + +<p><i>Amplectant</i>, embracing. <i>Amplexicaul</i> (leaves), clasping the stem by +the base.</p> + +<p><i>Ampullaceous</i>, swelling out like a bottle or bladder (<i>ampulla</i>).</p> + +<p><i>Amylaceous</i>, <i>Amyloid</i>, composed of starch (<i>amylum</i>), or starch-like.</p> + +<p><i>Anandrous</i>, without stamens.</p> + +<p><i>Anantherous</i>, without anthers.</p> + +<p><i>Ananthous</i>, destitute of flowers; flowerless.</p> + +<p><i>Anastomosing</i>, forming a net-work (<i>anastomosis</i>), as the veins of +leaves, <a href="#Page_50">50</a>.</p> + +<p><i>Anatropous</i> ovules or seeds, <a href="#Page_111">111</a>.</p> + +<p><i>Ancipital</i> (<i>anceps</i>), two-edged.</p> + +<p><i>Andrœcium</i>, a name for the stamens taken together, <a href="#Page_98">98</a>.</p> + +<span class="pagenum"><a name="Page_195" id="Page_195">[Pg 195]</a></span><p><i>Andro-diœcious</i>, flowers staminate on one plant, perfect on +another.</p> + +<p><i>Androgynous</i>, having both staminate and pistillate flowers in the same +cluster.</p> + +<p><i>Androphore</i>, a column of united stamens, as in a Mallow.</p> + +<p><i>Androus</i>, or <i>Ander</i>, <i>andra</i>, <i>andrum</i>, Greek in compounds for male, +or stamens.</p> + +<p><i>Anemophilous</i>, wind-loving, said of wind-fertilizable flowers, <a href="#Page_113">113</a>.</p> + +<p><i>Anfractuose</i>, bent hither and thither as the anthers of the Squash, &c.</p> + +<p><i>Angiospermæ</i>, <i>Angiospermous</i>, with seeds formed in an ovary or +pericarp, <a href="#Page_109">109</a>.</p> + +<p><i>Angular divergence</i> of leaves, <a href="#Page_69">69</a>.</p> + +<p><i>Anisos</i>, unequal. <i>Anisomerous</i>, parts unequal in number. +<i>Anisopetalous</i>, with unequal petals. <i>Anisophyllous</i>, the leaves +unequal in the pairs.</p> + +<p><i>Annual</i> (plant), flowering and fruiting the year it is raised from the +seed, and then dying, <a href="#Page_37">37</a>.</p> + +<p><i>Annular</i>, in the form of a ring, or forming a circle.</p> + +<p><i>Annulate</i>, marked by rings; or furnished with an</p> + +<p><i>Annulus</i>, or ring, like that of the spore-case of most Ferns. In Mosses +it is a ring of cells placed between the mouth of the spore-case and the +lid in many species.</p> + +<p><i>Annotinous</i>, yearly, or in yearly growths.</p> + +<p><i>Anterior</i>, in the blossom, is the part next the bract, i. e. external; +while the posterior side is that next the axis of inflorescence. Thus, +in the Pea, &c., the keel is <i>anterior</i>, and the standard <i>posterior</i>, +<a href="#Page_96">96</a>.</p> + +<p><i>Anthela</i>, an open paniculate cyme.</p> + +<p><i>Anther</i>, the essential part of the stamen, which contains the pollen, +<a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_101">101</a>.</p> + +<p><i>Antheridium</i> (plural <i>antheridia</i>), the organ in Cryptogams which +answers to the anther of Flowering Plants, <a href="#Page_150">150</a>.</p> + +<p><i>Antheriferous</i>, anther-bearing.</p> + +<p><i>Anthesis</i>, the period or the act of the expansion of a flower.</p> + +<p><i>Anthocarpus</i> (fruits), <a href="#Page_118">118</a>.</p> + +<p><i>Anthophore</i>, a stipe between calyx and corolla, <a href="#Page_113">113</a>.</p> + +<p><i>Anthos</i>, Greek for flower; in composition, <i>Monanthous</i>, one-flowered, +&c.</p> + +<p><i>Anticous</i>, same as anterior.</p> + +<p><i>Antrorse</i>, directed upwards or forwards.</p> + +<p><i>Apetalous</i>, destitute of petals, <a href="#Page_86">86</a>.</p> + +<p><i>Aphyllous</i>, leafless.</p> + +<p><i>Apical</i>, belonging to the apex or point.</p> + +<p><i>Apiculate</i>, pointleted; tipped with a small point.</p> + +<p><i>Apocarpous</i> (pistils), when the several pistils of the same flower are +separate.</p> + +<p><i>Apophysis</i>, any irregular swelling; the enlargement at the base of the +spore-case of the Umbrella-Moss.</p> + +<p><i>Apothecium</i>, the fructification of Lichens, <a href="#Page_171">171</a>.</p> + +<p><i>Appendage</i>, any superadded part. <i>Appendiculate</i>, provided with +appendages.</p> + +<p><i>Appressed</i>, close pressed to the stem, &c.</p> + +<p><i>Apricus</i>, growing in dry and sunny places.</p> + +<p><i>Apterous</i>, wingless.</p> + +<p><i>Aquatic</i> (<i>Aquatilis</i>), living or growing in water; applied to plants +whether growing under water, or with all but the base raised out of it.</p> + +<p><i>Arachnoid</i>, <i>Araneose</i>, cobwebby; clothed with, or consisting of, soft +downy fibres.</p> + +<p><i>Arboreous</i>, <i>Arborescent</i>, tree-like, in size or form, <a href="#Page_39">39</a>.</p> + +<p><i>Arboretum</i>, a collection of trees.</p> + +<p><i>Archegonium</i> (plural <i>archegonia</i>), the organ in Mosses, &c., which is +analogous to the pistil of Flowering Plants.</p> + +<p><i>Arcuate</i>, bent or curved like a bow.</p> + +<p><i>Arenose</i> (<i>Arenarius</i>), growing in sand.</p> + +<p><i>Areolate</i>, marked out into little spaces or <i>areolæ</i>.</p> + +<p><i>Argenteous</i>, or <i>Argentate</i>, silvery-like.</p> + +<p><i>Argillose</i>, growing in clay.</p> + +<p><i>Argos</i>, Greek for pure white; <i>Argophyllous</i> or <i>Argyrophyllous</i>, +white-leaved, &c.</p> + +<p><i>Argutus</i>, acutely dentate.</p> + +<span class="pagenum"><a name="Page_196" id="Page_196">[Pg 196]</a></span><p><i>Arillate</i> (seeds) furnished with an aril.</p> + +<p><i>Arilliform</i>, aril-like.</p> + +<p><i>Arillus</i>, or <i>Aril</i>, a fleshy growth from base of a seed, <a href="#Page_126">126</a>.</p> + +<p><i>Aristate</i>, awned, i. e. furnished with an <i>arista</i>, like the beard of +Barley, &c., <a href="#Page_54">54</a>.</p> + +<p><i>Aristulate</i>, diminutive of the last; short-awned.</p> + +<p><i>Arrect</i>, brought into upright position.</p> + +<p><i>Arrow-shaped</i> or <i>Arrow-headed</i>, same as <i>sagittate</i>, <a href="#Page_53">53</a>.</p> + +<p><i>Articulated</i>, jointed; furnished with joints or <i>articulations</i>, where +it separates or inclines to do so. <i>Articulated leaves</i>, <a href="#Page_57">57</a>.</p> + +<p><i>Artificial Classification</i>, <a href="#Page_181">181</a>.</p> + +<p><i>Ascending</i> (stems, &c.), <a href="#Page_39">39</a>; (seeds or ovules), <a href="#Page_110">110</a>.</p> + +<p><i>Ascidium</i>, a pitcher-shaped body, like leaves of Sarracenia.</p> + +<p><i>Ascus</i> (<i>asci</i>), a sac, the spore-case of Lichens and some Fungi.</p> + +<p><a name="Aspergilliform" id="Aspergilliform"></a><i>Aspergilliform</i>, shaped like the brush used to sprinkle holy water; as +the stigmas of many Grasses.</p> + +<p><i>Asperous</i>, rough to touch.</p> + +<p><i>Assimilation</i>, <a href="#Page_144">144</a>, <a href="#Page_147">147</a>.</p> + +<p><i>Assurgent</i>, same as ascending, <a href="#Page_39">39</a>.</p> + +<p><i>Atropous</i> or <i>Atropal</i> (ovules), same as orthotropous.</p> + +<p><i>Aurantiacous</i>, orange-colored.</p> + +<p><i>Aureous</i>, golden.</p> + +<p><a name="Auriculate" id="Auriculate"></a><i>Auriculate</i>, furnished with <i>auricles</i> or ear-like appendages, <a href="#Page_53">53</a>.</p> + +<p><i>Autogamy</i>, self-fertilization, <a href="#Page_115">115</a>.</p> + +<p><i>Awl-shaped</i>, sharp-pointed from a broader base, <a href="#Page_61">61</a>.</p> + +<p><i>Awn</i>, the bristle or beard of Barley, Oats, &c.; or any similar +appendage.</p> + +<p><i>Awned</i> or <i>Awn-pointed</i>, furnished with an awn or long bristle-shaped +tip, <a href="#Page_54">54</a>.</p> + +<p><i>Axil</i>, the angle on the upper side between a leaf and the stem, <a href="#Page_13">13</a>.</p> + +<p><i>Axile</i>, belonging to the axis, or occupying the axis.</p> + +<p><i>Axillary</i> (buds, &c.), occurring in an axil, <a href="#Page_27">27</a>.</p> + +<p><i>Axis</i>, the central line of any body; the organ round which others are +attached; the root and stem. <i>Ascending</i> and <i>Descending Axis</i>, <a href="#Page_38">38</a>.</p> +</div> + +<div class="index"> +<p><i>Baccate</i>, berried, berry-like, of a pulpy nature like a berry +(<i>bacca</i>).</p> + +<p><i>Badius</i>, chestnut-colored.</p> + +<p><i>Banner</i>, see <a href="#Standard">Standard</a>, <a href="#Page_92">92</a>.</p> + +<p><a name="Barbate" id="Barbate"></a><i>Barbate</i>, bearded; bearing tufts, spots, or lines of hairs.</p> + +<p><i>Barbed</i>, furnished with a <i>barb</i> or double hook; as the apex of the +bristle on the fruit of Echinospermum (Stickseed), &c.</p> + +<p><i>Barbellate</i>, said of the bristles of the pappus of some Compositæ when +beset with short, stiff hairs, longer than when denticulate, but shorter +than when plumose.</p> + +<p><i>Barbellulate</i>, diminutive of barbellate.</p> + +<p><i>Bark</i>, the covering of a stem outside of the wood, <a href="#Page_138">138</a>, <a href="#Page_140">140</a>.</p> + +<p><i>Basal</i>, belonging or attached to the</p> + +<p><i>Base</i>, that extremity of any organ by which it is attached to its +support.</p> + +<p><i>Basifixed</i>, attached by its base.</p> + +<p><i>Bast</i>, <i>Bast-fibres</i>, <a href="#Page_134">134</a>.</p> + +<p><i>Beaked</i>, ending in a prolonged narrow tip.</p> + +<p><i>Bearded</i>, see <a href="#Barbate"><i>barbate</i></a>. <i>Beard</i> is sometimes used for awn, more +commonly for long or stiff hairs of any sort.</p> + +<p><i>Bell-shaped</i>, of the shape of a bell, as the corolla of Harebell, <a href="#Page_90">90</a>.</p> + +<p><i>Berry</i>, a fruit pulpy or juicy throughout, as a grape, <a href="#Page_119">119</a>.</p> + +<p><i>Bi-</i> (or <i>Bis</i>), in compound words, twice; as</p> + +<p><i>Biarticulate</i>, twice-jointed, or two-jointed; separating into two +pieces.</p> + +<p><i>Biauriculate</i>, having two ears, as the leaf in fig. <a href="#Fig125">126</a>.</p> + +<p><i>Bicallose</i>, having two callosities or harder spots.</p> + +<p><i>Bicarinate</i>, two-keeled.</p> + +<p><i>Bicipital</i> (<i>Biceps</i>), two-headed; dividing into two parts.</p> + +<span class="pagenum"><a name="Page_197" id="Page_197">[Pg 197]</a></span><p><i>Biconjugate</i>, twice paired, as when a petiole forks twice.</p> + +<p><i>Bidentate</i>, having two teeth (not twice or doubly dentate).</p> + +<p><i>Biennial</i>, of two years' continuance; springing from the seed one +season, flowering and dying the next, <a href="#Page_38">38</a>.</p> + +<p><i>Bifarious</i>, two-ranked; arranged in two rows.</p> + +<p><i>Bifid</i>, two-cleft to about the middle.</p> + +<p><i>Bifoliolate</i>, a compound leaf of two leaflets, <a href="#Page_59">59</a>.</p> + +<p><i>Bifurcate</i>, twice forked; or more commonly, forked into two branches.</p> + +<p><i>Bijugate</i>, bearing two pairs (of leaflets, &c.).</p> + +<p><i>Bilabiate</i>, two-lipped, as the corolla of Labiatæ.</p> + +<p><i>Bilamellate</i>, of two plates (<i>lamellæ</i>), as the stigma of Mimulus.</p> + +<p><i>Bilobed</i>, the same as two-lobed.</p> + +<p><i>Bilocellate</i>, when a cell is divided into two <i>locelli</i>.</p> + +<p><i>Bilocular</i>, two-celled; as most anthers, the pod of Foxglove, &c.</p> + +<p><i>Binary</i>, in twos.</p> + +<p><i>Binate</i>, in couples, two together. <i>Bipartite</i>, the Latin form of +two-parted.</p> + +<p><i>Binodal</i>, of two nodes.</p> + +<p><i>Binomial</i>, of two words, as the name of genus and species taken +together, <a href="#Page_180">180</a>.</p> + +<p><i>Bipalmate</i>, twice palmately divided.</p> + +<p><i>Biparous</i>, bearing two.</p> + +<p><i>Bipinnate</i> (leaf), twice pinnate, <a href="#Page_58">58</a>. <i>Bipinnatifid</i>, twice pinnatifid, +<a href="#Page_57">57</a>.</p> + +<p><i>Bipinnatisect</i>, twice pinnately divided.</p> + +<p><i>Biplicate</i>, twice folded together.</p> + +<p><i>Biserial</i>, or <i>Biseriate</i>, occupying two rows, one within the other.</p> + +<p><i>Biserrate</i>, doubly serrate, as when the teeth of a leaf are themselves +serrate.</p> + +<p><i>Bisexual</i>, having both stamens and pistil.</p> + +<p><i>Biternate</i>, twice ternate; i. e. principal divisions three, each +bearing three leaflets, <a href="#Page_59">59</a>.</p> + +<p><i>Bladdery</i>, thin and inflated.</p> + +<p><i>Blade</i> of a leaf, its expanded portion, <a href="#Page_49">49</a>.</p> + +<p><i>Bloom</i>, the whitish powder on some fruits, leaves, &c.</p> + +<p><i>Boat-shaped</i>, concave within and keeled without, in shape like a small +boat.</p> + +<p><i>Border</i> of corolla, &c., <a href="#Page_89">89</a>.</p> + +<p><i>Brachiate</i>, with opposite branches at right angles to each other.</p> + +<p><i>Brachy-</i>, short, as <i>Brachycarpous</i>, short-fruited, &c.</p> + +<p><i>Bract</i> (<i>Bractea</i>), the leaf of an inflorescence. Specially, the bract +is the small leaf or scale from the axil of which a flower or its +pedicel proceeds, <a href="#Page_73">73</a>.</p> + +<p><i>Bracteate</i>, furnished with bracts.</p> + +<p><i>Bracteolate</i>, furnished with bractlets.</p> + +<p><i>Bracteose</i>, with numerous or conspicuous bracts.</p> + +<p><i>Bractlet</i> (<i>Bracteola</i>), or <i>Bracteole</i>, is a bract seated <i>on</i> the +pedicel or flower-stalk, <a href="#Page_73">73</a>.</p> + +<p><i>Branch</i>, <i>Branching</i>, <a href="#Page_27">27</a>.</p> + +<p><i>Breathing-pores</i>, <a href="#Page_144">144</a>.</p> + +<p><i>Bristles</i>, stiff, sharp hairs, or any very slender bodies of similar +appearance.</p> + +<p><i>Bristly</i>, beset with bristles. <i>Bristle-pointed</i>, <a href="#Page_54">54</a>.</p> + +<p><i>Brunneous</i>, brown.</p> + +<p><i>Brush-shaped</i>, see <a href="#Aspergilliform"><i>aspergilliform</i></a>.</p> + +<p><i>Bryology</i>, that part of botany which relates to Mosses.</p> + +<p><i>Bryophyta</i>, <i>Bryophytes</i>, <a href="#Page_163">163</a>.</p> + +<p><i>Bud</i>, a branch in its earliest or undeveloped state, <a href="#Page_27">27</a>. <i>Bud-scales</i>, +<a href="#Page_63">63</a>.</p> + +<p><i>Bulb</i>, a leaf-bud with fleshy scales, usually subterranean, <a href="#Page_46">46</a>.</p> + +<p><i>Bulbils</i>, diminutive bulbs.</p> + +<p><i>Bulbiferous</i>, bearing or producing bulbs. <i>Bulbose</i> or <i>bulbous</i>, +bulb-like in shape, &c.</p> + +<p><i>Bulblets</i>, small bulbs, borne above ground, <a href="#Page_46">46</a>.</p> + +<p><i>Bulb-scales</i>, <a href="#Page_46">46</a>.</p> + +<p><i>Bullate</i>, appearing as if blistered or bladdery (from <i>bulla</i>, a +bubble).</p> + +<p><i>Byssaceous</i>, composed of fine flax-like threads.</p> +</div> + +<div class="index"> +<span class="pagenum"><a name="Page_198" id="Page_198">[Pg 198]</a></span><p><i>Caducous</i>, dropping off very early, compared with other parts; as the +calyx in the Poppy, falling when the flower opens.</p> + +<p><i>Cæruleous</i>, blue. <i>Cærulescent</i>, becoming bluish.</p> + +<p><i>Cæspitose</i>, or <i>Cespitose</i>, growing in turf-like patches or tufts.</p> + +<p><i>Calathiform</i>, cup-shaped.</p> + +<p><i>Calcarate</i>, furnished with a spur (<i>calcar</i>), <a href="#Page_86">86</a>, <a href="#Page_87">87</a>.</p> + +<p><i>Calceolate</i> or <i>Calceiform</i>, slipper-shaped, like one petal of the +Lady's Slipper.</p> + +<p><i>Callose</i>, hardened; or furnished with callosities or thickened spots.</p> + +<p><i>Calvous</i>, bald or naked of hairs.</p> + +<p><i>Calyciflorus</i>, when petals and stamens are adnate to calyx.</p> + +<p><i>Calycine</i>, belonging to the calyx.</p> + +<p><i>Calyculate</i>, furnished with an outer accessory calyx (<i>calyculus</i>) or +set of bracts looking like a calyx, as in true Pinks.</p> + +<p><i>Calyptra</i>, the hood or veil of the capsule of a Moss, <a href="#Page_163">163</a>.</p> + +<p><i>Calyptrate</i>, having a calyptra.</p> + +<p><i>Calyptriform</i>, shaped like a calyptra or candle-extinguisher.</p> + +<p><i>Calyx</i>, the outer set of the floral envelopes or leaves of the flower, +<a href="#Page_14">14</a>, <a href="#Page_79">79</a>.</p> + +<p><i>Cambium</i>, <i>Cambium-layer</i>, <a href="#Page_140">140</a>.</p> + +<p><i>Campanulate</i>, bell-shaped, <a href="#Page_90">90</a>.</p> + +<p><i>Campylotropous</i>, or <i>Campylotropal</i>, curved ovules and seeds, <a href="#Page_111">111</a>. +<i>Campylospermous</i>, applied to fruits of Umbelliferæ when the seed is +curved in at the edges, forming a groove down the inner face; as in +Sweet Cicely.</p> + +<p><i>Canaliculate</i>, channelled, or with a deep longitudinal groove.</p> + +<p><i>Cancellate</i>, latticed, resembling lattice-work.</p> + +<p><i>Candidus</i>, Latin for pure white.</p> + +<p><a name="Canescent" id="Canescent"></a><i>Canescent</i>, grayish-white; hoary, usually because the surface is +covered with fine white hairs. <i>Incanous</i> is whiter still.</p> + +<p><i>Canous</i>, whitened with pubescence; see <a href="#Incanous"><i>incanous</i></a>.</p> + +<p><i>Capillaceous</i>, <i>Capillary</i>, hair-like in shape; as fine as hair or +slender bristles.</p> + +<p><i>Capitate</i>, having a globular apex, like the head on a pin.</p> + +<p><i>Capitellate</i>, diminutive of capitate.</p> + +<p><i>Capitulum</i>, a close rounded dense cluster or <i>head</i> of sessile flowers, +<a href="#Page_74">74</a>.</p> + +<p><i>Capreolate</i>, bearing tendrils (from <i>capreolus</i>, a tendril).</p> + +<p><i>Capsule</i>, a dry dehiscent seed-vessel of a compound pistil, <a href="#Page_122">122</a>.</p> + +<p><i>Capsular</i>, relating to, or like a capsule.</p> + +<p><i>Capture of insects</i>, <a href="#Page_154">154</a>.</p> + +<p><i>Carina</i>, a keel; the two anterior petals of a papilionaceous flower, +<a href="#Page_92">92</a>.</p> + +<p><i>Carinate</i>, keeled, furnished with a sharp ridge or projection on the +lower side.</p> + +<p><a name="Caryopsis" id="Caryopsis"></a><i>Cariopsis</i>, or <i>Caryopsis</i>, the one-seeded fruit or grain of Grasses, +<a href="#Page_121">121</a>.</p> + +<p><i>Carneous</i>, flesh-colored; pale red. <i>Carnose</i>, fleshy in texture.</p> + +<p><i>Carpel</i>, or <i>Carpidium</i>, a simple pistil or a pistil-leaf, <a href="#Page_106">106</a>.</p> + +<p><i>Carpellary</i>, pertaining to a carpel.</p> + +<p><i>Carpology</i>, that department of botany which relates to fruits.</p> + +<p><i>Carpophore</i>, the stalk or support of a pistil extending between its +carpels, <a href="#Page_113">113</a>.</p> + +<p><i>Carpos</i>, Greek for fruit.</p> + +<p><i>Cartilaginous</i>, or <i>Cartilagineous</i>, firm and tough in texture, like +cartilage.</p> + +<p><i>Caruncle</i>, an excrescence at the scar of some seeds, <a href="#Page_126">126</a>.</p> + +<p><i>Carunculate</i>, furnished with a caruncle.</p> + +<p><i>Caryophyllaceous</i>, pink-like: applied to a corolla of 5 long-clawed +petals.</p> + +<p><i>Cassideous</i>, helmet-shaped.</p> + +<p><i>Cassus</i>, empty and sterile.</p> + +<p><i>Catenate</i>, or <i>Catenulate</i>, end to end as in a chain.</p> + +<p><i>Catkin</i>, see <a href="#Ament">Ament</a>, <a href="#Page_75">75</a>.</p> + +<p><i>Caudate</i>, tailed, or tail-pointed.</p> + +<p><i>Caudex</i>, a sort of trunk, such as that of Palms; an upright rootstock, +<a href="#Page_39">39</a>, <a href="#Page_44">44</a>.</p> + +<p><i>Caudicle</i>, the stalk of a pollen-mass, &c.</p> + +<p><i>Caulescent</i>, having an obvious stem, <a href="#Page_36">36</a>.</p> + +<span class="pagenum"><a name="Page_199" id="Page_199">[Pg 199]</a></span><p><i>Caulicle</i>, a little stem, or rudimentary stem (of a seedling), <a href="#Page_11">11</a>, +<a href="#Page_127">127</a>.</p> + +<p><i>Cauline</i>, of or belonging to a stem, <a href="#Page_36">36</a>. <i>Caulis</i>, Latin name of stem.</p> + +<p><i>Caulocarpic</i>, equivalent to perennial.</p> + +<p><i>Caulome</i>, the cauline parts of a plant.</p> + +<p><i>Cell</i> (diminutive, <i>Cellule</i>), the cavity of an anther, ovary, &c.; one +of the anatomical elements, <a href="#Page_131">131</a>.</p> + +<p><i>Cellular Cryptogams</i>, <a href="#Page_162">162</a>.</p> + +<p><i>Cellular tissue</i>, <a href="#Page_131">131</a>.</p> + +<p><i>Cellulose</i>, <a href="#Page_131">131</a>.</p> + +<p><i>Cell-walls</i>, <a href="#Page_130">130</a>.</p> + +<p><i>Centrifugal</i> (inflorescence), produced or expanding in succession from +the centre outwards, <a href="#Page_77">77</a>.</p> + +<p><i>Centripetal</i>, the opposite of centrifugal, <a href="#Page_74">74</a>.</p> + +<p><i>Cephala</i>, Greek for head. In compounds, <i>Monocephalous</i>, with one head, +<i>Microcephalous</i>, small-headed, &c.</p> + +<p><i>Cereal</i>, belonging to corn, or corn-plants.</p> + +<p><i>Cernuous</i>, nodding; the summit more or less inclining.</p> + +<p><i>Chæta</i>, Greek for bristle.</p> + +<p><i>Chaff</i>, small membranous scales or bracts on the receptacle of +Compositæ; the glumes, &c., of grasses.</p> + +<p><i>Chaffy</i>, furnished with chaff, or of the texture of chaff.</p> + +<p><i>Chalaza</i>, that part of the ovule where all the parts grow together, +<a href="#Page_110">110</a>, <a href="#Page_126">126</a>.</p> + +<p><i>Channelled</i>, hollowed out like a gutter; same as <i>canaliculate</i>.</p> + +<p><i>Character</i>, a phrase expressing the essential marks of a species, +genus, &c., <a href="#Page_181">181</a>.</p> + +<p><i>Chartaceous</i>, of the texture of paper or parchment.</p> + +<p><i>Chloros</i>, Greek for green, whence <i>Chloranthous</i>, green-flowered; +<i>Chlorocarpous</i>, green-fruited, &c.</p> + +<p><i>Chlorophyll</i>, leaf green, <a href="#Page_136">136</a>.</p> + +<p><i>Chlorosis</i>, a condition in which naturally colored parts turn green.</p> + +<p><i>Choripetalous</i>, same as polypetalous.</p> + +<p><i>Chorisis</i>, separation of the normally united parts, or where two or +more parts take the place of one.</p> + +<p><i>Chromule</i>, coloring matter in plants, especially when not green, or +when liquid.</p> + +<p><i>Chrysos</i>, Greek for golden yellow, whence <i>Chrysanthous</i>, +yellow-flowered, &c.</p> + +<p><i>Cicatrix</i>, the scar left by the fall of a leaf or other organ.</p> + +<p><i>Ciliate</i>, beset on the margin with a fringe of <i>cilia</i>, i. e. of hairs +or bristles, like the eyelashes fringing the eyelids, whence the name.</p> + +<p><i>Cinereous</i>, or <i>Cineraceous</i>, ash-grayish; of the color of ashes.</p> + +<p><i>Circinate</i>, rolled inwards from the top, <a href="#Page_72">72</a>.</p> + +<p><i>Circumscissile</i>, or <i>Circumcissile</i>, divided by a circular line round +the sides, as the pods of Purslane, Plantain, &c., <a href="#Page_124">124</a>.</p> + +<p><i>Circumscription</i>, general outline.</p> + +<p><i>Cirrhiferous</i>, or <i>Cirrhose</i>, furnished with a tendril (Latin, +<i>Cirrhus</i>); as the Grape-vine. <i>Cirrhose</i> also means resembling or +coiling like tendrils, as the leaf-stalks of Virgin's-bower. More +properly <i>Cirrus</i> and <i>Cirrose</i>.</p> + +<p><i>Citreous</i>, lemon-yellow.</p> + +<p><i>Clados</i>, Greek for branch. <i>Cladophylla</i>, <a href="#Page_64">64</a>.</p> + +<p><i>Class</i>, <a href="#Page_178">178</a>, <a href="#Page_183">183</a>.</p> + +<p><i>Classification</i>, <a href="#Page_175">175</a>, <a href="#Page_183">183</a>.</p> + +<p><i>Clathrate</i>, latticed; same as <i>cancellate</i>.</p> + +<p><a name="Clavate" id="Clavate"></a><i>Clavate</i>, club-shaped; slender below and thickened upwards.</p> + +<p><i>Clavellate</i>, diminutive of clavate.</p> + +<p><i>Claviculate</i>, having <i>Claviculæ</i>, or little tendrils or hooks.</p> + +<p><i>Claw</i>, the narrow or stalk-like base of some petals, as of Pinks, <a href="#Page_91">91</a>.</p> + +<p><i>Cleistogamous</i> (<i>Cleistogamy</i>), fertilized in closed bud, <a href="#Page_115">115</a>.</p> + +<p><i>Cleft</i>, cut into lobes, <a href="#Page_55">55</a>.</p> + +<p><i>Close</i> fertilization, <a href="#Page_115">115</a>.</p> + +<p><i>Climbing</i>, rising by clinging to other objects, <a href="#Page_39">39</a>, <a href="#Page_151">151</a>.</p> + +<p><i>Club shaped</i>, see <a href="#Clavate"><i>clavate</i></a>.</p> + +<p><i>Clustered</i>, leaves, flowers, &c., aggregated or collected into a bunch.</p> + +<span class="pagenum"><a name="Page_200" id="Page_200">[Pg 200]</a></span><p><i>Clypeate</i>, buckler-shaped.</p> + +<p><i>Coadunate</i>, same as <i>connate</i>, i. e. united.</p> + +<p><i>Coalescent</i>, growing together. <i>Coalescence</i>, <a href="#Page_88">88</a>.</p> + +<p><i>Coarctate</i>, contracted or brought close together.</p> + +<p><i>Coated</i>, having an integument, or covered in layers. Coated bulb, <a href="#Page_46">46</a>.</p> + +<p><i>Cobwebby</i>, same as <i>arachnoid</i>; bearing hairs like cobwebs or gossamer.</p> + +<p><i>Coccineous</i>, scarlet-red.</p> + +<p><i>Coccus</i> (plural <i>cocci</i>), anciently a berry; now mostly used to denote +the separable carpels or nutlets of a dry fruit.</p> + +<p><i>Cochleariform</i>, spoon-shaped.</p> + +<p><i>Cochleate</i>, coiled or shaped like a snail-shell.</p> + +<p><i>Cœlospermous</i>, applied to those fruits of Umbelliferæ which have the +seed hollowed on the inner face, by incurving of top and bottom; as in +Coriander.</p> + +<p><i>Coherent</i>, usually the same as <i>connate</i>.</p> + +<p><i>Cohort</i>, name sometimes used for groups between order and class, <a href="#Page_178">178</a>.</p> + +<p><i>Coleorhiza</i>, a root-sheath.</p> + +<p><i>Collateral</i>, side by side.</p> + +<p><i>Collective fruits</i>, <a href="#Page_118">118</a>.</p> + +<p><i>Collum</i> or <i>Collar</i>, the neck or junction of stem and root.</p> + +<p><i>Colored</i>, parts of a plant which are other-colored than green.</p> + +<p><i>Columella</i>, the axis to which the carpels of a compound pistil are +often attached, as in Geranium (<a href="#Page_112">112</a>), or which is left when a pod opens, +as in Azalea.</p> + +<p><i>Column</i>, the united stamens, as in Mallow, or the stamens and pistils +united into one body, as in the Orchis family.</p> + +<p><i>Columnar</i>, shaped like a column or pillar.</p> + +<p><i>Coma</i>, a tuft of any sort (literally, a head of hair), <a href="#Page_125">125</a>.</p> + +<p><i>Comose</i>, tufted; bearing a tuft of hairs, as the seeds of Milkweed, +<a href="#Page_126">126</a>.</p> + +<p><i>Commissure</i>, the line of junction of two carpels, as in the fruit of +Umbelliferæ.</p> + +<p><i>Complanate</i>, flattened.</p> + +<p><i>Compound leaf</i>, <a href="#Page_54">54</a>, <a href="#Page_57">57</a>. <i>Compound pistil</i>, <a href="#Page_107">107</a>. <i>Compound umbel</i>, <a href="#Page_75">75</a>, +&c.</p> + +<p><i>Complete</i> (flower), <a href="#Page_81">81</a>.</p> + +<p><i>Complicate</i>, folded upon itself.</p> + +<p><i>Compressed</i>, flattened on opposite sides.</p> + +<p><i>Conceptacle</i>, <a href="#Page_168">168</a>.</p> + +<p><i>Concinnous</i>, neat.</p> + +<p><i>Concolor</i>, all of one color.</p> + +<p><i>Conchiform</i>, shell- or half-shell-shaped.</p> + +<p><i>Conduplicate</i>, folded upon itself lengthwise, <a href="#Page_71">71</a>.</p> + +<p><i>Cone</i>, the fruit of the Pine family, <a href="#Page_124">124</a>. <i>Coniferous</i>, cone-bearing.</p> + +<p><i>Confertus</i>, much crowded.</p> + +<p><i>Conferruminate</i>, stuck together, as the cotyledons in a horse-chestnut.</p> + +<p><i>Confluent</i>, blended together; or the same as <i>coherent</i>.</p> + +<p><i>Conformed</i>, similar to another thing it is associated with or compared +to; or closely fitted to it, as the skin to the kernel of a seed.</p> + +<p><i>Congested</i>, <i>Conglomerate</i>, crowded together.</p> + +<p><i>Conglomerate</i>, crowded into a glomerule.</p> + +<p><i>Conjugate</i>, coupled; in single pairs. <i>Conjugation</i>, <a href="#Page_170">170</a>.</p> + +<p><i>Connate</i>, united or grown together from the first formation, <a href="#Page_96">96</a>.</p> + +<p><i>Connate-perfoliate</i>, when a pair of leaves are connate round a stem, +<a href="#Page_60">60</a>.</p> + +<p><i>Connective</i>, <i>Connectivum</i>, the part of the anther connecting its two +cells, <a href="#Page_101">101</a>.</p> + +<p><i>Connivent</i>, converging, or brought close together.</p> + +<p><i>Consolidation</i> (floral), <a href="#Page_94">94</a>.</p> + +<p><i>Consolidated</i> forms of vegetation, <a href="#Page_47">47</a>.</p> + +<p><i>Contents</i> of cells, <a href="#Page_136">136</a>.</p> + +<p><i>Continuous</i>, the reverse of interrupted or articulated.</p> + +<p><i>Contorted</i>, twisted together. <i>Contorted æstivation</i>, same as +<i>convolute</i>, <a href="#Page_97">97</a>.</p> + +<p><i>Contortuplicate</i>, twisted back upon itself.</p> + +<p><i>Contracted</i>, either narrowed or shortened.</p> + +<span class="pagenum"><a name="Page_201" id="Page_201">[Pg 201]</a></span><p><i>Contrary</i>, turned in opposite direction to the ordinary.</p> + +<p><i>Convolute</i>, rolled up lengthwise, as the leaves of the Plum in +vernation, <a href="#Page_72">72</a>. In æstivation, same as <i>contorted</i>, <a href="#Page_97">97</a>.</p> + +<p><i>Cordate</i>, heart-shaped, <a href="#Page_53">53</a>.</p> + +<p><i>Coriaceous</i>, resembling leather in texture.</p> + +<p><i>Corky</i>, of the texture of cork. <i>Corky layer</i> of bark, <a href="#Page_141">141</a>.</p> + +<p><i>Corm</i>, a solid bulb, like that of Crocus, <a href="#Page_45">45</a>.</p> + +<p><i>Corneous</i>, of the consistence or appearance of horn.</p> + +<p><i>Corniculate</i>, furnished with a small horn or spur.</p> + +<p><i>Cornute</i>, horned; bearing a horn-like projection or appendage.</p> + +<p><i>Corolla</i>, the leaves of the flower within the calyx, <a href="#Page_14">14</a>, <a href="#Page_79">79</a>.</p> + +<p><i>Corollaceous</i>, <i>Corolline</i>, like or belonging to a corolla.</p> + +<p><a name="Corona" id="Corona"></a><i>Corona</i>, a coronet or crown; an appendage at the top of the claw of +some petals, <a href="#Page_91">91</a>.</p> + +<p><a name="Coronate" id="Coronate"></a><i>Coronate</i>, crowned; furnished with a crown.</p> + +<p><i>Cortex</i>, bark. <i>Cortical</i>, belonging to the bark (<i>cortex</i>).</p> + +<p><i>Corticate</i>, coated with bark or bark-like covering.</p> + +<p><i>Corymb</i>, a flat or convex indeterminate flower-cluster, <a href="#Page_74">74</a>.</p> + +<p><i>Corymbiferous</i>, bearing corymbs.</p> + +<p><i>Corymbose</i>, in corymbs, approaching the form of a corymb, or branched +in that way.</p> + +<p><i>Costa</i>, a rib; the midrib of a leaf, &c. <i>Costate</i>, ribbed.</p> + +<p><a name="Cotyledons" id="Cotyledons"></a><i>Cotyledons</i>, the proper leaves of the embryo, <a href="#Page_11">11</a>, <a href="#Page_127">127</a>.</p> + +<p><i>Crateriform</i>, goblet-shaped or deep saucer-shaped.</p> + +<p><i>Creeping</i> (stems), growing flat on or beneath the ground and rooting, +<a href="#Page_39">39</a>.</p> + +<p><i>Cremocarp</i>, a half-fruit, or one of the two carpels of Umbelliferæ, +<a href="#Page_121">121</a>.</p> + +<p><i>Crenate</i>, or <i>Crenelled</i>, the edge scalloped into rounded teeth, <a href="#Page_55">55</a>.</p> + +<p><i>Crenulate</i>, minutely or slightly crenate.</p> + +<p><i>Crested</i>, or <i>Cristate</i>, bearing any elevated appendage like a crest.</p> + +<p><i>Cretaceous</i>, chalky or chalk-like.</p> + +<p><i>Cribrose</i>, or <i>cribriform</i>, pierced like a sieve with small apertures.</p> + +<p><i>Crinite</i>, bearing long hairs.</p> + +<p><i>Crispate</i>, curled or crispy.</p> + +<p><i>Croceous</i>, saffron-color, deep reddish-yellow.</p> + +<p><i>Cross-breeds</i>, the progeny of interbred varieties, <a href="#Page_176">176</a>.</p> + +<p><i>Cross fertilization</i>, <a href="#Page_115">115</a>.</p> + +<p><i>Crown</i>, see <a href="#Corona"><i>corona</i></a>. <i>Crowned</i>, see <a href="#Coronate"><i>coronate</i></a>.</p> + +<p><i>Cruciate</i>, or <i>Cruciform</i>, cross-shaped. <i>Cruciform Corolla</i>, <a href="#Page_86">86</a>.</p> + +<p><i>Crustaceous</i>, hard and brittle in texture; crust-like.</p> + +<p><i>Cryptogamous Plants</i>, <i>Cryptogams</i>, <a href="#Page_10">10</a>, <a href="#Page_156">156</a>.</p> + +<p><i>Cryptos</i>, concealed, as <i>Cryptopetalous</i>, with concealed petals, &c.</p> + +<p><i>Crystals</i> in plants, <a href="#Page_137">137</a>.</p> + +<p><a name="Cucullate" id="Cucullate"></a><i>Cucullate</i>, hooded, or hood-shaped, rolled up like a cornet of paper, +or a hood (<i>cucullus</i>), as the spathe of Indian Turnip, <a href="#Page_75">75</a>.</p> + +<p><i>Culm</i>, a straw; the stem of Grasses and Sedges, <a href="#Page_39">39</a>.</p> + +<p><i>Cultrate</i>, shaped like a trowel or broad knife.</p> + +<p><i>Cuneate</i>, <i>Cuneiform</i>, wedge-shaped, <a href="#Page_53">53</a>.</p> + +<p><i>Cup-shaped</i>, same as cyathiform or near it.</p> + +<p><i>Cupule</i>, a little cup; the cup to the acorn of the Oak, <a href="#Page_122">122</a>.</p> + +<p><i>Cupular</i>, or <i>Cupulate</i>, provided with a cupule.</p> + +<p><i>Cupuliferous</i>, cupule-bearing.</p> + +<p><i>Curviveined</i>, with curved ribs or veins.</p> + +<p><i>Curviserial</i>, in oblique or spiral ranks.</p> + +<p><i>Cushion</i>, the enlargement at the insertion or base of a petiole.</p> + +<p><i>Cuspidate</i>, tipped with a sharp and stiff point or <i>cusp</i>, <a href="#Page_54">54</a>.</p> + +<p><i>Cut</i>, same as incised, or applied generally to any sharp and deep +division, <a href="#Page_55">55</a>.</p> + +<p><i>Cuticle</i>, the skin of plants, or more strictly its external pellicle.</p> + +<p><i>Cyaneous</i>, bright blue.</p> + +<p><i>Cyathiform</i>, in the shape of a cup, or particularly of a wine-glass.</p> + +<p><i>Cycle</i>, one complete turn of a spire, or a circle, <a href="#Page_70">70</a>.</p> + +<span class="pagenum"><a name="Page_202" id="Page_202">[Pg 202]</a></span><p><i>Cyclical</i>, rolled up circularly, or coiled into a complete circle.</p> + +<p><i>Cyclosis</i>, circulation in closed cells, <a href="#Page_149">149</a>.</p> + +<p><i>Cylindraceous</i>, approaching to the <i>Cylindrical</i> form, terete and not +tapering.</p> + +<p><i>Cymbæform</i>, or <i>Cymbiform</i>, same as boat-shaped.</p> + +<p><i>Cyme</i>, a cluster of centrifugal inflorescence, <a href="#Page_77">77</a>.</p> + +<p><i>Cymose</i>, furnished with cymes, or like a cyme.</p> + +<p><i>Cymule</i>, a partial or diminutive cyme, <a href="#Page_77">77</a>.</p> +</div> + +<div class="index"> +<p><i>Deca-</i> (in words of Greek derivation), ten; as</p> + +<p><i>Decagynous</i>, with 10 pistils or styles, <i>Decamerous</i>, of 10 parts, +<i>Decandrous</i>, with 10 stamens, &c.</p> + +<p><i>Deciduous</i>, falling off, or subject to fall; said of leaves which fall +in autumn, and of a calyx and corolla which fall before the fruit forms.</p> + +<p><i>Declinate</i>, <i>declined</i>, turned to one side, or downwards.</p> + +<p><i>Decompound</i>, several times compounded or divided, <a href="#Page_59">59</a>.</p> + +<p><i>Decumbent</i>, reclined on the ground, the summit tending to rise, <a href="#Page_39">39</a>.</p> + +<p><i>Decurrent</i> (leaves), prolonged on the stem beneath the insertion, as in +Thistles.</p> + +<p><i>Decussate</i>, arranged in pairs which successively cross each other, <a href="#Page_71">71</a>.</p> + +<p><i>Deduplication</i>, same as chorisis.</p> + +<p><i>Definite</i>, when of a uniform number, and not above twelve or so.</p> + +<p><i>Definite Inflorescence</i>, <a href="#Page_72">72</a>.</p> + +<p><i>Deflexed</i>, bent downwards.</p> + +<p><i>Deflorate</i>, past the flowering state, as an anther after it has +discharged its pollen.</p> + +<p><i>Dehiscence</i>, the regular splitting open of capsule or anther, <a href="#Page_103">103</a>, <a href="#Page_119">119</a>.</p> + +<p><i>Dehiscent</i>, opening by regular dehiscence, <a href="#Page_119">119</a>, <a href="#Page_123">123</a>.</p> + +<p><i>Deliquescent</i>, branching off so that the stem is lost in the branches, +<a href="#Page_32">32</a>.</p> + +<p><i>Deltoid</i>, of a triangular shape, like the Greek capital Δ.</p> + +<p><i>Demersed</i>, growing below the surface of water.</p> + +<p><i>Dendroid</i>, <i>Dendritic</i>, tree-like in form or appearance.</p> + +<p><i>Dendron</i>, Greek for tree.</p> + +<p><i>Deni</i>, ten together.</p> + +<p><i>Dens</i>, Latin for tooth.</p> + +<p><i>Dentate</i>, toothed, <a href="#Page_55">55</a>. <i>Denticulate</i>, furnished with denticulations, or +little teeth.</p> + +<p><i>Depauperate</i>, impoverished or starved, and so below the natural size.</p> + +<p><i>Depressed</i>, flattened or as if pressed down from above.</p> + +<p><i>Derma</i>, Greek for skin.</p> + +<p><i>Descending</i>, tending gradually downwards. <i>Descending axis</i>, the root.</p> + +<p><i>Desmos</i>, Greek for things connected or bound together.</p> + +<p><i>Determinate Inflorescence</i>, <a href="#Page_72">72</a>.</p> + +<p><i>Dextrorse</i>, turned to the right hand.</p> + +<p><i>Di- Dis</i> (in Greek compounds) two, as</p> + +<p><i>Diadelphous</i> (stamens), united by their filaments in two sets, <a href="#Page_99">99</a>.</p> + +<p><i>Diagnosis</i>, a short distinguishing character or descriptive phrase.</p> + +<p><i>Dialypetalous</i>, same as polypetalous.</p> + +<p><i>Diandrous</i>, having two stamens, &c.</p> + +<p><i>Diaphanous</i>, transparent or translucent.</p> + +<p><i>Dicarpellary</i>, of two carpels.</p> + +<p><i>Dichlamydeous</i> (flower), having both calyx and corolla.</p> + +<p><i>Dichogamous</i>, <i>Dichogamy</i>, <a href="#Page_116">116</a>.</p> + +<p><i>Dichotomous</i>, two-forked.</p> + +<p><i>Diclinous</i>, having the stamens in one flower, the pistils in another, +<a href="#Page_85">85</a>.</p> + +<p><i>Dicoccous</i> (fruit), splitting into two <i>cocci</i> or closed carpels.</p> + +<p><i>Dicotyls</i>, <a href="#Page_23">23</a>.</p> + +<p><i>Dicotyledonous</i> (embryo), having a pair of cotyledons, <a href="#Page_23">23</a>. +<i>Dicotyledonous Plants</i>, <a href="#Page_23">23</a>, <a href="#Page_182">182</a>.</p> + +<p><i>Didymous</i>, twin.</p> + +<p><i>Didynamous</i> (stamens), having four stamens in two pairs, <a href="#Page_100">100</a>.</p> + +<p><i>Diffuse</i>, spreading widely and irregularly.</p> + +<span class="pagenum"><a name="Page_203" id="Page_203">[Pg 203]</a></span><p><i>Digitate</i> (fingered), where the leaflets of a compound leaf are all +borne on the apex of the petiole, <a href="#Page_58">58</a>.</p> + +<p><i>Digynous</i> (flower), having two pistils or styles, <a href="#Page_105">105</a>.</p> + +<p><i>Dimerous</i>, made up of two parts, or its organs in twos.</p> + +<p><i>Dimidiate</i>, halved; as where a leaf or leaflet has only one side +developed.</p> + +<p><i>Dimorphism</i>, <a href="#Page_117">117</a>. <i>Dimorphous</i>, <i>Dimorphic</i>, of two forms, <a href="#Page_117">117</a>.</p> + +<p><i>Diœcious</i>, or <i>Dioicous</i>, with stamens and pistils on different +plants, <a href="#Page_85">85</a>.</p> + +<p><i>Dipetalous</i>, of two petals.</p> + +<p><i>Diphyllous</i>, two-leaved.</p> + +<p><i>Dipterous</i>, two-winged.</p> + +<p><i>Diplo-</i>, Greek for double, as <i>Diplostemonous</i>, with two sets of +stamens.</p> + +<p><i>Disciform</i> or <i>Disk-shaped</i>, flat and circular, like a disk or quoit.</p> + +<p><i>Discoidal</i>, or <i>Discoid</i>, belonging to or like a disk.</p> + +<p><i>Discolor</i>, of two different colors or hues.</p> + +<p><i>Discrete</i>, separate, opposite of concrete.</p> + +<p><i>Disepalous</i>, of two sepals.</p> + +<p><i>Disk</i>, the face of any flat body; the central part of a head of +flowers, like the Sunflower, or Coreopsis, as opposed to the ray or +margin; a fleshy expansion of the receptacle of a flower, <a href="#Page_113">113</a>.</p> + +<p><i>Disk-flowers</i>, those of the disk in Compositæ.</p> + +<p><i>Dissected</i>, cut deeply into many lobes or divisions.</p> + +<p><i>Dissepiments</i>, the partitions of a compound ovary or a fruit, <a href="#Page_108">108</a>.</p> + +<p><i>Dissilient</i>, bursting in pieces.</p> + +<p><i>Distichous</i>, two-ranked.</p> + +<p><i>Distinct</i>, uncombined with each other, <a href="#Page_95">95</a>.</p> + +<p><i>Dithecous</i>, of two thecæ or anther-cells.</p> + +<p><i>Divaricate</i>, straddling; very widely divergent.</p> + +<p><i>Divided</i> (leaves, &c.), cut into divisions down to the base or midrib, +<a href="#Page_55">55</a>.</p> + +<p><i>Dodeca</i>, Greek for twelve; as <i>Dodecagynous</i>, with twelve pistils or +styles, <i>Dodecandrous</i>, with twelve stamens.</p> + +<p><i>Dodrans</i>, span-long.</p> + +<p><i>Dolabriform</i>, axe-shaped.</p> + +<p><i>Dorsal</i>, pertaining to the back (<i>dorsum</i>) of an organ. <i>Dorsal +Suture</i>, <a href="#Page_106">106</a>.</p> + +<p><i>Dotted Ducts</i>, <a href="#Page_148">148</a>.</p> + +<p><i>Double Flowers</i>, where the petals are multiplied unduly, <a href="#Page_79">79</a>.</p> + +<p><i>Downy</i>, clothed with a coat of soft and short hairs.</p> + +<p><i>Drupaceous</i>, like or pertaining to a drupe.</p> + +<p><i>Drupe</i>, a stone-fruit, <a href="#Page_120">120</a>. <i>Drupelet</i> or <i>Drupel</i>, a little drupe.</p> + +<p><i>Ducts</i>, the so-called vessels of plants, <a href="#Page_134">134</a>.</p> + +<p><i>Dumose</i>, bushy, or relating to bushes.</p> + +<p><i>Duramen</i>, the heart-wood, <a href="#Page_142">142</a>.</p> + +<p><i>Dwarf</i>, remarkably low in stature.</p> +</div> + +<div class="index"> +<p><i>E-</i>, as a prefix of Latin compound words, means destitute of; as +<i>ecostate</i>, without a rib or midrib; <i>exalbuminous</i>, without albumen, +&c.</p> + +<p><i>Eared</i>, see <a href="#Auriculate"><i>auriculate</i></a>, <a href="#Page_53">53</a>.</p> + +<p><i>Ebracteate</i>, destitute of bracts. <i>Ebracteolate</i>, destitute of +bractlets.</p> + +<p><i>Eburneous</i>, ivory-white.</p> + +<p><i>Echinate</i>, armed with prickles (like a hedgehog). <i>Echinulate</i>, a +diminutive of it.</p> + +<p><i>Edentate</i>, toothless.</p> + +<p><i>Effete</i>, past bearing, &c.; said of anthers which have discharged their +pollen.</p> + +<p><i>Effuse</i>, very loosely branched and spreading.</p> + +<p><i>Eglandulose</i>, destitute of glands.</p> + +<p><i>Elaters</i>, threads mixed with the spores of Liverworts, <a href="#Page_165">165</a>.</p> + +<p><i>Ellipsoidal</i>, approaching an elliptical figure.</p> + +<p><i>Elliptical</i>, oval or oblong, with the ends regularly rounded, <a href="#Page_52">52</a>.</p> + +<p><i>Emarginate</i>, notched at the summit, <a href="#Page_54">54</a>.</p> + +<p><i>Embryo</i>, the rudimentary plantlet in a seed, <a href="#Page_11">11</a>, <a href="#Page_127">127</a>.</p> + +<p><i>Embryonal</i>, belonging or relating to the embryo.</p> + +<p><i>Embryo-sac</i>, <a href="#Page_117">117</a>.</p> + +<span class="pagenum"><a name="Page_204" id="Page_204">[Pg 204]</a></span><p><i>Emersed</i>, raised out of water.</p> + +<p><i>Endecagynous</i>, with eleven pistils or styles.</p> + +<p><i>Endecandrous</i>, with eleven stamens.</p> + +<p><i>Endemic</i>, peculiar to the country geographically.</p> + +<p><i>Endocarp</i>, the inner layer of a pericarp or fruit, <a href="#Page_120">120</a>.</p> + +<p><i>Endochrome</i>, the coloring matter of Algæ and the like.</p> + +<p><i>Endogenous Stems</i>, <a href="#Page_138">138</a>. <i>Endogenous plants</i>, an old name for +monocotyledons.</p> + +<p><i>Endopleura</i>, inner seed-coat.</p> + +<p><i>Endorhizal</i>, radicle or root sheathed in germination.</p> + +<p><i>Endosperm</i>, the albumen of a seed, <a href="#Page_21">21</a>.</p> + +<p><i>Endostome</i>, the orifice in the inner coat of an ovule.</p> + +<p><i>Ennea-</i>, nine. <i>Enneagynous</i>, with nine petals or styles. +<i>Enneandrous</i>, nine-stamened.</p> + +<p><i>Ensate</i>, <i>Ensiform</i>, sword-shaped.</p> + +<p><i>Entire</i>, the margins not at all toothed, notched, or divided, but even, +<a href="#Page_55">55</a>.</p> + +<p><i>Entomophilous</i>, said of flowers frequented and fertilized by insects, +<a href="#Page_113">113</a>.</p> + +<p><i>Ephemeral</i>, lasting for a day or less, as the corolla of Purslane, &c.</p> + +<p><i>Epi-</i>, Greek for upon.</p> + +<p><i>Epicalyx</i>, such an involucel as that of Malvaceæ.</p> + +<p><i>Epicarp</i>, the outermost layer of a fruit, <a href="#Page_120">120</a>.</p> + +<p><i>Epidermal</i>, relating to the <i>Epidermis</i>, or skin of a plant, <a href="#Page_50">50</a>, <a href="#Page_141">141</a>, +<a href="#Page_143">143</a>.</p> + +<p><i>Epigæous</i>, growing on the earth, or close to the ground.</p> + +<p><i>Epigynous</i>, upon the ovary, <a href="#Page_95">95</a>, <a href="#Page_99">99</a>.</p> + +<p><i>Epipetalous</i>, borne on the petals or the corolla, <a href="#Page_99">99</a>.</p> + +<p><i>Epiphyllous</i>, borne on a leaf.</p> + +<p><i>Epiphyte</i>, a plant growing on another plant, but not nourished by it, +<a href="#Page_36">36</a>.</p> + +<p><i>Epiphytic</i> or <i>Epiphytal</i>, relating to <i>Epiphytes</i>.</p> + +<p><i>Epipterous</i>, winged at top.</p> + +<p><i>Episperm</i>, the skin or coat of a seed, especially the outer coat.</p> + +<p><i>Equal</i>, alike in number or length.</p> + +<p><i>Equally pinnate</i>, same as abruptly pinnate, <a href="#Page_57">57</a>.</p> + +<p><i>Equitant</i> (riding straddle), <a href="#Page_60">60</a>.</p> + +<p><i>Erion</i>, Greek for wool. <i>Erianthous</i>, woolly-flowered. <i>Eriophorous</i>, +wool-bearing, &c.</p> + +<p><i>Erose</i>, eroded, as if gnawed.</p> + +<p><i>Erostrate</i>, not beaked.</p> + +<p><i>Erythros</i>, Greek for red. <i>Erythrocarpous</i>, red-fruited, &c.</p> + +<p><i>Essential Organs</i> of the flower, <a href="#Page_80">80</a>.</p> + +<p><i>Estivation</i>, see <a href="#aestivation"><i>æstivation</i></a>.</p> + +<p><i>Etiolated</i>, blanched by excluding the light, as the stalks of Celery.</p> + +<p><i>Eu</i>, Greek prefix, meaning very, or much.</p> + +<p><i>Evergreen</i>, holding the leaves over winter and until new ones appear, +or longer.</p> + +<p><i>Ex</i>, Latin prefix; privative in place of "e" when next letter is a +vowel. So <i>Exalate</i>, wingless; <i>Exalbuminous</i> (seed), without albumen, +<a href="#Page_21">21</a>.</p> + +<p><i>Excurrent</i>, running out, as when a midrib projects beyond the apex of a +leaf, or a trunk is continued to the very top of a tree, <a href="#Page_32">32</a>.</p> + +<p><i>Exiguous</i>, puny.</p> + +<p><i>Exilis</i>, lank or meagre.</p> + +<p><i>Eximius</i>, distinguished for size or beauty.</p> + +<p><i>Exo-</i>, in Greek compounds, outward, as in</p> + +<p><i>Exocarp</i>, outer layer of a pericarp, <a href="#Page_120">120</a>.</p> + +<p><i>Exogenous</i>, outward growing. <i>Exogenous stems</i>, <a href="#Page_139">139</a>.</p> + +<p><i>Exorhizal</i>, radicle in germination not sheathed.</p> + +<p><i>Exostome</i>, the orifice in the outer coat of the ovule.</p> + +<p><i>Explanate</i>, spread or flattened out.</p> + +<p><i>Exserted</i>, protruding out of, as the stamens out of the corolla.</p> + +<p><i>Exstipulate</i>, destitute of stipules.</p> + +<p><i>Extine</i>, outer coat of a pollen-grain.</p> + +<p><i>Extra-axillary</i>, said of a branch or bud somewhat out of the axil, <a href="#Page_31">31</a>.</p> + +<p><i>Extrorse</i>, turned outwards; the anther is extrorse when fastened to the +filament on the side next the pistil, and opening on the outer side, +<a href="#Page_101">101</a>.</p> +</div> + +<div class="index"> +<span class="pagenum"><a name="Page_205" id="Page_205">[Pg 205]</a></span><p><i>Falcate</i>, scythe-shaped; a flat body curved, its edges parallel.</p> + +<p><i>False Racemes</i>, <a href="#Page_78">78</a>.</p> + +<p><i>Family</i>, in botany same as Order, <a href="#Page_177">177</a>.</p> + +<p><i>Farina</i>, meal or starchy matter, <a href="#Page_136">136</a>.</p> + +<p><a name="Farinaceous" id="Farinaceous"></a><i>Farinaceous</i>, mealy in texture. <i>Farinose</i>, covered with a mealy +powder.</p> + +<p><i>Fasciate</i>, banded; also applied to monstrous stems which grow flat.</p> + +<p><i>Fascicle</i>, a close cluster, <a href="#Page_77">77</a>.</p> + +<p><i>Fascicled</i>, <i>Fasciculated</i>, growing in a bundle or tuft, as the leaves +of Larch, <a href="#Page_68">68</a>, and roots of Peony, <a href="#Page_35">35</a>.</p> + +<p><i>Fastigiate</i>, close, parallel, and upright, as the branches of Lombardy +Poplar.</p> + +<p><i>Faux</i> (plural, <i>fauces</i>), the throat of a calyx, corolla, &c., <a href="#Page_89">89</a>.</p> + +<p><i>Faveolate</i>, <i>Favose</i>, honeycombed; same as <i>alveolate</i>.</p> + +<p><i>Feather-veined</i>, with veins of a leaf all springing from the sides of a +midrib, <a href="#Page_51">51</a>.</p> + +<p><i>Fecula</i> or <i>Fæcula</i>, starch, <a href="#Page_136">136</a>.</p> + +<p><i>Female flower</i> or <i>plant</i>, one bearing pistils only.</p> + +<p><i>Fenestrate</i>, pierced with one or more large holes, like windows.</p> + +<p><i>Ferrugineous</i>, or <i>Ferruginous</i>, resembling iron-rust; red-grayish.</p> + +<p><i>Fertile</i>, fruit-bearing, or capable of it; also said of anthers +producing good pollen.</p> + +<p><i>Fertilization</i>, the process by which pollen causes the embryo to be +formed, <a href="#Page_114">114</a>.</p> + +<p><i>Fibre</i> (woody), <a href="#Page_133">133</a>. <i>Fibrous</i>, containing much fibre, or composed of +fibres.</p> + +<p><i>Fibrillose</i>, formed of small fibres, or <i>Fibrillæ</i>.</p> + +<p><i>Fibro-vascular</i> bundle or tissue, formed of fibres and vessels.</p> + +<p><a name="Fiddleshaped" id="Fiddleshaped"></a><i>Fiddle-shaped</i>, obovate with a deep recess on each side.</p> + +<p><i>Fidus</i>, Latin suffix for cleft, as <i>Bifid</i>, two-cleft.</p> + +<p><i>Filament</i>, the stalk of a stamen, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_101">101</a>; also any slender +thread-shaped body.</p> + +<p><i>Filamentose</i>, or <i>Filamentous</i>, bearing or formed of slender threads.</p> + +<p><i>Filiform</i>, thread-shaped; long, slender, and cylindrical.</p> + +<p><i>Fimbriate</i>, fringed; furnished with fringes (<i>fimbriæ</i>).</p> + +<p><i>Fimbrillate</i>, <i>Fimbrilliferous</i>, bearing small <i>fimbriæ</i>, i. e. +<i>fimbrillæ</i>.</p> + +<p><i>Fissiparous</i>, multiplying by division of one body into two.</p> + +<p><i>Fissus</i>, Latin for split or divided.</p> + +<p><i>Fistular</i>, or <i>Fistulose</i>, hollow and cylindrical, as the leaves of the +Onion.</p> + +<p><i>Flabelliform</i>, or <i>Flabellate</i>, fan-shaped.</p> + +<p><i>Flagellate</i>, or <i>Flagelliform</i>, long, narrow, and flexible, like the +thong of a whip; or like the runners (<i>flagellæ</i>) of the Strawberry.</p> + +<p><i>Flavescent</i>, yellowish, or turning yellow.</p> + +<p><i>Flavus</i>, Latin for yellow.</p> + +<p><i>Fleshy</i>, composed of firm pulp or flesh.</p> + +<p><i>Flexuose</i>, or <i>Flexuous</i>, bending in opposite directions, in a zigzag +way.</p> + +<p><i>Floating</i>, swimming on the surface of water.</p> + +<p><i>Floccose</i>, composed of or bearing tufts of woolly or long and soft +hairs.</p> + +<p><i>Flora</i> (the goddess of flowers), the plants of a country or district, +taken together, or a work systematically describing them, <a href="#Page_9">9</a>.</p> + +<p><i>Floral Envelopes</i>, or <i>Flower-leaves</i>, <a href="#Page_79">79</a>.</p> + +<p><i>Floret</i>, a diminutive flower, one of a mass or cluster.</p> + +<p><i>Floribund</i>, abundantly floriferous.</p> + +<p><i>Florula</i>, the flora of a small district.</p> + +<p><i>Flos</i>, <i>floris</i>, Latin for flower.</p> + +<p><i>Flosculus</i>, diminutive, same as floret.</p> + +<p><i>Flower</i>, the whole organs of reproduction of Phænogamous plants, <a href="#Page_14">14</a>, +<a href="#Page_72">72</a>.</p> + +<p><i>Flower-bud</i>, an unopened flower.</p> + +<p><i>Flowering Plants</i>, <a href="#Page_10">10</a>, <a href="#Page_156">156</a>. <i>Flowerless Plants</i>, <a href="#Page_10">10</a>, <a href="#Page_156">156</a>.</p> + +<p><i>Fly-trap leaves</i>, <a href="#Page_65">65</a>.</p> + +<p><i>Fluitans</i>, Latin for floating. <i>Fluviatile</i>, belonging to a river or +stream.</p> + +<p><i>Foliaceous</i>, belonging to, or of the texture or nature of, a leaf +(<i>folium</i>).</p> + +<p><i>Foliate</i>, provided with leaves. Latin prefixes denote the number of +leaves, as <i>bifoliate</i>, <i>trifoliate</i>, &c. <i>Foliose</i>, leafy; abounding in +leaves.</p> + +<p><i>Foliolate</i>, relating to or bearing leaflets (<i>foliola</i>); +<i>trifoliolate</i>, with three leaflets, &c.</p> + +<span class="pagenum"><a name="Page_206" id="Page_206">[Pg 206]</a></span><p><i>Folium</i> (plural, <i>folia</i>), Latin for leaf.</p> + +<p><i>Follicle</i>, a simple pod, opening down the inner suture, <a href="#Page_122">122</a>.</p> + +<p><i>Follicular</i>, resembling or belonging to a follicle.</p> + +<p><i>Food of Plants</i>, <a href="#Page_144">144</a>.</p> + +<p><i>Foot-stalk</i>, either petiole or peduncle, <a href="#Page_49">49</a>.</p> + +<p><i>Foramen</i>, a hole or orifice, as that of the ovule, <a href="#Page_110">110</a>.</p> + +<p><i>Foraminose</i>, <i>Foraminulose</i>, pierced with holes.</p> + +<p><i>Forked</i>, branched in two or three or more.</p> + +<p><i>Fornicate</i>, bearing fornices.</p> + +<p><i>Fornix</i>, little arched scales in the throat of some corollas, as of +Comfrey.</p> + +<p><i>Foveate</i>, deeply pitted. <i>Foveolate</i>, diminutive of <i>foveate</i>.</p> + +<p><i>Free</i>, not united with any other parts of a different sort, <a href="#Page_95">95</a>.</p> + +<p><i>Fringed</i>, the margin beset with slender appendages, bristles, &c.</p> + +<p><i>Frond</i>, what answers to leaves in Ferns, &c., <a href="#Page_157">157</a>; or to the stem and +leaves fused into one, as in Liverwort.</p> + +<p><i>Frondescence</i>, the bursting into leaf.</p> + +<p><i>Frondose</i>, frond-bearing; like a frond, or sometimes used for leafy.</p> + +<p><i>Fructification</i>, the state or result of fruiting.</p> + +<p><i>Fructus</i>, Latin for fruit.</p> + +<p><i>Fruit</i>, the matured ovary and all it contains or is connected with, +<a href="#Page_117">117</a>.</p> + +<p><i>Fruit-dots</i> in Ferns; see <a href="#Sorus"><i>Sorus</i></a>.</p> + +<p><i>Frustulose</i>, consisting of a chain of similar pieces, or <i>Frustules</i>.</p> + +<p><i>Frutescent</i>, somewhat shrubby; becoming a shrub (<i>Frutex</i>), <a href="#Page_39">39</a>.</p> + +<p><i>Fruticulose</i>, like a small shrub, or <i>Fruticulus</i>. <i>Fruticose</i>, +shrubby, <a href="#Page_39">39</a>.</p> + +<p><i>Fugacious</i>, soon falling off or perishing.</p> + +<p><i>Fulcrate</i>, having accessory organs or <i>fulcra</i>, i. e. props.</p> + +<p><i>Fulvous</i>, tawny; dull yellow with gray.</p> + +<p><i>Fungus</i>, <i>Fungi</i>, <a href="#Page_172">172</a>.</p> + +<p><i>Funicle</i>, <i>Funiculus</i>, the stalk of a seed or ovule, <a href="#Page_110">110</a>.</p> + +<p><i>Funnelform</i>, or <i>funnel-shaped</i>, expanding gradually upwards into an +open mouth, like a funnel or tunnel, <a href="#Page_90">90</a>.</p> + +<p><i>Furcate</i>, forked.</p> + +<p><i>Furfuraceous</i>, covered with bran-like fine scurf.</p> + +<p><i>Furrowed</i>, marked by longitudinal channels or grooves.</p> + +<p><i>Fuscous</i>, deep gray-brown.</p> + +<p><i>Fusiform</i>, spindle-shaped, <a href="#Page_36">36</a>.</p> +</div> + +<div class="index"> +<p><i>Galbalus</i>, the fleshy or at length woody cone of Juniper and Cypress.</p> + +<p><i>Galea</i>, a helmet-shaped body, as the upper sepal of the Monkshood, <a href="#Page_87">87</a>.</p> + +<p><i>Galeate</i>, shaped like a helmet.</p> + +<p><i>Gamopetalous</i>, of united petals, <a href="#Page_89">89</a>.</p> + +<p><i>Gamophyllous</i>, formed of united leaves.</p> + +<p><i>Gamosepalous</i>, formed of united sepals, <a href="#Page_89">89</a>.</p> + +<p><i>Geminate</i>, twin; in pairs.</p> + +<p><i>Gemma</i>, Latin for a bud.</p> + +<p><i>Gemmation</i>, the state of budding; budding growth.</p> + +<p><i>Gemmule</i>, a small bud; the plumule, <a href="#Page_13">13</a>.</p> + +<p><i>Genera</i>, plural of genus.</p> + +<p><i>Geniculate</i>, bent abruptly, like a knee (<i>genu</i>), as many stems.</p> + +<p><i>Generic Names</i>, <a href="#Page_179">179</a>.</p> + +<p><i>Genus</i>, a kind of a rank above species, <a href="#Page_177">177</a>.</p> + +<p><i>Germ</i>, a growing point; a young bud; sometimes the same as embryo, <a href="#Page_127">127</a>.</p> + +<p><i>Germen</i>, the old name for ovary.</p> + +<p><i>Germination</i>, the development of a plantlet from the seed, <a href="#Page_12">12</a>.</p> + +<p><i>Gerontogæous</i>, inhabiting the Old World.</p> + +<p><i>Gibbous</i>, more tumid at one place or on one side than the other.</p> + +<p><i>Gilvous</i>, dirty reddish-yellow.</p> + +<p><i>Glabrate</i>, becoming glabrous with age, or almost glabrous.</p> + +<p><i>Glabrous</i>, smooth, in the sense of having no hairs, bristles, or other +pubescence.</p> + +<span class="pagenum"><a name="Page_207" id="Page_207">[Pg 207]</a></span><p><i>Gladiate</i>, sword-shaped, as the leaves of Iris.</p> + +<p><i>Glands</i>, small cellular organs which secrete oily or aromatic or other +products; they are sometimes sunk in the leaves or rind, as in the +Orange, Prickly Ash, &c.; sometimes on the surface as small projections; +sometimes raised on hairs or bristles (<i>glandular hairs, &c.</i>), as in +the Sweetbrier and Sundew. The name is also given to any small +swellings, &c., whether they secrete anything or not; so that the word +is loosely used.</p> + +<p><i>Glandular</i>, <i>Glandulose</i>, furnished with glands, or gland-like.</p> + +<p><i>Glans</i> (<i>Gland</i>), the acorn or mast of Oak and similar fruits.</p> + +<p><i>Glareose</i>, growing in gravel.</p> + +<p><i>Glaucescent</i>, slightly glaucous, or bluish-gray.</p> + +<p><i>Glaucous</i>, covered with a <i>bloom</i>, viz. with a fine white powder of wax +that rubs off, like that on a fresh plum, or a cabbage-leaf.</p> + +<p><i>Globose</i>, spherical in form, or nearly so. <i>Globular</i>, nearly globose.</p> + +<p><i>Glochidiate</i>, or <i>Glochideous</i>, (bristles) barbed; tipped with barbs, +or with a double hooked point.</p> + +<p><i>Glomerate</i>, closely aggregated into a dense cluster.</p> + +<p><i>Glomerule</i>, a dense head-like cluster, <a href="#Page_77">77</a>.</p> + +<p><i>Glossology</i>, the department of botany in which technical terms are +explained.</p> + +<p><i>Glumaceous</i>, glume-like, or glume-bearing.</p> + +<p><i>Glume</i>; Glumes are the husks or floral coverings of Grasses, or, +particularly, the outer husks or bracts of each spikelet.</p> + +<p><i>Glumelles</i>, the inner husks of Grasses.</p> + +<p><i>Gonophore</i>, a stipe below stamens, <a href="#Page_113">113</a>.</p> + +<p><i>Gossypine</i>, cottony, flocculent.</p> + +<p><i>Gracilis</i>, Latin for slender.</p> + +<p><i>Grain</i>, see <a href="#Caryopsis"><i>Caryopsis</i></a>, <a href="#Page_121">121</a>.</p> + +<p><i>Gramineous</i>, grass-like.</p> + +<p><i>Granular</i>, composed of grains. <i>Granule</i>, a small grain.</p> + +<p><i>Graveolent</i>, heavy-scented.</p> + +<p><i>Griseous</i>, gray or bluish-gray.</p> + +<p><i>Growth</i>, <a href="#Page_129">129</a>.</p> + +<p><i>Grumous</i>, or <i>Grumose</i>, formed of coarse clustered grains.</p> + +<p><i>Guttate</i>, spotted, as if by drops of something colored.</p> + +<p><i>Gymnos</i>, Greek for naked, as</p> + +<p><i>Gymnocarpous</i>, naked-fruited.</p> + +<p><i>Gymnospermous</i>, naked-seeded, <a href="#Page_109">109</a>.</p> + +<p><i>Gymnospermous gynœcium</i>, <a href="#Page_109">109</a>.</p> + +<p><i>Gymnospermæ</i>, or <i>Gymnospermous Plants</i>, <a href="#Page_183">183</a>.</p> + +<p><i>Gynandrous</i>, with stamens borne on, i. e. united with, the pistil, <a href="#Page_99">99</a>.</p> + +<p><i>Gynœcium</i>, a name for the pistils of a flower taken altogether, <a href="#Page_105">105</a>.</p> + +<p><i>Gynobase</i>, a depressed receptacle or support of the pistil or carpels, +<a href="#Page_114">114</a>.</p> + +<p><i>Gynophore</i>, a stalk raising a pistil above the stamens, <a href="#Page_113">113</a>.</p> + +<p><i>Gynostegium</i>, a sheath around pistils, of whatever nature.</p> + +<p><i>Gynostemium</i>, name of the column in Orchids, &c., consisting of style +and stigma with stamens combined.</p> + +<p><i>Gyrate</i>, coiled or moving circularly.</p> + +<p><i>Gyrose</i>, strongly bent to and fro.</p> +</div> + +<div class="index"> +<p><i>Habit</i>, the general aspect of a plant, or its mode of growth.</p> + +<p><i>Habitat</i>, the situation or country in which a plant grows in a wild +state.</p> + +<p><i>Hairs</i>, hair-like growths on the surface of plants.</p> + +<p><i>Hairy</i>, beset with hairs, especially longish ones.</p> + +<p><i>Halberd-shaped</i>, see <a href="#Hastate"><i>hastate</i></a>, <a href="#Page_53">53</a>.</p> + +<p><i>Halved</i>, when appearing as if one half of the body were cut away.</p> + +<p><i>Hamate</i>, or <i>Hamose</i>, hooked; the end of a slender body bent round.</p> + +<p><i>Hamulose</i>, bearing a small hook; a diminutive of the last.</p> + +<p><i>Haplo-</i>, in Greek compounds, single; as <i>Haplostemonous</i>, having only +one series of stamens.</p> + +<span class="pagenum"><a name="Page_208" id="Page_208">[Pg 208]</a></span><p><a name="Hastate" id="Hastate"></a><i>Hastate</i>, or <i>Hastile</i>, shaped like a halberd; furnished with a +spreading lobe on each side at the base, <a href="#Page_53">53</a>.</p> + +<p><i>Head</i>, capitulum, a form of inflorescence, <a href="#Page_74">74</a>.</p> + +<p><i>Heart-shaped</i>, of the shape of a heart as painted on cards, <a href="#Page_53">53</a>.</p> + +<p><i>Heart-wood</i>, the older or matured wood of exogenous trees, <a href="#Page_142">142</a>.</p> + +<p><i>Helicoid</i>, coiled like a <i>helix</i> or snail-shell, <a href="#Page_77">77</a>.</p> + +<p><i>Helmet</i>, the upper sepal of Monkshood is so called.</p> + +<p><i>Helvolous</i>, grayish-yellow.</p> + +<p><i>Hemi-</i> in compounds from the Greek, half; e. g. <i>Hemispherical</i>, &c.</p> + +<p><i>Hemicarp</i>, half-fruit, one carpel of an Umbelliferous plant, <a href="#Page_121">121</a>.</p> + +<p><i>Hemitropous</i> (ovule or seed), nearly same as <i>amphitropous</i>, <a href="#Page_123">123</a>.</p> + +<p><i>Hepta-</i> (in words of Greek origin), seven; as <i>Heptagynous</i>, with seven +pistils or styles. <i>Heptamerous</i>, its parts in sevens. <i>Heptandrous</i>, +having seven stamens.</p> + +<p><i>Herb</i>, plant not woody, at least above ground.</p> + +<p><i>Herbaceous</i>, of the texture of an herb; not woody, <a href="#Page_39">39</a>.</p> + +<p><i>Herbarium</i>, the botanist's arranged collection of dried plants, <a href="#Page_186">186</a>.</p> + +<p><i>Herborization</i>, <a href="#Page_184">184</a>.</p> + +<p><i>Hermaphrodite</i> (flower), having stamens and pistils in the same +blossom, <a href="#Page_81">81</a>.</p> + +<p><i>Hesperidium</i>, orange-fruit, a hard-rinded berry.</p> + +<p><i>Hetero-</i>, in Greek compounds, means of two or more sorts, as</p> + +<p><i>Heterocarpous</i>, bearing fruit of two kinds or shapes.</p> + +<p><i>Heterogamous</i>, bearing two or more sorts of flowers in one cluster.</p> + +<p><i>Heterogony</i>, <i>Heterogone</i>, or <i>Heterogonous</i>, with stamens and pistil +reciprocally of two sorts, <a href="#Page_116">116</a>. <i>Heterostyled</i> is same.</p> + +<p><i>Heteromorphous</i>, of two or more shapes.</p> + +<p><i>Heterophyllous</i>, with two sorts of leaves.</p> + +<p><i>Heterotropous</i> (ovule), the same as <i>amphitropous</i>, <a href="#Page_123">123</a>.</p> + +<p><i>Hexa-</i> (in Greek compounds), six; as <i>Hexagonal</i>, six-angled. +<i>Hexagynous</i>, with six pistils or styles. <i>Hexamerous</i>, its parts in +sixes. <i>Hexandrous</i>, with six stamens. <i>Hexapterous</i>, six-winged.</p> + +<p><i>Hibernaculum</i>, a winter bud.</p> + +<p><a name="Hiemal" id="Hiemal"></a><i>Hiemal</i>, relating to winter.</p> + +<p><i>Hilar</i>, belonging to the hilum.</p> + +<p><i>Hilum</i>, the scar of the seed; its place of attachment, <a href="#Page_110">110</a>, <a href="#Page_126">126</a>.</p> + +<p><i>Hippocrepiform</i>, horseshoe-shaped.</p> + +<p><i>Hirsute</i>, clothed with stiffish or beard-like hairs.</p> + +<p><i>Hirtellous</i>, minutely hirsute.</p> + +<p><i>Hispid</i>, bristly, beset with stiff hairs. <i>Hispidulous</i>, diminutive of +hispid.</p> + +<p><i>Histology</i>, <a href="#Page_9">9</a>.</p> + +<p><i>Hoary</i>, grayish-white; see <a href="#Canescent"><i>canescent</i></a>, &c.</p> + +<p><i>Holosericeous</i>, all over sericeous or silky.</p> + +<p><i>Homo-</i>, in Greek compounds, all alike or of one sort.</p> + +<p><i>Homodromous</i>, running in one direction.</p> + +<p><i>Homogamous</i>, a head or cluster with flowers all of one kind.</p> + +<p><i>Homogeneous</i>, uniform in nature; all of one kind.</p> + +<p><i>Homogone</i>, or <i>Homogonous</i>, counterpart of <i>Heterogone</i> or +<i>Homostyled</i>.</p> + +<p><i>Homologous</i>, of same type; thus petals and sepals are the homologues of +leaves.</p> + +<p><i>Homomallous</i> (leaves, &c.), originating all round an axis, but all bent +or curved to one side.</p> + +<p><i>Homorphous</i>, all of one shape.</p> + +<p><i>Homotropous</i> (embryo), curved with the seed; curved only one way.</p> + +<p><i>Hood</i>, same as <i>helmet</i> or <i>galea</i>. <i>Hooded</i>, hood-shaped; see +<a href="#Cucullate"><i>cucullate</i></a>.</p> + +<p><i>Hooked</i>, same as <i>hamate</i>.</p> + +<p><i>Horn</i>, a spur or some similar appendage. <i>Horny</i>, of the texture of +horn.</p> + +<p><i>Hortensis</i>, pertaining to the garden.</p> + +<p><i>Hortus Siccus</i>, an herbarium, or collection of dried plants, <a href="#Page_186">186</a>.</p> + +<p><i>Humifuse</i>, <i>Humistrate</i>, spread over the surface of the ground.</p> + +<p><i>Humilis</i>, low in stature.</p> + +<span class="pagenum"><a name="Page_209" id="Page_209">[Pg 209]</a></span><p><i>Hyaline</i>, transparent, or partly so.</p> + +<p><i>Hybrid</i>, a cross-breed between two allied species, <a href="#Page_176">176</a>.</p> + +<p><i>Hydrophytes</i>, water-plants.</p> + +<p><i>Hyemal</i>, see <a href="#Hiemal"><i>hiemal</i></a>.</p> + +<p><i>Hymenium</i> of a Mushroom, <a href="#Page_172">172</a>.</p> + +<p><i>Hypanthium</i>, a hollow flower-receptacle, such as that of Rose.</p> + +<p><i>Hypo-</i>, Greek prefix for under, or underneath.</p> + +<p><i>Hypocotyle</i>, or <i>Hypocotyl</i>, part of stem below the cotyledons, <a href="#Page_11">11</a>.</p> + +<p><i>Hypocrateriform</i>, properly <i>Hypocraterimorphous</i>, salver-shaped.</p> + +<p><i>Hypogæan</i>, or <i>Hypogæous</i>, produced under ground, <a href="#Page_19">19</a>.</p> + +<p><i>Hypogynous</i>, inserted under the pistil, <a href="#Page_95">95</a>, <a href="#Page_99">99</a>.</p> + +<p><i>Hysteranthous</i>, with the blossoms developed earlier than the leaves.</p> +</div> + +<div class="index"> +<p><i>Icosandrous</i>, having 20 (or 12 or more) stamens inserted on the calyx.</p> + +<p><i>Imberbis</i>, Latin for beardless.</p> + +<p><i>Imbricate</i>, <i>Imbricated</i>, <i>Imbricative</i>, overlapping one another, like +tiles or shingles on a roof, as the bud-scales of Horse-chestnut and +Hickory, <a href="#Page_27">27</a>. In æstivation, where some leaves of the calyx or corolla +are overlapped on both sides by others, <a href="#Page_98">98</a>.</p> + +<p><i>Immarginate</i>, destitute of a rim or border.</p> + +<p><i>Immersed</i>, growing wholly under water.</p> + +<p><i>Impari-pinnate</i>, pinnate with a single leaflet at the apex, <a href="#Page_57">57</a>.</p> + +<p><i>Imperfect flowers</i>, wanting either stamens or pistils, <a href="#Page_85">85</a>.</p> + +<p><i>Inæquilateral</i>, unequal-sided, as the leaf of a Begonia.</p> + +<p><i>Inane</i>, empty, said of an anther which produces no pollen, &c.</p> + +<p><i>Inappendiculate</i>, not appendaged.</p> + +<p><a name="Incanous" id="Incanous"></a><i>Incanous</i>, <i>Incanescent</i>, hoary with soft white pubescence.</p> + +<p><i>Incarnate</i>, flesh-colored.</p> + +<p><i>Incised</i>, cut rather deeply and irregularly, <a href="#Page_58">58</a>.</p> + +<p><i>Included</i>, enclosed; when the part in question does not project beyond +another.</p> + +<p><i>Incomplete Flower</i>, wanting calyx or corolla, <a href="#Page_86">86</a>.</p> + +<p><i>Incrassated</i>, thickened.</p> + +<p><i>Incubous</i>, with tip of one leaf lying flat over the base of the next +above.</p> + +<p><i>Incumbent</i>, leaning or resting upon; the cotyledons are incumbent when +the back of one of them lies against the radicle, <a href="#Page_128">128</a>; the anthers are +incumbent when turned or looking inwards.</p> + +<p><i>Incurved</i>, gradually curving inwards.</p> + +<p><i>Indefinite</i>, not uniform in number, or too numerous to mention (over +12).</p> + +<p><i>Indefinite</i> or <i>Indeterminate Inflorescence</i>, <a href="#Page_72">72</a>.</p> + +<p><i>Indehiscent</i>, not splitting open; i. e. not dehiscent, <a href="#Page_119">119</a>.</p> + +<p><i>Indigenous</i>, native to the country.</p> + +<p><i>Individuals</i>, <a href="#Page_175">175</a>.</p> + +<p><i>Indumentum</i>, any hairy coating or pubescence.</p> + +<p><i>Induplicate</i>, with the edges turned inwards, <a href="#Page_97">97</a>.</p> + +<p><i>Induviate</i>, clothed with old and withered parts or <i>induviæ</i>.</p> + +<p><i>Indusium</i>, the shield or covering of a fruit-dot of a Fern, <a href="#Page_159">159</a>.</p> + +<p><i>Inermis</i>, Latin for unarmed, not prickly.</p> + +<p><i>Inferior</i>, growing below some other organ, <a href="#Page_96">96</a>.</p> + +<p><i>Infertile</i>, not producing seed, or pollen, as the case may be.</p> + +<p><i>Inflated</i>, turgid and bladdery.</p> + +<p><i>Inflexed</i>, bent inwards.</p> + +<p><i>Inflorescence</i>, the arrangement of flowers on the stem, <a href="#Page_72">72</a>.</p> + +<p><i>Infra-axillary</i>, situated beneath the axil.</p> + +<p><i>Infundibuliform</i> or <i>Infundibular</i>, funnel-shaped, <a href="#Page_90">90</a>.</p> + +<p><i>Innate</i> (anther), attached by its base to the very apex of the +filament, <a href="#Page_101">101</a>.</p> + +<p><i>Innovation</i>, a young shoot, or new growth.</p> + +<p><i>Insertion</i>, the place or the mode of attachment of an organ to its +support, <a href="#Page_95">95</a>, <a href="#Page_99">99</a>.</p> + +<p><i>Integer</i>, entire, not lobed. <i>Integerrimus</i>, quite entire, not serrate.</p> + +<span class="pagenum"><a name="Page_210" id="Page_210">[Pg 210]</a></span><p><i>Intercellular Passages</i> or <i>Spaces</i>, <a href="#Page_131">131</a>, <a href="#Page_143">143</a>.</p> + +<p><i>Interfoliaceous</i>, between the leaves of a pair or whorl.</p> + +<p><i>Internode</i>, the part of a stem between two nodes, <a href="#Page_13">13</a>.</p> + +<p><i>Interpetiolar</i>, between petioles.</p> + +<p><i>Interruptedly pinnate</i>, pinnate with small leaflets intermixed with +larger.</p> + +<p><i>Intine</i>, inner coat of a pollen grain.</p> + +<p><i>Intrafoliaceous</i> (stipules, &c.), placed between the leaf or petiole +and the stem.</p> + +<p><i>Introrse</i>, turned or facing inwards; i. e. towards the axis of the +flower, <a href="#Page_101">101</a>.</p> + +<p><i>Intruse</i>, as it were pushed inwards.</p> + +<p><i>Inversed</i> or <i>Inverted</i>, where the apex is in the direction opposite to +that of the organ it is compared with.</p> + +<p><i>Involucel</i>, a partial or small involucre, <a href="#Page_76">76</a>.</p> + +<p><i>Involucellate</i>, furnished with an involucel. <i>Involucrate</i>, furnished +with an involucre.</p> + +<p><i>Involucre</i>, a whorl or set of bracts around a flower, umbel, or head, +&c., <a href="#Page_74">74</a>, <a href="#Page_75">75</a>.</p> + +<p><i>Involute</i>, in vernation, <a href="#Page_72">72</a>; rolled inwards from the edges, <a href="#Page_97">97</a>.</p> + +<p><i>Irregular Flowers</i>, <a href="#Page_86">86</a>, <a href="#Page_91">91</a>.</p> + +<p><i>Isos</i>, Greek for equal in number. <i>Isomerous</i>, the same number in the +successive circles or sets. <i>Isostemonous</i>, the stamens equal in number +to the sepals or petals.</p> +</div> + +<div class="index"> +<p><i>Jointed</i>, separate or separable at one or more places into pieces, <a href="#Page_64">64</a>, +&c.</p> + +<p><i>Jugum</i> (plural <i>Juga</i>), Latin for a pair, as of leaflets,—thus +<i>Unijugate</i>, of a single pair; <i>Bijugate</i>, of two pairs, &c.</p> + +<p><i>Julaceus</i>, like a catkin or <i>Julus</i>.</p> +</div> + +<div class="index"> +<p><i>Keel</i>, a projecting ridge on a surface, like the keel of a boat; the +two anterior petals of a papilionaceous corolla, <a href="#Page_92">92</a>.</p> + +<p><i>Keeled</i>, furnished with a keel or sharp longitudinal ridge.</p> + +<p><i>Kermesine</i>, Carmine-red.</p> + +<p><i>Kernel</i> of the ovule and seed, <a href="#Page_110">110</a>.</p> + +<p><i>Key</i>, or <i>Key-fruit</i>, a Samara, <a href="#Page_122">122</a>.</p> + +<p><i>Kidney-shaped</i>, resembling the outline of a kidney, <a href="#Page_53">53</a>.</p> +</div> + +<div class="index"> +<p><i>Labellum</i>, the odd petal in the Orchis Family.</p> + +<p><i>Labiate</i>, same as <i>bilabiate</i> or two-lipped, <a href="#Page_92">92</a>.</p> + +<p><i>Labiatiflorous</i>, having flowers with bilabiate corolla.</p> + +<p><i>Labium</i> (plural, <i>Labia</i>), Latin for lip.</p> + +<p><i>Lacerate</i>, with margin appearing as if torn.</p> + +<p><i>Laciniate</i>, slashed; cut into deep narrow lobes or <i>Laciniæ</i>.</p> + +<p><i>Lactescent</i>, producing milky juice, as does the Milkweed, &c.</p> + +<p><i>Lacteus</i>, Latin for milk-white.</p> + +<p><i>Lacunose</i>, full of holes or gaps.</p> + +<p><i>Lacustrine</i>, belonging to lakes.</p> + +<p><i>Lævigate</i>, smooth as if polished. Latin, <i>Lævis</i>, smooth, as opposed to +rough.</p> + +<p><i>Lageniform</i>, gourd-shaped.</p> + +<p><i>Lagopous</i>, Latin, hare-footed; densely clothed with long soft hairs.</p> + +<p><i>Lamellar</i> or <i>Lamellate</i>, consisting of flat plates, <i>Lamellæ</i>.</p> + +<p><i>Lamina</i>, a plate or blade, the blade of a leaf, &c., <a href="#Page_49">49</a>.</p> + +<p><i>Lanate</i>, <i>Lanose</i>, woolly; clothed with long and soft entangled hairs.</p> + +<p><i>Lanceolate</i>, lance-shaped, <a href="#Page_52">52</a>.</p> + +<p><i>Lanuginous</i>, cottony or woolly.</p> + +<p><i>Latent buds</i>, concealed or undeveloped buds, <a href="#Page_30">30</a>.</p> + +<p><i>Lateral</i>, belonging to the side.</p> + +<p><i>Latex</i>, the milky juice, &c., of plants, <a href="#Page_135">135</a>.</p> + +<p><i>Lax</i> (<i>Laxus</i>), loose in texture, or sparse; the opposite of crowded.</p> + +<p><i>Leaf</i>, <a href="#Page_49">49</a>. <i>Leaf-buds</i>, <a href="#Page_31">31</a>.</p> + +<p><i>Leaflet</i>, one of the divisions or blades of a compound leaf, <a href="#Page_57">57</a>.</p> + +<p><i>Leaf-like</i>, same as <i>foliaceous</i>.</p> + +<p><i>Leathery</i>, of about the consistence of leather; coriaceous.</p> + +<span class="pagenum"><a name="Page_211" id="Page_211">[Pg 211]</a></span><p><i>Legume</i>, a simple pod which dehisces in two pieces, like that of the +Pea, <a href="#Page_122">122</a>.</p> + +<p><i>Leguminous</i>, belonging to legumes, or to the Leguminous Family.</p> + +<p><i>Lenticular</i>, lens-shaped; i. e. flattish and convex on both sides.</p> + +<p><i>Lappaceous</i>, bur-like.</p> + +<p><i>Lasio</i>, Greek for woolly or hairy, as <i>Lasianthus</i>, woolly-flowered.</p> + +<p><i>Lateritious</i>, brick-colored.</p> + +<p><i>Laticiferous</i>, containing latex, <a href="#Page_135">135</a>.</p> + +<p><i>Latus</i>, Latin for broad, as <i>Latifolius</i>, broad-leaved.</p> + +<p><i>Leaf-scar</i>, <i>Leaf-stalk</i>, petiole.</p> + +<p><i>Lenticels</i>, lenticular dots on young bark.</p> + +<p><i>Lentiginose</i>, as if freckled.</p> + +<p><i>Lepal</i>, a made-up word for a staminode.</p> + +<p><i>Lepis</i>, Greek for a scale, whence <i>Lepidote</i>, leprous; covered with +scurfy scales.</p> + +<p><i>Leptos</i>, Greek for slender; so <i>Leptophyllous</i>, slender-leaved.</p> + +<p><i>Leukos</i>, Greek for white; whence <i>Leucanthous</i>, white-flowered, &c.</p> + +<p><i>Liber</i>, the inner bark of Exogenous stems, <a href="#Page_140">140</a>.</p> + +<p><i>Lid</i>, see <a href="#Operculum"><i>operculum</i></a>.</p> + +<p><i>Ligneous</i>, or <i>Lignose</i>, woody in texture.</p> + +<p><i>Ligulate</i>, furnished with a ligule, <a href="#Page_93">93</a>.</p> + +<p><i>Ligule</i>, <i>Ligula</i>, the strap-shaped corolla in many Compositæ, <a href="#Page_93">93</a>; the +membranous appendage at the summit of the leaf-sheaths of most Grasses, +<a href="#Page_67">67</a>.</p> + +<p><i>Limb</i>, the border of a corolla, &c., <a href="#Page_89">89</a>.</p> + +<p><i>Limbate</i>, bordered (Latin, <i>Limbus</i>, a border).</p> + +<p><i>Line</i>, the twelfth of an inch; or French lines, the tenth.</p> + +<p><i>Linear</i>, narrow and flat, the margins parallel, <a href="#Page_52">52</a>.</p> + +<p><i>Lineate</i>, marked with parallel lines. <i>Lineolate</i>, marked with minute +lines.</p> + +<p><i>Lingulate</i>, <i>Linguiform</i>, tongue-shaped.</p> + +<p><i>Lip</i>, the principal lobes of a bilabiate corolla or calyx, <a href="#Page_92">92</a>.</p> + +<p><i>Litoral</i> or <i>Littoral</i>, belonging to the shore.</p> + +<p><i>Livid</i>, pale lead-colored.</p> + +<p><i>Lobe</i>, any projection or division (especially a rounded one) of a leaf, +&c.</p> + +<p><i>Lobed</i> or <i>Lobate</i>, cut into lobes, <a href="#Page_55">55</a>, <a href="#Page_56">56</a>; <i>Lobulate</i>, into small +lobes.</p> + +<p><i>Locellate</i>, having <i>Locelli</i>, i. e. compartments in a cell: thus an +anther-cell is often <i>bilocellate</i>.</p> + +<p><i>Loculament</i>, same as <i>loculus</i>.</p> + +<p><i>Locular</i>, relating to the cell or compartment (<i>Loculus</i>) of an ovary, +&c.</p> + +<p><i>Loculicidal</i> (dehiscence), splitting down through the back of each +cell, <a href="#Page_123">123</a>.</p> + +<p><i>Locusta</i>, a name for the spikelet of Grasses.</p> + +<p><i>Lodicule</i>, one of the scales answering to perianth-leaves in +Grass-flowers.</p> + +<p><i>Loment</i>, a pod which separates transversely into joints, <a href="#Page_122">122</a>.</p> + +<p><i>Lomentaceous</i>, pertaining to or resembling a loment.</p> + +<p><i>Lorate</i>, thong-shaped.</p> + +<p><i>Lunate</i>, crescent-shaped. <i>Lunulate</i>, diminutive of <i>lunate</i>.</p> + +<p><i>Lupuline</i>, like hops.</p> + +<p><i>Lusus</i>, Latin for a sport or abnormal variation.</p> + +<p><i>Luteolus</i>, yellowish; diminutive of</p> + +<p><i>Luteus</i>, Latin for yellow. <i>Lutescent</i>, verging to yellow.</p> + +<p><i>Lyrate</i>, lyre-shaped; a pinnatifid leaf of an obovate or spatulate +outline, the end-lobe large and roundish, and the lower lobes small, as +in fig. <a href="#Fig148">149</a>.</p> +</div> + +<div class="index"> +<p><i>Macros</i>, Greek for long, sometimes also used for large: thus +<i>Macrophyllous</i>, long or large-leaved, &c.</p> + +<p><i>Macrospore</i>, the large kind of spore, when there are two kinds, <a href="#Page_160">160</a>, +<a href="#Page_161">161</a>.</p> + +<p><i>Maculate</i>, spotted or blotched.</p> + +<p><i>Male</i> (flowers or plants), having stamens but no pistil.</p> + +<p><i>Mammose</i>, breast-shaped.</p> + +<p><i>Marcescent</i>, withering without falling off.</p> + +<p><i>Marginal</i>, belonging to margin.</p> + +<span class="pagenum"><a name="Page_212" id="Page_212">[Pg 212]</a></span><p><i>Marginate</i>, margined with an edge different from the rest.</p> + +<p><i>Marginicidal dehiscence</i>, <a href="#Page_123">123</a>.</p> + +<p><i>Maritime</i>, belonging to sea-coasts.</p> + +<p><i>Marmorate</i>, marbled.</p> + +<p><i>Mas.</i>, <i>Masc.</i>, <i>Masculine</i>, male.</p> + +<p><i>Masked</i>, see <a href="#Personate"><i>personate</i></a>.</p> + +<p><i>Mealy</i>, see <a href="#Farinaceous"><i>farinaceous</i></a>.</p> + +<p><i>Median</i>, <i>Medial</i>, belonging to the middle.</p> + +<p><i>Medifixed</i>, attached by the middle.</p> + +<p><i>Medullary</i>, belonging to, or of the nature of, pith (<i>Medulla</i>); pithy.</p> + +<p><i>Medullary Rays</i>, the silver-grain of wood, <a href="#Page_140">140</a>, <a href="#Page_141">141</a>.</p> + +<p><i>Medullary Sheath</i>, a set of ducts just around the pith, <a href="#Page_140">140</a>.</p> + +<p><i>Meiostemonous</i>, having fewer stamens than petals.</p> + +<p><i>Membranaceous</i> or <i>Membranous</i>, of the texture of membrane; thin and +soft.</p> + +<p><i>Meniscoid</i>, crescent-shaped.</p> + +<p><i>Mericarp</i>, one carpel of the fruit of an Umbelliferous plant, <a href="#Page_121">121</a>.</p> + +<p><i>Merismatic</i>, separating into parts by the formation of partitions +across.</p> + +<p><i>Merous</i>, from the Greek for part; used with numeral prefix to denote +the number of pieces in a set or circle: as <i>Monomerous</i>, of only one, +<i>Dimerous</i>, with two, <i>Trimerous</i>, with three parts (sepals, petals, +stamens, &c.) in each circle.</p> + +<p><i>Mesocarp</i>, the middle part of a pericarp, when that is distinguishable +into three layers, <a href="#Page_120">120</a>.</p> + +<p><i>Mesophlœum</i>, the middle or green bark.</p> + +<p><i>Micropyle</i>, the closed orifice of the seed, <a href="#Page_110">110</a>, <a href="#Page_126">126</a>.</p> + +<p><i>Microspore</i>, the smaller kind of spore when there are two kinds, <a href="#Page_161">161</a>.</p> + +<p><i>Midrib</i>, the middle or main rib of a leaf, <a href="#Page_50">50</a>.</p> + +<p><i>Milk-vessels</i>, <a href="#Page_135">135</a>.</p> + +<p><i>Miniate</i>, vermilion-colored.</p> + +<p><i>Mitriform</i>, mitre-shaped: in the form of a peaked cap, or one cleft at +the top.</p> + +<p><a name="Moniliform" id="Moniliform"></a><i>Moniliform</i>, necklace-shaped; a cylindrical body contracted at +intervals.</p> + +<p><i>Monocarpic</i> (duration), flowering and seeding but once, <a href="#Page_38">38</a>.</p> + +<p><i>Monochlamydeous</i>, having only one floral envelope.</p> + +<p><i>Monocotyledonous</i> (embryo), with only one cotyledon, <a href="#Page_24">24</a>.</p> + +<p><i>Monocotyledonous Plants</i>, <a href="#Page_24">24</a>. <i>Monocotyls</i>, <a href="#Page_24">24</a>.</p> + +<p><i>Monœcious</i>, or <i>Monoicous</i> (flower), having stamens or pistils only, +<a href="#Page_85">85</a>.</p> + +<p><i>Monogynous</i> (flower), having only one pistil, or one style, <a href="#Page_105">105</a>.</p> + +<p><i>Monopetalous</i> (flower), with the corolla of one piece, <a href="#Page_89">89</a>.</p> + +<p><i>Monophyllous</i>, one-leaved, or of one piece.</p> + +<p><i>Monos</i>, Greek for solitary or only one; thus <a name="Monadelphous" id="Monadelphous"></a><i>Monadelphous</i>, stamens +united by their filaments into one set, <a href="#Page_99">99</a>; <i>Monandrous</i> (flower), +having only one stamen, <a href="#Page_100">100</a>.</p> + +<p><i>Monosepalous</i>, a calyx of one piece; i. e. with the sepals united into +one body.</p> + +<p><i>Monospermous</i>, one-seeded.</p> + +<p><i>Monstrosity</i>, an unnatural deviation from the usual structure or form.</p> + +<p><i>Morphology</i>, <i>Morphological Botany</i>, <a href="#Page_9">9</a>; the department of botany which +treats of the forms which an organ may assume.</p> + +<p><i>Moschate</i>, Musk-like in odor.</p> + +<p><i>Movements</i>, <a href="#Page_149">149</a>.</p> + +<p><i>Mucronate</i>, tipped with an abrupt short point (<i>Mucro</i>), <a href="#Page_54">54</a>.</p> + +<p><i>Mucronulate</i>, tipped with a minute abrupt point; a diminutive of the +last.</p> + +<p><i>Multi-</i>, in composition, many; as <i>Multangular</i>, many-angled; +<i>Multicipital</i>, many-headed, &c.; <i>Multifarious</i>, in many rows or ranks; +<i>Multifid</i>, many-cleft; <i>Multilocular</i>, many-celled; <i>Multiserial</i>, in +many rows.</p> + +<p><i>Multiple Fruits</i>, <a href="#Page_118">118</a>, <a href="#Page_124">124</a>.</p> + +<p><i>Muricate</i>, beset with short and hard or prickly points.</p> + +<p><i>Muriform</i>, wall-like; resembling courses of bricks in a wall.</p> + +<p><i>Muticous</i>, pointless, blunt, unarmed.</p> + +<p><i>Mycelium</i>, the spawn of Fungi; i. e. the filaments from which +Mushrooms, &c., originate, <a href="#Page_172">172</a>.</p> +</div> + +<div class="index"> +<span class="pagenum"><a name="Page_213" id="Page_213">[Pg 213]</a></span><p><i>Naked</i>, wanting some usual covering, as achlamydeous flowers, <a href="#Page_86">86</a>; +gymnospermous seeds, <a href="#Page_109">109</a>, <a href="#Page_125">125</a>, &c.</p> + +<p><i>Names</i> in botany, <a href="#Page_179">179</a>.</p> + +<p><i>Nanus</i>, Latin for dwarf.</p> + +<p><i>Napiform</i>, turnip-shaped, <a href="#Page_35">35</a>.</p> + +<p><i>Natural System</i>, <a href="#Page_182">182</a>.</p> + +<p><i>Naturalized</i>, introduced from a foreign country, and flourishing wild.</p> + +<p><i>Navicular</i>, boat-shaped, like the glumes of most Grasses.</p> + +<p><i>Necklace-shaped</i>, looking like a string of beads; see <a href="#Moniliform"><i>moniliform</i></a>.</p> + +<p><i>Nectar</i>, the sweet secretion in flowers from which bees make honey, &c.</p> + +<p><i>Nectariferous</i>, honey-bearing; or having a nectary.</p> + +<p><i>Nectary</i>, the old name for petals and other parts of the flower when of +unusual shape, especially when honey-bearing. So the hollow spur-shaped +petals of Columbine were called nectaries; also the curious long-clawed +petals of Monkshood, <a href="#Page_87">87</a>, &c.</p> + +<p><i>Needle-shaped</i>, long, slender, and rigid, like the leaves of Pines.</p> + +<p><i>Nemorose</i> or <i>Nemoral</i>, inhabiting groves.</p> + +<p><i>Nerve</i>, a name for the ribs or veins of leaves when simple and +parallel, <a href="#Page_50">50</a>.</p> + +<p><i>Nerved</i>, furnished with nerves, or simple and parallel ribs or veins, +<a href="#Page_50">50</a>.</p> + +<p><i>Nervose</i>, conspicuously nerved. <i>Nervulose</i>, minutely nervose.</p> + +<p><i>Netted-veined</i>, furnished with branching veins forming network, <a href="#Page_50">50</a>, <a href="#Page_51">51</a>.</p> + +<p><i>Neuter</i>, <i>Neutral</i>, sexless. <i>Neutral flower</i>, <a href="#Page_79">79</a>.</p> + +<p><i>Niger</i>, Latin for black. <i>Nigricans</i>, Latin for verging to black.</p> + +<p><i>Nitid</i>, shining.</p> + +<p><i>Nival</i>, living in or near snow. <i>Niveus</i>, snow-white.</p> + +<p><i>Nodding</i>, bending so that the summit hangs downward.</p> + +<p><i>Node</i>, a knot; the "joints" of a stem, or the part whence a leaf or a +pair of leaves springs, <a href="#Page_13">13</a>.</p> + +<p><i>Nodose</i>, knotty or knobby. <i>Nodulose</i>, furnished with little knobs or +knots.</p> + +<p><i>Nomenclature</i>, <a href="#Page_175">175</a>, <a href="#Page_179">179</a>.</p> + +<p><i>Normal</i>, according to rule, natural.</p> + +<p><i>Notate</i>, marked with spots or lines of a different color.</p> + +<p><i>Nucamentaceous</i>, relating to or resembling a small nut.</p> + +<p><i>Nuciform</i>, nut-shaped or nut-like.</p> + +<p><i>Nucleus</i>, the kernel of an ovule (<a href="#Page_110">110</a>) or seed (<a href="#Page_127">127</a>) of a cell.</p> + +<p><i>Nucule</i>, same as nutlet.</p> + +<p><i>Nude</i>, (Latin, <i>Nudus</i>), naked. So <i>Nudicaulis</i>, naked-stemmed, &c.</p> + +<p><i>Nut</i>, Latin <i>Nux</i>, a hard, mostly one-seeded indehiscent fruit; as a +chestnut, butternut, acorn, <a href="#Page_121">121</a>.</p> + +<p><i>Nutant</i>, nodding.</p> + +<p><i>Nutlet</i>, a little nut; or the stone of a drupe.</p> +</div> + +<div class="index"> +<p><i>Ob-</i> (meaning over against), when prefixed to words signifies +inversion; as, <i>Obcompressed</i>, flattened the opposite of the usual way; +<i>Obcordate</i>, heart-shaped, with the broad and notched end at the apex +instead of the base, <a href="#Page_54">54</a>; <i>Oblanceolate</i>, lance-shaped with the tapering +point downwards, <a href="#Page_52">52</a>.</p> + +<p><i>Oblique</i>, applied to leaves, &c., means unequal-sided.</p> + +<p><i>Oblong</i>, from two to four times as long as broad, <a href="#Page_52">52</a>.</p> + +<p><i>Obovate</i>, inversely ovate, the broad end upward, <a href="#Page_53">53</a>. <i>Obovoid</i>, solid +obovate.</p> + +<p><i>Obtuse</i>, blunt or round at the end, <a href="#Page_54">54</a>.</p> + +<p><i>Obverse</i>, same as <i>inverse</i>.</p> + +<p><i>Obvolute</i> (in the bud), when the margins of one piece or leaf +alternately overlap those of the opposite one.</p> + +<p><i>Ocellate</i>, with a circular colored patch, like an eye.</p> + +<p><i>Ochroleucous</i>, yellowish-white; dull cream-color.</p> + +<p><i>Ocreate</i>, furnished with <i>Ocreæ</i> (boots), or stipules in the form of +sheaths, <a href="#Page_67">67</a>.</p> + +<p><i>Octo-</i>, Latin for eight, enters into the composition of <i>Octagynous</i>, +with eight pistils or styles; <i>Octamerous</i>, its parts in eights; +<i>Octandrous</i>, with eight stamens, &c.</p> + +<span class="pagenum"><a name="Page_214" id="Page_214">[Pg 214]</a></span><p><i>Oculate</i>, with eye-shaped marking.</p> + +<p><i>Officinal</i>, used in medicine, therefore kept in the shops.</p> + +<p><i>Offset</i>, short branches next the ground which take root, <a href="#Page_40">40</a>.</p> + +<p><i>Oides</i>, termination, from the Greek, to denote likeness; so +<i>Dianthoides</i>, Pink-like.</p> + +<p><i>Oleraceous</i>, esculent, as a pot-herb.</p> + +<p><i>Oligos</i>, Greek for few; thus <i>Oliganthous</i>, few-flowered, &c.</p> + +<p><i>Olivaceous</i>, olive-green.</p> + +<p><i>Oophoridium</i>, a name for spore-case containing macrospores.</p> + +<p><i>Opaque</i>, applied to a surface, means dull, not shining.</p> + +<p><i>Operculate</i>, furnished with a lid (<a name="Operculum" id="Operculum"></a><i>Operculum</i>), as the spore-case of +Mosses, <a href="#Page_163">163</a>.</p> + +<p><i>Opposite</i>, said of leaves and branches when on opposite sides of the +stem from each other (i. e. in pairs), <a href="#Page_29">29</a>, <a href="#Page_68">68</a>. Stamens are opposite the +petals, &c., when they stand before them.</p> + +<p><i>Oppositifolius</i>, situated opposite a leaf.</p> + +<p><i>Orbicular</i>, <i>Orbiculate</i>, circular in outline, or nearly so, <a href="#Page_52">52</a>.</p> + +<p><i>Order</i>, group below class, <a href="#Page_178">178</a>. <i>Ordinal names</i>, <a href="#Page_180">180</a>.</p> + +<p><i>Organ</i>, any member of the plant, as a leaf, a stamen, &c.</p> + +<p><i>Organography</i>, study of organs, <a href="#Page_9">9</a>. <i>Organogenesis</i>, that of the +development of organs.</p> + +<p><i>Orgyalis</i>, of the height of a man.</p> + +<p><i>Orthos</i>, Greek for straight; thus, <i>Orthocarpous</i>, with straight fruit; +<i>Orthostichous</i>, straight-ranked.</p> + +<p><i>Orthotropous</i> (ovule or seed), <a href="#Page_111">111</a>.</p> + +<p><i>Osseous</i>, of a bony texture.</p> + +<p><i>Outgrowths</i>, growths from the surface of a leaf, petal, &c.</p> + +<p><i>Oval</i>, broadly elliptical, <a href="#Page_52">52</a>.</p> + +<p><i>Ovary</i>, that part of the pistil containing the ovules or future seeds, +<a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_105">105</a>.</p> + +<p><i>Ovate</i>, shaped like an egg, with the broader end downwards; or, in +plain surfaces, such as leaves, like the section of an egg lengthwise, +<a href="#Page_52">52</a>.</p> + +<p><i>Ovoid</i>, ovate or oval in a solid form.</p> + +<p><i>Ovule</i>, the body which is destined to become a seed, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_105">105</a>, <a href="#Page_110">110</a>.</p> + +<p><i>Ovuliferous</i>, ovule-bearing.</p> +</div> + +<div class="index"> +<p><i>Palate</i>, a projection of the lower lip of a labiate corolla into the +throat, as in Snapdragon, &c.</p> + +<p><i>Palea</i> (plural <i>paleæ</i>), chaff; the inner husks of Grasses; the chaff +or bracts on the receptacle of many Compositæ, as Coreopsis, and +Sunflower.</p> + +<p><i>Paleaceous</i>, furnished with chaff, or chaffy in texture.</p> + +<p><i>Paleolate</i>, having <i>Paleolæ</i> or paleæ of a second order, or narrow +paleæ.</p> + +<p><i>Palet</i>, English term for palea.</p> + +<p><i>Palmate</i>, when leaflets or the divisions of a leaf all spread from the +apex of the petiole, like the hand with the outspread fingers, <a href="#Page_57">57</a>, <a href="#Page_58">58</a>.</p> + +<p><i>Palmately</i> (veined, lobed, &c.), in a palmate manner, <a href="#Page_51">51</a>, <a href="#Page_56">56</a>.</p> + +<p><i>Palmatifid</i>, <i>-lobed</i>, <i>-sect</i>, palmately cleft, or lobed, or divided.</p> + +<p><i>Paludose</i>, inhabiting marshes. <i>Palustrine</i>, same.</p> + +<p><i>Panduriform</i>, or <i>Pandurate</i>, <a href="#Fiddleshaped">fiddle-shaped</a> (which see).</p> + +<p><i>Panicle</i>, an open and branched cluster, <a href="#Page_81">81</a>.</p> + +<p><i>Panicled</i>, <i>Paniculate</i>, arranged in panicles, or like a panicle.</p> + +<p><i>Pannose</i>, covered with a felt of woolly hairs.</p> + +<p><i>Papery</i>, of about the consistence of letter-paper.</p> + +<p><i>Papilionaceous</i>, butterfly-shaped; applied to such a corolla as that of +the Pea, <a href="#Page_91">91</a>.</p> + +<p><i>Papilla</i> (plural <i>papillæ</i>), little nipple-shaped protuberances.</p> + +<p><i>Papillate</i>, <i>Papillose</i>, covered with papillæ.</p> + +<p><i>Pappus</i>, thistle-down. The down crowning the achenium of the Thistle, +Groundsel, &c., and whatever in Compositæ answers to calyx, whether +hairs, teeth, or scales, <a href="#Page_121">121</a>.</p> + +<p><i>Papyraceous</i>, like parchment in texture.</p> + +<p><i>Parallel-veined</i> or <i>nerved</i> (leaves), <a href="#Page_50">50</a>.</p> + +<span class="pagenum"><a name="Page_215" id="Page_215">[Pg 215]</a></span><p><i>Paraphyses</i>, jointed filaments mixed with the antheridia of Mosses.</p> + +<p><i>Parasitic</i>, living as a parasite, i. e. on another plant or animal, <a href="#Page_37">37</a>.</p> + +<p><i>Parenchemytous</i>, composed of parenchyma.</p> + +<p><i>Parenchyma</i>, soft cellular tissue of plants, like the green pulp of +leaves, <a href="#Page_132">132</a>.</p> + +<p><i>Parietal</i> (placentæ, &c.), attached to the walls (<i>parietes</i>) of the +ovary.</p> + +<p><i>Paripinnate</i>, pinnate with an even number of leaflets.</p> + +<p><i>Parted</i>, separated or cleft into parts almost to the base, <a href="#Page_55">55</a>.</p> + +<p><i>Parthenogenesis</i>, producing seed without fertilization.</p> + +<p><i>Partial involucre</i>, same as an <i>involucel</i>; <i>partial petiole</i>, a +division of a main leaf-stalk or the stalk of a leaflet; <i>partial +peduncle</i>, a branch of a peduncle; <i>partial umbel</i>, an umbellet, <a href="#Page_76">76</a>.</p> + +<p><i>Partition</i>, a segment of a <i>parted</i> leaf; or an internal wall in an +ovary, anther, &c.</p> + +<p><i>Patelliform</i>, disk-shaped, like the <i>patella</i> or kneepan.</p> + +<p><i>Patent</i>, spreading, open. <i>Patulous</i>, moderately spreading.</p> + +<p><i>Pauci-</i>, in composition, few; as <i>pauciflorous</i>, few-flowered, &c.</p> + +<p><i>Pear-shaped</i>, solid obovate, the shape of a pear.</p> + +<p><i>Pectinate</i>, pinnatifid or pinnately divided into narrow and close +divisions, like the teeth of a comb.</p> + +<p><i>Pedate</i>, like a bird's foot; palmate or palmately cleft, with the side +divisions again cleft, as in Viola pedata, &c.</p> + +<p><i>Pedicel</i>, the stalk of each particular flower of a cluster, <a href="#Page_73">73</a>.</p> + +<p><i>Pedicellate</i>, <i>Pedicelled</i>, borne on a pedicel.</p> + +<p><i>Pedalis</i>, Latin for a foot high or long.</p> + +<p><i>Peduncle</i>, a flower-stalk, whether of a single flower or of a +flower-cluster, <a href="#Page_73">73</a>.</p> + +<p><i>Peduncled</i>, <i>Pedunculate</i>, furnished with a peduncle.</p> + +<p><i>Peloria</i>, an abnormal return to regularity and symmetry in an irregular +flower; commonest in Snapdragon.</p> + +<p><i>Peltate</i>, shield-shaped; said of a leaf, whatever its shape, when the +petiole is attached to the lower side, somewhere within the margin, <a href="#Page_53">53</a>.</p> + +<p><i>Pelviform</i>, basin-shaped.</p> + +<p><i>Pendent</i>, hanging. <i>Pendulous</i>, somewhat hanging or drooping.</p> + +<p><i>Penicillate</i>, <i>Penicilliform</i>, tipped with a tuft of fine hairs, like a +painter's pencil; as the stigmas of some Grasses.</p> + +<p><i>Pennate</i>, same as pinnate. <i>Penninerved</i> and <i>Penniveined</i>, pinnately +veined, <a href="#Page_51">51</a>.</p> + +<p><i>Penta-</i> (in words of Greek composition), five; as <i>Pentadelphous</i>, <a href="#Page_99">99</a>; +<i>Pentagynous</i>, with five pistils or styles; <i>Pentamerous</i>, with its +parts in fives, or on the plan of five; <i>Pentandrous</i>, having five +stamens, <a href="#Page_112">112</a>; <i>Pentastichous</i>, in five ranks, &c.</p> + +<p><i>Pepo</i>, a fruit like the Melon and Cucumber, <a href="#Page_119">119</a>.</p> + +<p><i>Perennial</i>, lasting from year to year, <a href="#Page_38">38</a>.</p> + +<p><i>Perfect</i> (flower), having both stamens and pistils, <a href="#Page_81">81</a>.</p> + +<p><i>Perfoliate</i>, passing through the leaf, in appearance, <a href="#Page_60">60</a>.</p> + +<p><i>Perforate</i>, pierced with holes, or with transparent dots resembling +holes, as an Orange-leaf.</p> + +<p><i>Peri-</i>, Greek for around; from which are such terms as</p> + +<p><i>Perianth</i>, the leaves of the flower collectively, <a href="#Page_79">79</a>.</p> + +<p><i>Pericarp</i>, the ripened ovary; the walls of the fruit, <a href="#Page_117">117</a>.</p> + +<p><i>Pericarpic</i>, belonging to the pericarp.</p> + +<p><i>Perigonium</i>, <i>Perigone</i>, same as <i>perianth</i>.</p> + +<p><i>Perigynium</i>, bodies around the pistil; applied to the closed cup or +bottle-shaped body (of bracts) which encloses the ovary of Sedges, and +to the bristles, little scales, &c., of the flowers of some other +Cyperaceæ.</p> + +<p><i>Perigynous</i>, the petals and stamens borne on the calyx, <a href="#Page_95">95</a>, <a href="#Page_99">99</a>.</p> + +<p><i>Peripheric</i>, around the outside, or periphery, of any organ.</p> + +<p><i>Perisperm</i>, a name for the albumen of a seed.</p> + +<p><i>Peristome</i>, the fringe of teeth to the spore-case of Mosses, <a href="#Page_163">163</a>.</p> + +<p><i>Persistent</i>, remaining beyond the period when such parts commonly fall, +as the leaves of evergreens, and the calyx of such flowers as persist +during the growth of the fruit.</p> + +<span class="pagenum"><a name="Page_216" id="Page_216">[Pg 216]</a></span><p><a name="Personate" id="Personate"></a><i>Personate</i>, masked; a bilabiate corolla with a <i>palate</i> in the throat, +<a href="#Page_92">92</a>.</p> + +<p><i>Pertuse</i>, perforated with a hole or slit.</p> + +<p><i>Perulate</i>, having scales (<i>Perulæ</i>), such as bud-scales.</p> + +<p><i>Pes</i>, <i>pedis</i>, Latin for the foot or support, whence <i>Longipes</i>, +long-stalked, &c.</p> + +<p><i>Petal</i>, a leaf of the corolla, <a href="#Page_14">14</a>, <a href="#Page_79">79</a>.</p> + +<p><i>Petalody</i>, metamorphosis of stamens, &c., into petals.</p> + +<p><i>Petaloid</i>, <i>Petaline</i>, petal-like; resembling or colored like petals.</p> + +<p><i>Petiole</i>, a footstalk of a leaf; a leaf-stalk, <a href="#Page_49">49</a>.</p> + +<p><i>Petioled</i>, <i>Petiolate</i>, furnished with a petiole.</p> + +<p><i>Petiolulate</i>, said of a leaflet when raised on its own partial +leaf-stalk.</p> + +<p><i>Petræus</i>, Latin for growing on rocks.</p> + +<p><i>Phalanx</i>, <i>phalanges</i>, bundles of stamens.</p> + +<p><i>Phænogamous</i>, or <i>Phanerogamous</i>, plants bearing flowers and producing +seeds; same as Flowering Plants. <i>Phænogams</i>, <i>Phanerogams</i>, <a href="#Page_10">10</a>.</p> + +<p><i>Phlœum</i>, Greek name for bark, whence <i>Endophlœum</i>, inner bark, +&c.</p> + +<p><i>Phœniceous</i>, deep red verging to scarlet.</p> + +<p><i>Phycology</i>, the botany of Algæ.</p> + +<p><i>Phyllocladia</i>, branches assuming the form and function of leaves.</p> + +<p><i>Phyllodium</i> (plural, <i>phyllodia</i>), a leaf where the seeming blade is a +dilated petiole, as in New Holland Acacias, <a href="#Page_61">61</a>.</p> + +<p><i>Phyllome</i>, foliar parts, those answering to leaves in their nature.</p> + +<p><i>Phyllon</i> (plural, <i>phylla</i>), Greek for leaf and leaves; used in many +compound terms and names.</p> + +<p><i>Phyllotaxis</i>, or <i>Phyllotaxy</i>, the arrangement of leaves on the stem, +<a href="#Page_67">67</a>.</p> + +<p><i>Physiological Botany</i>, <a href="#Page_9">9</a>.</p> + +<p><i>Phytography</i>, relates to characterizing and describing plants.</p> + +<p><i>Phyton</i>, or <i>Phytomer</i>, a name used to designate the pieces which by +their repetition make up a plant, theoretically, viz. a joint of stem +with its leaf or pair of leaves.</p> + +<p><i>Pileus</i> of a mushroom, <a href="#Page_172">172</a>.</p> + +<p><i>Piliferous</i>, bearing a slender bristle or hair (<i>pilum</i>), or beset with +hairs.</p> + +<p><i>Pilose</i>, hairy; clothed with soft slender hairs.</p> + +<p><i>Pinna</i>, a primary division with its leaflets of a bipinnate or +tripinnate leaf.</p> + +<p><i>Pinnule</i>, a secondary division of a bipinnate or tripinnate leaf, <a href="#Page_66">66</a>.</p> + +<p><i>Pinnate</i> (leaf), when leaflets are arranged along the sides of a common +petiole, <a href="#Page_57">57</a>.</p> + +<p><i>Pinnately lobed</i>, <i>cleft</i>, <i>parted</i>, <i>divided</i>, <i>veined</i>, <a href="#Page_56">56</a>.</p> + +<p><i>Pinnatifid</i>, <i>Pinnatisect</i>, same as pinnately cleft and pinnately +parted, <a href="#Page_56">56</a>.</p> + +<p><i>Pisiform</i>, pea-shaped.</p> + +<p><i>Pistil</i>, the seed-bearing organ of the flower, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_105">105</a>.</p> + +<p><i>Pistillate</i>, having a pistil, <a href="#Page_85">85</a>.</p> + +<p><i>Pistillidium</i>, the body which in Mosses answers to the pistil, <a href="#Page_159">159</a>, +<a href="#Page_164">164</a>.</p> + +<p><i>Pitchers</i>, <a href="#Page_64">64</a>.</p> + +<p><i>Pith</i>, the cellular centre of an exogenous stem, <a href="#Page_138">138</a>.</p> + +<p><i>Placenta</i>, the surface or part of the ovary to which the ovules are +attached, <a href="#Page_107">107</a>.</p> + +<p><i>Placentiform</i>, nearly same as quoit-shaped.</p> + +<p><i>Plaited</i> (in the bud), or <i>Plicate</i>, folded, <a href="#Page_72">72</a>, <a href="#Page_98">98</a>.</p> + +<p><i>Platy-</i>, Greek for broad, in compounds, such as <i>Platyphyllous</i>, +broad-leaved, &c.</p> + +<p><i>Pleio-</i>, Greek for full or abounding, used in compounds, such as +<i>Pleiopetalous</i>, of many petals, &c.</p> + +<p><i>Plumbeus</i>, lead-colored.</p> + +<p><i>Plumose</i>, feathery; when any slender body (such as a bristle of a +pappus or a style) is beset with hairs along its sides, like the plume +of a feather.</p> + +<p><i>Plumule</i>, the bud or first shoot of a germinating plantlet above the +cotyledons, <a href="#Page_13">13</a>.</p> + +<p><i>Pluri-</i>, in composition, many or several; as <i>Plurifoliolate</i>, with +several leaflets.</p> + +<p><i>Pod</i>, specially a legume, <a href="#Page_122">122</a>; also may be applied to any sort of +capsule.</p> + +<p><i>Podium</i>, a footstalk or stipe, used only in Greek compounds, as +(suffixed) <i>Leptopodus</i>, slender-stalked, or (prefixed) <i>Podocephalus</i>, +with a stalked head, and in <i>Podosperm</i>, a seed stalk or funiculus.</p> + +<p><i>Pogon</i>, Greek for beard, comes into various compounds.</p> + +<span class="pagenum"><a name="Page_217" id="Page_217">[Pg 217]</a></span><p><i>Pointless</i>, destitute of any pointed tip, such as a <i>mucro</i>, <i>awn</i>, +<i>acumination</i>, &c.</p> + +<p><i>Pollen</i>, the fertilizing powder contained in the anther, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_103">103</a>.</p> + +<p><i>Pollen-growth</i>, <a href="#Page_117">117</a>. <i>Polleniferous</i>, pollen-bearing.</p> + +<p><i>Pollen-mass</i>, <i>Pollinium</i>, the united mass of pollen, <a href="#Page_104">104</a>, as in +Milkweed and Orchis.</p> + +<p><i>Pollicaris</i>, Latin for an inch long.</p> + +<p><i>Pollination</i>, the application of pollen to the stigma, <a href="#Page_114">114</a>.</p> + +<p><i>Poly-</i>, in compound words of Greek origin, same as <i>multi-</i> in those of +Latin origin viz. many, as</p> + +<p><i>Polyadelphous</i>, stamens united by their filaments into several bundles, +<a href="#Page_100">100</a>.</p> + +<p><i>Polyandrous</i>, with numerous stamens (inserted on the receptacle), <a href="#Page_100">100</a>.</p> + +<p><i>Polycarpic</i>, term used by DeCandolle in the sense of perennial.</p> + +<p><i>Polycotyledonous</i>, having many (more than two) cotyledons, as Pines, +<a href="#Page_23">23</a>.</p> + +<p><i>Polygamous</i>, having some perfect and some unisexual flowers, <a href="#Page_85">85</a>.</p> + +<p><i>Polygonal</i>, many-angled.</p> + +<p><i>Polygynous</i>, with many pistils or styles, <a href="#Page_105">105</a>.</p> + +<p><i>Polymerous</i>, formed of many parts of each set.</p> + +<p><i>Polymorphous</i>, of several or varying forms.</p> + +<p><i>Polypetalous</i>, when the petals are distinct or separate (whether few or +many), <a href="#Page_89">89</a>.</p> + +<p><i>Polyphyllous</i>, many-leaved; formed of several distinct pieces.</p> + +<p><i>Polysepalous</i>, same as the last when applied to the calyx, <a href="#Page_89">89</a>.</p> + +<p><i>Polyspermous</i>, many-seeded.</p> + +<p><i>Pome</i>, the apple, pear, and similar fleshy fruits, <a href="#Page_119">119</a>.</p> + +<p><i>Pomiferous</i>, pome-bearing.</p> + +<p><i>Porrect</i>, outstretched.</p> + +<p><i>Posterior</i> side or portion of a flower (when axillary) is that toward +the axis, <a href="#Page_96">96</a>.</p> + +<p><i>Pouch</i>, the silicle or short pod, as of Shepherd's Purse, <a href="#Page_123">123</a>.</p> + +<p><i>Præcocious</i> (Latin, <i>præcox</i>), unusually early in development.</p> + +<p><i>Præfloration</i>, same as <i>æstivation</i>, <a href="#Page_97">97</a>.</p> + +<p><i>Præfoliation</i>, same as <i>vernation</i>, <a href="#Page_71">71</a>.</p> + +<p><i>Præmorse</i>, ending abruptly, as if bitten off.</p> + +<p><i>Pratensis</i>, Latin for growing in meadows.</p> + +<p><i>Prickles</i>, sharp elevations of the bark, coming off with it, as of the +Rose.</p> + +<p><i>Prickly</i>, bearing prickles, or sharp projections like them.</p> + +<p><i>Primine</i>, the outer coat of the covering of the ovule, <a href="#Page_110">110</a>.</p> + +<p><i>Primordial</i>, earliest formed; primordial leaves are the first after the +cotyledons.</p> + +<p><i>Prismatic</i>, prism-shaped; having three or more angles bounding flat +sides.</p> + +<p><i>Procerous</i>, tall, or tall and slim.</p> + +<p><i>Process</i>, any projection from the surface or edge of a body.</p> + +<p><i>Procumbent</i>, trailing on the ground, <a href="#Page_39">39</a>.</p> + +<p><i>Procurrent</i>, running through but not projecting.</p> + +<p><i>Produced</i>, extended or projecting; the upper sepal of a Larkspur is +<i>produced</i> above into a spur, <a href="#Page_87">87</a>.</p> + +<p><i>Proliferous</i> (literally, bearing offspring), where a new branch rises +from an older one, or one head or cluster of flowers out of another.</p> + +<p><i>Propaculum</i> or <i>Propagulum</i>, a shoot for propagation.</p> + +<p><i>Prosenchyma</i>, a tissue of wood-cells.</p> + +<p><i>Prostrate</i>, lying flat on the ground, <a href="#Page_39">39</a>.</p> + +<p><i>Protandrous</i> or <i>Proterandrous</i>, the anthers first maturing, <a href="#Page_116">116</a>.</p> + +<p><i>Proteranthous</i>, flowering before leafing.</p> + +<p><i>Proterogynous</i> or <i>Protogynous</i>, the stigmas first to mature, <a href="#Page_116">116</a>.</p> + +<p><i>Prothallium</i> or <i>Prothallus</i>, <a href="#Page_160">160</a>.</p> + +<p><i>Protoplasm</i>, the soft nitrogenous lining or contents, or living part, +of cells, <a href="#Page_129">129</a>.</p> + +<p><i>Protos</i>, Greek for first; in various compounds.</p> + +<p><i>Pruinose</i>, <i>Pruinate</i>, frosted; covered with a powder like hoar-frost.</p> + +<p><i>Pseudo-</i>, Greek for false. <i>Pseudo-bulb</i>, the aerial corms of epiphytic +Orchids, &c.</p> + +<p><i>Psilos</i>, Greek for bare or naked, used in many compounds.</p> + +<p><i>Pteridophyta</i>, <i>Pteridophytes</i>, <a href="#Page_156">156</a>.</p> + +<p><i>Pteris</i>, Greek for wing, and general name for Fern, enters into many +compounds.</p> + +<span class="pagenum"><a name="Page_218" id="Page_218">[Pg 218]</a></span><p><i>Puberulent</i>, covered with fine and short or almost imperceptible down.</p> + +<p><i>Pubescent</i>, hairy or downy, especially with fine and soft hairs or +<i>pubescence</i>.</p> + +<p><i>Pulverulent</i> or <i>Pulveraceous</i>, as if dusted with fine powder.</p> + +<p><i>Pulvinate</i>, cushioned, or shaped like a cushion.</p> + +<p><i>Pumilus</i>, low or little.</p> + +<p><i>Punctate</i>, dotted, either with minute holes or what look as such.</p> + +<p><i>Puncticulate</i>, minutely punctate.</p> + +<p><i>Pungent</i>, prickly-tipped.</p> + +<p><i>Puniceous</i>, carmine-red.</p> + +<p><i>Purpureus</i>, originally red or crimson, more used for duller or +bluish-red.</p> + +<p><i>Pusillus</i>, weak and small, tiny.</p> + +<p><i>Putamen</i>, the stone of a drupe, or the shell of a nut, <a href="#Page_120">120</a>.</p> + +<p><i>Pygmæus</i>, Latin for dwarf.</p> + +<p><i>Pyramidal</i>, shaped like a pyramid.</p> + +<p><i>Pyrene</i>, <i>Pyrena</i>, a seed-like nutlet or stone of a small drupe.</p> + +<p><i>Pyriform</i>, pear-shaped.</p> + +<p><i>Pyxidate</i>, furnished with a lid.</p> + +<p><i>Pyxis</i>, <i>Pyxidium</i>, a pod opening round horizontally by a lid, <a href="#Page_124">124</a>.</p> +</div> + +<div class="index"> +<p><i>Quadri-</i>, in words of Latin origin, four; as <i>Quadrangular</i>, +four-angled; <i>Quadrifoliate</i>, four-leaved; <i>Quadrifid</i>, four-cleft. +<i>Quaternate</i> in fours.</p> + +<p><i>Quinate</i>, in fives. <i>Quinque</i>, five.</p> + +<p><i>Quincuncial</i>, in a quincunx; when the parts in æstivation are five, two +of them outside, two inside, and one half out and half in.</p> + +<p><i>Quintuple</i>, five-fold.</p> +</div> + +<div class="index"> +<p><i>Race</i>, a marked variety which may be perpetuated from seed, <a href="#Page_176">176</a>.</p> + +<p><i>Raceme</i>, a flower-cluster, with one-flowered pedicels arranged along +the sides of a general peduncle, <a href="#Page_73">73</a>.</p> + +<p><i>Racemose</i>, bearing racemes, or raceme-like.</p> + +<p><i>Rachis</i>, see <a href="#Rhachis"><i>rhachis</i></a>.</p> + +<p><i>Radial</i>, belonging to the ray.</p> + +<p><i>Radiate</i>, or <i>Radiant</i>, furnished with ray-flowers, <a href="#Page_94">94</a>.</p> + +<p><i>Radiate-veined</i>, <a href="#Page_52">52</a>.</p> + +<p><i>Radical</i>, belonging to the root, or apparently coming from the root.</p> + +<p><i>Radicant</i>, rooting, taking root on or above the ground.</p> + +<p><i>Radicels</i>, little roots or rootlets.</p> + +<p><i>Radicle</i>, the stem part of the embryo, the lower end of which forms the +root, <a href="#Page_11">11</a>, <a href="#Page_127">127</a>.</p> + +<p><i>Rameal</i>, belonging to a branch. <i>Ramose</i>, full of branches (<i>rami</i>).</p> + +<p><i>Ramentaceous</i>, beset with thin chaffy scales (<i>Ramenta</i>), as the stalks +of many Ferns.</p> + +<p><i>Ramification</i>, branching, <a href="#Page_27">27</a>.</p> + +<p><i>Ramulose</i>, full of branchlets (<i>ramuli</i>).</p> + +<p><i>Raphe</i>, see <a href="#Rhaphe"><i>rhaphe</i></a>.</p> + +<p><i>Ray</i>, parts diverging from a centre, the marginal flowers of a head (as +of Coreopsis, <a href="#Page_94">94</a>), or cluster, as of Hydrangea (<a href="#Page_78">78</a>), when different from +the rest, especially when ligulate and diverging (like rays or +sunbeams); also the branches of an umbel, <a href="#Page_74">74</a>.</p> + +<p><i>Ray-flowers</i>, <a href="#Page_94">94</a>.</p> + +<p><i>Receptacle</i>, the axis or support of a flower, <a href="#Page_81">81</a>, <a href="#Page_112">112</a>; also the common +axis or support of a head of flowers, <a href="#Page_73">73</a>.</p> + +<p><i>Reclined</i>, turned or curved downwards; nearly recumbent.</p> + +<p><i>Rectinerved</i>, with straight nerves or veins.</p> + +<p><i>Recurved</i>, curved outwards or backwards.</p> + +<p><i>Reduplicate</i> (in æstivation), valvate with the margins turned outwards, +<a href="#Page_97">97</a>.</p> + +<p><i>Reflexed</i>, bent outwards or backwards.</p> + +<p><i>Refracted</i>, bent suddenly, so as to appear broken at the bend.</p> + +<p><i>Regular</i>, all the parts similar in shape, <a href="#Page_82">82</a>.</p> + +<p><i>Reniform</i>, kidney-shaped, <a href="#Page_53">53</a>.</p> + +<span class="pagenum"><a name="Page_219" id="Page_219">[Pg 219]</a></span><p><i>Repand</i>, wavy-margined, <a href="#Page_55">55</a>.</p> + +<p><i>Repent</i>, creeping, i. e. prostrate and rooting underneath.</p> + +<p><i>Replum</i>, the frame of some pods (as of Prickly Poppy and Cress), +persistent after the valves fall away.</p> + +<p><i>Reptant</i>, same as repent.</p> + +<p><i>Resupinate</i>, inverted, or appearing as if upside down, or reversed.</p> + +<p><i>Reticulated</i>, the veins forming network, <a href="#Page_50">50</a>. <i>Retiform</i>, in network.</p> + +<p><i>Retinerved</i>, reticulate-veined.</p> + +<p><i>Retroflexed</i>, bent backwards; same as <i>reflexed</i>.</p> + +<p><i>Retuse</i>, blunted; the apex not only obtuse but somewhat indented, <a href="#Page_54">54</a>.</p> + +<p><i>Revolute</i>, rolled backwards, as the margins of many leaves, <a href="#Page_72">72</a>.</p> + +<p><a name="Rhachis" id="Rhachis"></a><i>Rhachis</i> (the backbone), the axis of a spike or other body, <a href="#Page_73">73</a>.</p> + +<p><a name="Rhaphe" id="Rhaphe"></a><i>Rhaphe</i>, the continuation of the seed-stalk along the side of an +anatropous ovule or seed, <a href="#Page_112">112</a>, <a href="#Page_126">126</a>.</p> + +<p><i>Rhaphides</i>, crystals, especially needle-shaped ones, in the tissues of +plants, <a href="#Page_137">137</a>.</p> + +<p><i>Rhizanthous</i>, flowering from the root.</p> + +<p><i>Rhizoma</i>, <i>Rhizome</i>, a rootstock, <a href="#Page_42">42</a>-<a href="#Page_44">44</a>.</p> + +<p><i>Rhombic</i>, in the shape of a rhomb. <i>Rhomboidal</i>, approaching that +shape.</p> + +<p><i>Rib</i>, the principal piece, or one of the principal pieces of the +framework of a leaf, or any similar elevated line along a body, <a href="#Page_49">49</a>, <a href="#Page_50">50</a>.</p> + +<p><i>Rimose</i>, having chinks or cracks.</p> + +<p><i>Ring</i>, an elastic band on the spore-cases of Ferns, <a href="#Page_159">159</a>.</p> + +<p><i>Ringent</i>, grinning; gaping open, <a href="#Page_92">92</a>.</p> + +<p><i>Riparious</i>, on river-banks.</p> + +<p><i>Rivalis</i>, Latin for growing along brooks; or <i>Rivularis</i>, in rivulets.</p> + +<p><i>Root</i>, <a href="#Page_33">33</a>.</p> + +<p><i>Root-hairs</i>, <a href="#Page_35">35</a>.</p> + +<p><i>Rootlets</i>, small roots, or root-branches, <a href="#Page_33">33</a>.</p> + +<p><i>Rootstock</i>, root-like trunks or portions of stems on or under ground, +<a href="#Page_42">42</a>.</p> + +<p><i>Roridus</i>, dewy.</p> + +<p><i>Rosaceous</i>, arranged like the petals of a rose.</p> + +<p><i>Rostellate</i>, bearing a small beak (<i>Rostellum</i>).</p> + +<p><i>Rostrate</i>, bearing a beak (<i>Rostrum</i>) or a prolonged appendage.</p> + +<p><i>Rosulate</i>, in a rosette or cluster of spreading leaves.</p> + +<p><i>Rotate</i>, wheel-shaped, <a href="#Page_89">89</a>.</p> + +<p><i>Rotund</i>, rounded or roundish in outline.</p> + +<p><i>Ruber</i>, Latin for red in general. <i>Rubescent</i>, <i>Rubicund</i>, reddish or +blushing.</p> + +<p><i>Rudimentary</i>, imperfectly developed, or in an early state of +development.</p> + +<p><i>Rufous</i>, <i>Rufescent</i>, brownish-red or reddish-brown.</p> + +<p><i>Rugose</i>, wrinkled; roughened with wrinkles.</p> + +<p><i>Ruminated</i> (albumen), penetrated with irregular channels or portions, +as a nutmeg, looking as if chewed.</p> + +<p><i>Runcinate</i>, coarsely saw-toothed or cut, the pointed teeth turned +towards the base of the leaf, as the leaf of a Dandelion.</p> + +<p><i>Runner</i>, a slender and prostrate branch, rooting at the end, or at the +joints, <a href="#Page_40">40</a>.</p> +</div> + +<div class="index"> +<p><i>Sabulose</i>, growing in sand.</p> + +<p><i>Sac</i>, any closed membrane, or a deep purse-shaped cavity.</p> + +<p><i>Saccate</i>, sac-shaped.</p> + +<p><i>Sagittate</i>, arrowhead-shaped, <a href="#Page_53">53</a>.</p> + +<p><i>Salsuginous</i>, growing in brackish soil.</p> + +<p><i>Salver-shaped</i>, or <i>Salver-form</i>, with a border spreading at right +angles to a slender tube, <a href="#Page_89">89</a>.</p> + +<p><i>Samara</i>, a wing-fruit, or key, <a href="#Page_122">122</a>.</p> + +<p><i>Samaroid</i>, like a samara or key-fruit.</p> + +<p><i>Sap</i>, the juices of plants generally, <a href="#Page_136">136</a>. <i>Sapwood</i>, <a href="#Page_142">142</a>.</p> + +<p><i>Saprophytes</i>, <a href="#Page_37">37</a>.</p> + +<p><i>Sarcocarp</i>, the fleshy part of a stone-fruit, <a href="#Page_120">120</a>.</p> + +<span class="pagenum"><a name="Page_220" id="Page_220">[Pg 220]</a></span><p><i>Sarmentaceous</i>, <i>Sarmentose</i>, bearing long and flexible twigs +(<i>Sarments</i>), either spreading or procumbent.</p> + +<p><i>Saw-toothed</i>, see <a href="#Serrate"><i>serrate</i></a>, <a href="#Page_55">55</a>.</p> + +<p><i>Scabrous</i>, rough or harsh to the touch.</p> + +<p><i>Scalariform</i>, with cross-bands, resembling the steps of a ladder, <a href="#Page_134">134</a>.</p> + +<p><i>Scales</i>, of buds, <a href="#Page_28">28</a>; of bulbs, &c., <a href="#Page_46">46</a>.</p> + +<p><i>Scalloped</i>, same as <i>crenate</i>, <a href="#Page_55">55</a>.</p> + +<p><i>Scaly</i>, furnished with scales, or scale-like in texture.</p> + +<p><i>Scandent</i>, climbing, <a href="#Page_39">39</a>.</p> + +<p><i>Scape</i>, a peduncle rising from the ground or near it, as in many +Violets.</p> + +<p><i>Scapiform</i>, scape-like.</p> + +<p><i>Scapigerous</i>, scape-bearing.</p> + +<p><i>Scar</i> of the seed, <a href="#Page_126">126</a>. <i>Leaf-scars</i>, <a href="#Page_27">27</a>, <a href="#Page_28">28</a>.</p> + +<p><i>Scarious</i> or <i>Scariose</i>, thin, dry, and membranous.</p> + +<p><i>Scion</i>, a shoot or slip used for grafting.</p> + +<p><i>Scleros</i>, Greek for hard, hence <i>Sclerocarpous</i>, hard-fruited.</p> + +<p><i>Scobiform</i>, resembling sawdust.</p> + +<p><i>Scorpioid</i> or <i>Scorpioidal</i>, curved or circinate at the end, <a href="#Page_77">77</a>.</p> + +<p><i>Scrobiculate</i>, pitted; excavated into shallow pits.</p> + +<p><i>Scurf</i>, <i>Scurfiness</i>, minute scales on the surface of many leaves, as +of Goosefoot.</p> + +<p><i>Scutate</i>, <i>Scutiform</i>, buckler-shaped.</p> + +<p><i>Scutellate</i>, or <i>Scutelliform</i>, saucer-shaped or platter-shaped.</p> + +<p><i>Secund</i>, one-sided; i. e. where flowers, leaves, &c., are all turned to +one side.</p> + +<p><i>Secundine</i>, the inner coat of the ovule, <a href="#Page_110">110</a>.</p> + +<p><i>Seed</i>, <a href="#Page_125">125</a>. <i>Seed-leaves</i>, see <a href="#Cotyledons"><i>cotyledons</i></a>. <i>Seed-vessel</i>, <a href="#Page_127">127</a>.</p> + +<p><i>Segment</i>, a subdivision or lobe of any cleft body.</p> + +<p><i>Segregate</i>, separated from each other.</p> + +<p><i>Semi-</i>, in compound words of Latin origin, half; as</p> + +<p><i>Semi-adherent</i>, as the calyx or ovary of Purslane; <i>Semicordate</i>, +half-heart-shaped; <i>Semilunar</i>, like a half-moon; <i>Semiovate</i>, +half-ovate, &c.</p> + +<p><i>Seminal</i>, relating to the seed (<i>Semen</i>). <i>Seminiferous</i>, seed-bearing.</p> + +<p><i>Sempervirent</i>, evergreen.</p> + +<p><i>Sensitiveness</i> in plants, <a href="#Page_149">149</a>, <a href="#Page_152">152</a>.</p> + +<p><i>Senary</i>, in sixes.</p> + +<p><i>Sepal</i>, a leaf or division of the calyx, <a href="#Page_14">14</a>, <a href="#Page_79">79</a>.</p> + +<p><i>Sepaloid</i>, sepal-like. <i>Sepaline</i>, relating to the sepals.</p> + +<p><i>Separated Flowers</i>, those having stamens or pistils only, <a href="#Page_85">85</a>.</p> + +<p><i>Septate</i>, divided by partitions.</p> + +<p><i>Septenate</i>, with parts in sevens.</p> + +<p><i>Septicidal</i>, where dehiscence is through the partitions, <a href="#Page_123">123</a>.</p> + +<p><i>Septiferous</i>, bearing the partition.</p> + +<p><i>Septifragal</i>, where the valves in dehiscence break away from the +partitions, <a href="#Page_123">123</a>.</p> + +<p><i>Septum</i> (plural <i>septa</i>), a partition or dissepiment.</p> + +<p><i>Serial</i>, or <i>Seriate</i>, in rows; as <i>biserial</i>, in two rows, &c.</p> + +<p><i>Sericeous</i>, silky; clothed with satiny pubescence.</p> + +<p><i>Serotinous</i>, late in the season.</p> + +<p><a name="Serrate" id="Serrate"></a><i>Serrate</i>, the margin cut into teeth (<i>Serratures</i>) pointing forwards, +<a href="#Page_55">55</a>.</p> + +<p><i>Serrulate</i>, same as the last, but with fine teeth.</p> + +<p><i>Sessile</i>, sitting; without any stalk.</p> + +<p><i>Sesqui-</i>, Latin for one and a half; so <i>Sesquipedalis</i>, a foot and a +half long.</p> + +<p><i>Seta</i>, a bristle, or a slender body or appendage resembling a bristle.</p> + +<p><i>Setaceous</i>, bristle-like. <i>Setiform</i>, bristle-shaped.</p> + +<p><i>Setigerous</i>, bearing bristles. <i>Setose</i>, beset with bristles or bristly +hairs.</p> + +<p><i>Setula</i>, a diminutive bristle. <i>Setulose</i>, provided with such.</p> + +<p><i>Sex</i>, six. <i>Sexangular</i>, six-angled. <i>Sexfarious</i>, six-faced.</p> + +<p><i>Sheath</i>, the base of such leaves as those of Grasses, which are</p> + +<p><i>Sheathing</i>, wrapped round the stem.</p> + +<p><i>Shield-shaped</i>, same as <i>scutate</i>, or as <i>peltate</i>, <a href="#Page_53">53</a>.</p> + +<span class="pagenum"><a name="Page_221" id="Page_221">[Pg 221]</a></span><p><i>Shrub</i>, <i>Shrubby</i>, <a href="#Page_39">39</a>.</p> + +<p><i>Sieve-cells</i>, <a href="#Page_140">140</a>.</p> + +<p><i>Sigmoid</i>, curved in two directions, like the letter S, or the Greek +<i>sigma</i>.</p> + +<p><i>Silicle</i>, a pouch, or short pod of the Cress Family, <a href="#Page_123">123</a>.</p> + +<p><i>Siliculose</i>, bearing a silicle, or a fruit resembling it.</p> + +<p><i>Silique</i>, capsule of the Cress Family, <a href="#Page_123">123</a>.</p> + +<p><i>Siliquose</i>, bearing siliques or pods which resemble siliques.</p> + +<p><i>Silky</i>, glossy with a coat of fine and soft, close-pressed, straight +hairs.</p> + +<p><i>Silver-grain</i>, the medullary rays of wood, <a href="#Page_139">139</a>.</p> + +<p><i>Silvery</i>, shining white or bluish-gray, usually from a silky +pubescence.</p> + +<p><i>Simple</i>, of one piece; opposed to <i>compound</i>.</p> + +<p><i>Sinistrorse</i>, turned to the left.</p> + +<p><i>Sinuate</i>, with margin alternately bowed inwards and outwards, <a href="#Page_55">55</a>.</p> + +<p><i>Sinus</i>, a recess or bay; the re-entering angle between two lobes or +projections.</p> + +<p><i>Sleep of Plants</i> (so called), <a href="#Page_151">151</a>.</p> + +<p><i>Smooth</i>, properly speaking not rough, but often used for glabrous, i. +e. not pubescent.</p> + +<p><i>Soboliferous</i>, bearing shoots (<i>Soboles</i>) from near the ground.</p> + +<p><i>Solitary</i>, single; not associated with others.</p> + +<p><i>Sordid</i>, dull or dirty in hue.</p> + +<p><i>Sorediate</i>, bearing patches on the surface.</p> + +<p><i>Sorosis</i>, name of a multiple fruit, like a pine-apple.</p> + +<p><a name="Sorus" id="Sorus"></a><i>Sorus</i>, a fruit-dot of Ferns, <a href="#Page_159">159</a>.</p> + +<p><i>Spadiceous</i>, chestnut-colored. Also spadix-bearing.</p> + +<p><i>Spadix</i>, a fleshy spike of flowers, <a href="#Page_75">75</a>.</p> + +<p><i>Span</i>, the distance between the tip of the thumb and of little finger +outstretched, six or seven inches.</p> + +<p><i>Spathaceous</i>, resembling or furnished with a</p> + +<p><i>Spathe</i>, a bract which inwraps an inflorescence, <a href="#Page_75">75</a>.</p> + +<p><i>Spatulate</i>, or <i>Spathulate</i>, shaped like a spatula, <a href="#Page_52">52</a>.</p> + +<p><i>Species</i>, <a href="#Page_175">175</a>.</p> + +<p><i>Specific Names</i>, <a href="#Page_179">179</a>.</p> + +<p><i>Specimens</i>, <a href="#Page_184">184</a>.</p> + +<p><i>Spermaphore</i>, or <i>Spermophore</i>, one of the names of the placenta.</p> + +<p><i>Spermum</i>, Latin form of Greek word for seed; much used in composition.</p> + +<p><i>Spica</i>, Latin for spike; hence <i>Spicate</i>, in a spike, <i>Spiciform</i>, in +shape resembling a spike.</p> + +<p><i>Spike</i>, an inflorescence like a raceme, only the flowers are sessile, +<a href="#Page_74">74</a>.</p> + +<p><i>Spikelet</i>, a small or a secondary spike; the inflorescence of Grasses.</p> + +<p><i>Spine</i>, <a href="#Page_41">41</a>, <a href="#Page_64">64</a>.</p> + +<p><i>Spindle-shaped</i>, tapering to each end, like a radish, <a href="#Page_36">36</a>.</p> + +<p><i>Spinescent</i>, tipped by or degenerating into a thorn.</p> + +<p><i>Spinose</i>, or <i>Spiniferous</i>, thorny.</p> + +<p><i>Spiral Vessels</i> or <i>ducts</i>, <a href="#Page_135">135</a>.</p> + +<p><i>Spithameous</i>, span-high.</p> + +<p><i>Spora</i>, Greek name for seed, used in compound words.</p> + +<p><i>Sporadic</i>, widely dispersed.</p> + +<p><i>Sporangium</i>, a spore-case in Ferns, &c., <a href="#Page_158">158</a>.</p> + +<p><i>Spore</i>, a body resulting from the fructification of Cryptogamous +plants, in them the analogue of a seed.</p> + +<p><i>Spore-case</i> (<i>Sporangium</i>), <a href="#Page_158">158</a>.</p> + +<p><i>Sporocarp</i>, <a href="#Page_162">162</a>.</p> + +<p><i>Sport</i>, a newly appeared variation, <a href="#Page_176">176</a>.</p> + +<p><i>Sporule</i>, same as a spore, or a small spore.</p> + +<p><i>Spumescent</i>, appearing like froth.</p> + +<p><i>Spur</i>, any projecting appendage of the flower, looking like a spur but +hollow, as that of Larkspur, fig. <a href="#Fig239">239</a>.</p> + +<p><i>Squamate</i>, <i>Squamose</i>, or <i>Squamaceous</i>, furnished with scales +(<i>squamæ</i>).</p> + +<span class="pagenum"><a name="Page_222" id="Page_222">[Pg 222]</a></span><p><i>Squamellate</i>, or <i>Squamulose</i>, furnished with little scales +(<i>Squamellæ</i>, or <i>Squamulæ</i>).</p> + +<p><i>Squamiform</i>, shaped like a scale.</p> + +<p><i>Squarrose</i>, where scales, leaves, or any appendages spread widely from +the axis on which they are thickly set.</p> + +<p><i>Squarrulose</i>, diminutive of <i>squarrose</i>; slightly squarrose.</p> + +<p><i>Stachys</i>, Greek for spike.</p> + +<p><i>Stalk</i>, the stem, petiole, peduncle, &c., as the case may be.</p> + +<p><i>Stamen</i>, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_98">98</a>.</p> + +<p><i>Staminate</i>, furnished with stamens, <a href="#Page_86">86</a>. <i>Stamineal</i>, relating to the +stamens.</p> + +<p><i>Staminodium</i>, an abortive stamen, or other body in place of a stamem.</p> + +<p><a name="Standard" id="Standard"></a><i>Standard</i>, the upper petal of a papilionaceous corolla, <a href="#Page_92">92</a>.</p> + +<p><i>Starch</i>, <a href="#Page_136">136</a>, <a href="#Page_163">163</a>.</p> + +<p><i>Station</i>, the particular kind of situation in which a plant naturally +occurs.</p> + +<p><i>Stellate</i>, <i>Stellular</i>, starry or star-like; where several similar +parts spread out from a common centre, like a star.</p> + +<p><i>Stem</i>, <a href="#Page_39">39</a>. <i>Stemlet</i>, diminutive stem.</p> + +<p><i>Stemless</i>, destitute or apparently destitute of stem.</p> + +<p><i>Stenos</i>, Greek for narrow; hence <i>Stenophyllous</i>, narrow-leaved, &c.</p> + +<p><i>Sterile</i>, barren or imperfect.</p> + +<p><i>Stigma</i>, the part of the pistil which receives the pollen, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_105">105</a>.</p> + +<p><i>Stigmatic</i>, or <i>Stigmatose</i>, belonging to the stigma.</p> + +<p><i>Stipe</i> (Latin <i>Stipes</i>), the stalk of a pistil, &c., when it has any, +<a href="#Page_112">112</a>; also of a Fern, <a href="#Page_158">158</a>, and of a Mushroom, <a href="#Page_172">172</a>.</p> + +<p><i>Stipel</i>, a stipule of a leaflet, as of the Bean, &c.</p> + +<p><i>Stipellate</i>, furnished with stipels, as in the Bean tribe.</p> + +<p><i>Stipitate</i>, furnished with a stipe.</p> + +<p><i>Stipulaceous</i>, belonging to stipules. <i>Stipulate</i>, furnished with +stipules.</p> + +<p><i>Stipules</i>, the appendages on each side of the base of certain leaves, +<a href="#Page_66">66</a>.</p> + +<p><i>Stirps</i> (plural, <i>stirpes</i>), Latin for race.</p> + +<p><i>Stock</i>, used for race or source. Also for any root-like base from which +the herb grows up.</p> + +<p><i>Stole</i>, or <i>Stolon</i>, a trailing or reclined and rooting shoot, <a href="#Page_40">40</a>.</p> + +<p><i>Stoloniferous</i>, producing stolons.</p> + +<p><i>Stomate</i> (Latin <i>Stoma</i>, plural <i>Stomata</i>), the breathing-pores of +leaves, <a href="#Page_144">144</a>.</p> + +<p><i>Stone-fruit</i>, <a href="#Page_119">119</a>.</p> + +<p><i>Storage-leaves</i>, <a href="#Page_62">62</a>.</p> + +<p><i>Stramineous</i>, straw-like, or straw-colored.</p> + +<p><i>Strap-shaped</i>, long, flat, and narrow.</p> + +<p><i>Striate</i>, or <i>Striated</i>, marked with slender longitudinal grooves or +stripes.</p> + +<p><i>Strict</i>, close and narrow; straight and narrow.</p> + +<p><i>Strigillose</i>, <i>Strigose</i>, beset with stout and appressed, stiff or +rigid bristles.</p> + +<p><i>Strobilaceous</i>, relating to or resembling a</p> + +<p><i>Strobile</i>, a multiple fruit in the form of a cone or head, <a href="#Page_124">124</a>.</p> + +<p><i>Strombuliform</i>, twisted, like a spiral shell.</p> + +<p><i>Strophiole</i>, same as <i>caruncle</i>, <a href="#Page_126">126</a>. <i>Strophiolate</i>, furnished with a +strophiole.</p> + +<p><i>Struma</i>, a wen; a swelling or protuberance of any organ.</p> + +<p><i>Strumose</i>, bearing a struma.</p> + +<p><i>Stupose</i>, like tow.</p> + +<p><i>Style</i>, a stalk between ovary and stigma, <a href="#Page_14">14</a>, <a href="#Page_80">80</a>, <a href="#Page_105">105</a>.</p> + +<p><i>Styliferous</i>, <i>Stylose</i>, bearing styles or conspicuous ones.</p> + +<p><i>Stylopodium</i>, an epigynous disk, or an enlargement at the base of the +style.</p> + +<p><i>Sub-</i>, as a prefix, about, nearly, somewhat; as <i>Subcordate</i>, slightly +cordate; <i>Subserrate</i>, slightly serrate; <i>Subaxillary</i>, just beneath the +axil, &c.</p> + +<p><i>Subclass</i>, <i>Suborder</i>, <i>Subtribe</i>, <a href="#Page_178">178</a>.</p> + +<p><i>Suberose</i>, corky or cork-like in texture.</p> + +<p><i>Subulate</i>, awl-shaped; tapering from a broadish or thickish base to a +sharp point.</p> + +<p><i>Succise</i>, as if cut off at lower end.</p> + +<p><i>Succubous</i>, when crowded leaves are each covered by base of next above.</p> + +<span class="pagenum"><a name="Page_223" id="Page_223">[Pg 223]</a></span><p><i>Suckers</i>, shoots from subterranean branches, <a href="#Page_39">39</a>.</p> + +<p><i>Suffrutescent</i>, slightly shrubby or woody at the base only, <a href="#Page_39">39</a>.</p> + +<p><i>Suffruticose</i>, rather more than suffrutescent, <a href="#Page_37">37</a>, <a href="#Page_39">39</a>.</p> + +<p><i>Sulcate</i>, grooved longitudinally with deep furrows.</p> + +<p><i>Superior</i>, above, <a href="#Page_96">96</a>; sometimes equivalent to posterior, <a href="#Page_96">96</a>.</p> + +<p><i>Supernumerary Buds</i>, <a href="#Page_30">30</a>, <a href="#Page_31">31</a>.</p> + +<p><i>Supervolute</i>, plaited and convolute in bud, <a href="#Page_97">97</a>.</p> + +<p><i>Supine</i>, lying flat, with face upward.</p> + +<p><i>Supra-axillary</i>, borne above the axil, as some buds, <a href="#Page_31">31</a>.</p> + +<p><i>Supra-decompound</i>, many times compounded or divided.</p> + +<p><i>Surculose</i>, producing suckers (<i>Surculi</i>) or shoots resembling them.</p> + +<p><i>Suspended</i>, hanging down. Suspended ovules or seeds hang from the very +summit of the cell which contains them.</p> + +<p><i>Sutural</i>, belonging or relating to a suture.</p> + +<p><i>Suture</i>, the line of junction of contiguous parts grown together, <a href="#Page_106">106</a>.</p> + +<p><i>Sword-shaped</i>, applied to narrow leaves, with acute parallel edges, +tapering above.</p> + +<p><i>Syconium</i>, the fig-fruit, <a href="#Page_124">124</a>.</p> + +<p><i>Sylvestrine</i>, growing in woods.</p> + +<p><i>Symmetrical Flower</i>, similar in the number of parts of each set, <a href="#Page_82">82</a>.</p> + +<p><i>Sympetalous</i>, same as gamopetalous.</p> + +<p><i>Sympode</i>, <i>Sympodium</i>, a stem composed of a series of superposed +branches in such a way as to imitate a simple axis, as in Grape-vine.</p> + +<p><i>Synantherous</i> or <i>Syngenesious</i>, where stamens are united by their +anthers, <a href="#Page_100">100</a>.</p> + +<p><i>Syncarpous</i> (fruit or pistil), composed of several carpels consolidated +into one.</p> + +<p><i>Synonym</i>, an equivalent superseded name.</p> + +<p><i>Synsepalous</i>, same as gamosepalous.</p> + +<p><i>System</i> (artificial and natural), <a href="#Page_182">182</a>, <a href="#Page_183">183</a>.</p> + +<p><i>Systematic Botany</i>, the study of plants after their kinds, <a href="#Page_9">9</a>.</p> +</div> + +<div class="index"> +<p><i>Tabescent</i>, wasting or shrivelling.</p> + +<p><i>Tail</i>, any long and slender prolongation of an organ.</p> + +<p><i>Taper-pointed</i>, same as acuminate, <a href="#Page_54">54</a>.</p> + +<p><i>Tap-root</i>, a root with a stout tapering body, <a href="#Page_32">32</a>-<a href="#Page_35">35</a>.</p> + +<p><i>Tawny</i>, dull yellowish, with a tinge of brown.</p> + +<p><i>Taxonomy</i>, the part of botany which treats of classification.</p> + +<p><i>Tegmen</i>, a name for the inner seed-coat.</p> + +<p><i>Tendril</i>, a thread-shaped organ used for climbing, <a href="#Page_40">40</a>.</p> + +<p><i>Terete</i>, long and round; same as <i>cylindrical</i>, only it may taper.</p> + +<p><i>Terminal</i>, borne at, or belonging to, the extremity or summit.</p> + +<p><i>Terminology</i> treats of technical terms; same as <i>Glossology</i>, <a href="#Page_181">181</a>.</p> + +<p><i>Ternate</i>, <i>Ternately</i>, in threes.</p> + +<p><i>Tessellate</i>, in checker-work.</p> + +<p><i>Testa</i>, the outer (and usually the harder) coat or shell of the seed, +<a href="#Page_125">125</a>.</p> + +<p><i>Testaceous</i>, the color of unglazed pottery.</p> + +<p><i>Tetra-</i> (in words of Greek composition), four; as, <i>Tetracoccous</i>, of +four cocci.</p> + +<p><i>Tetradynamous</i>, where a flower has six stamens, two shorter than the +four, <a href="#Page_101">101</a>.</p> + +<p><i>Tetragonal</i>, four-angled. <i>Tetragynous</i>, with four pistils or styles. +<i>Tetramerous</i>, with its parts or sets in fours. <i>Tetrandrous</i>, with four +stamens, <a href="#Page_100">100</a>.</p> + +<p><i>Tetraspore</i>, a quadruple spore, <a href="#Page_169">169</a>.</p> + +<p><i>Thalamaflorous</i>, with petals and stamens inserted on the torus or +<i>Thalamus</i>.</p> + +<p><i>Thallophyta</i>, <i>Thallophytes</i>, <a href="#Page_165">165</a>.</p> + +<p><i>Thallus</i>, a stratum, in place of stem and leaves, <a href="#Page_165">165</a>.</p> + +<p><i>Theca</i>, a case; the cells or lobes of the anther.</p> + +<p><i>Thecaphore</i>, the stipe of a carpel, <a href="#Page_113">113</a>.</p> + +<p><i>Thorn</i>, an indurated pointed branch, <a href="#Page_41">41</a>, <a href="#Page_42">42</a>.</p> + +<p><i>Thread-shaped</i>, slender and round or roundish, like a thread.</p> + +<p><i>Throat</i>, the opening or gorge of a monopetalous corolla, &c., where the +border and the tube join, and a little below, <a href="#Page_89">89</a>.</p> + +<span class="pagenum"><a name="Page_224" id="Page_224">[Pg 224]</a></span><p><i>Thyrse</i> or <i>Thyrsus</i>, a compact and pyramidal panicle of cymes or +cymules, <a href="#Page_79">79</a>.</p> + +<p><i>Tomentose</i>, clothed with matted woolly hairs (<i>tomentum</i>).</p> + +<p><i>Tongue-shaped</i>, long and flat, but thickish and blunt.</p> + +<p><i>Toothed</i>, furnished with teeth or short projections of any sort on the +margin; used especially when these are sharp, like saw-teeth, and do not +point forwards, <a href="#Page_55">55</a>.</p> + +<p><i>Top-shaped</i>, shaped like a top, or a cone with apex downwards.</p> + +<p><i>Torose</i>, <i>Torulose</i>, knobby; where a cylindrical body is swollen at +intervals.</p> + +<p><i>Torus</i>, the receptacle of the flower, <a href="#Page_81">81</a>, <a href="#Page_112">112</a>.</p> + +<p><i>Trachea</i>, a spiral duct.</p> + +<p><i>Trachys</i>, Greek for rough; used in compounds, as, <i>Trachyspermous</i>, +rough-seeded.</p> + +<p><i>Transverse</i>, across, standing right and left instead of fore and aft.</p> + +<p><i>Tri-</i> (in composition), three; as,</p> + +<p><i>Triadelphous</i>, stamens united by their filaments into three bundles, +<a href="#Page_99">99</a>.</p> + +<p><i>Triandrous</i>, where the flower has three stamens, <a href="#Page_112">112</a>.</p> + +<p><i>Tribe</i>, <a href="#Page_178">178</a>.</p> + +<p><i>Trichome</i>, of the nature of hair or pubescence.</p> + +<p><i>Trichotomous</i>, three-forked.</p> + +<p><i>Tricoccous</i>, of three cocci or roundish carpels.</p> + +<p><i>Tricolor</i>, having three colors.</p> + +<p><i>Tricostate</i>, having three ribs.</p> + +<p><i>Tricuspidate</i>, three-pointed.</p> + +<p><i>Tridentate</i>, three-toothed.</p> + +<p><i>Triennial</i>, lasting for three years.</p> + +<p><i>Trifarious</i>, in three vertical rows; looking three ways.</p> + +<p><i>Trifid</i>, three-cleft, <a href="#Page_56">56</a>.</p> + +<p><i>Trifoliate</i>, three-leaved. <i>Trifoliolate</i>, of three leaflets.</p> + +<p><i>Trifurcate</i>, three-forked.</p> + +<p><i>Trigonous</i>, three-angled, or triangular.</p> + +<p><i>Trigynous</i>, with three pistils or styles, <a href="#Page_116">116</a>.</p> + +<p><i>Trijugate</i>, in three pairs (<i>jugi</i>).</p> + +<p><i>Trilobed</i> or <i>Trilobate</i>, three-lobed, <a href="#Page_55">55</a>.</p> + +<p><i>Trilocular</i>, three-celled, as the pistils or pods in fig. <a href="#Fig326">328-330</a>.</p> + +<p><i>Trimerous</i>, with its parts in threes.</p> + +<p><i>Trimorphism</i>, <a href="#Page_117">117</a>. <i>Trimorphic</i> or <i>Trimorphous</i>, in three forms.</p> + +<p><i>Trinervate</i>, three-nerved, or with three slender ribs.</p> + +<p><i>Triœcious</i>, where there are three sorts of flowers on the same or +different individuals, as in Red Maple. A form of Polygamous.</p> + +<p><i>Tripartible</i>, separable into three pieces. <i>Tripartite</i>, three-parted, +<a href="#Page_55">55</a>.</p> + +<p><i>Tripetalous</i>, having three petals.</p> + +<p><i>Triphyllous</i>, three-leaved; composed of three pieces.</p> + +<p><i>Tripinnate</i>, thrice pinnate, <a href="#Page_59">59</a>. <i>Tripinnatifid</i>, thrice pinnately +cleft, <a href="#Page_57">57</a>.</p> + +<p><i>Triple-ribbed</i>, <i>Triple-nerved</i>, &c., where a midrib branches into +three, near the base of the leaf.</p> + +<p><i>Triquetrous</i>, sharply three-angled; and especially with the sides +concave, like a bayonet.</p> + +<p><i>Triserial</i>, or <i>Triseriate</i>, in three rows, under each other.</p> + +<p><i>Tristichous</i>, in three longitudinal or perpendicular ranks.</p> + +<p><i>Tristigmatic</i>, or <i>Tristigmatose</i>, having three stigmas.</p> + +<p><i>Trisulcate</i>, three-grooved.</p> + +<p><i>Triternate</i>, three times ternate, <a href="#Page_59">59</a>.</p> + +<p><i>Trivial Name</i>, the specific name.</p> + +<p><i>Trochlear</i>, pulley-shaped.</p> + +<p><i>Trumpet-shaped</i>, tubular; enlarged at or towards the summit.</p> + +<p><i>Truncate</i>, as if cut off at the top.</p> + +<p><i>Trunk</i>, the main stem or general body of a stem or tree.</p> + +<p><i>Tube</i> (of corolla, &c.), <a href="#Page_89">89</a>.</p> + +<p><i>Tuber</i>, a thickened portion of a subterranean stem or branch, provided +with eyes (buds) on the sides, <a href="#Page_44">44</a>.</p> + +<p><i>Tubercle</i>, a small excrescence.</p> + +<p><i>Tubercled</i>, or <i>Tuberculate</i>, bearing excrescences or pimples.</p> + +<p><i>Tubæform</i>, trumpet-shaped.</p> + +<p><i>Tuberous</i>, resembling a tuber. <i>Tuberiferous</i>, bearing tubers.</p> + +<p><i>Tubular</i>, hollow and of an elongated form; hollowed like a pipe, <a href="#Page_91">91</a>.</p> + +<span class="pagenum"><a name="Page_225" id="Page_225">[Pg 225]</a></span><p><i>Tubuliflorous</i>, bearing only tubular flowers.</p> + +<p><i>Tunicate</i>, coated; invested with layers, as an onion, <a href="#Page_46">46</a>.</p> + +<p><i>Turbinate</i>, top-shaped.</p> + +<p><i>Turio</i> (plural <i>turiones</i>), strong young shoots or suckers springing +out of the ground; as Asparagus-shoots.</p> + +<p><i>Turnip-shaped</i>, broader than high, abruptly narrowed below, <a href="#Page_35">35</a>.</p> + +<p><i>Twining</i>, ascending by coiling round a support, <a href="#Page_39">39</a>.</p> + +<p><i>Type</i>, the ideal pattern, <a href="#Page_10">10</a>.</p> + +<p><i>Typical</i>, well exemplifying the characteristics of a species, genus, +&c.</p> +</div> + +<div class="index"> +<p><i>Uliginose</i>, growing in swamps.</p> + +<p><i>Umbel</i>, the umbrella-like form of inflorescence, <a href="#Page_74">74</a>.</p> + +<p><i>Umbellate</i>, in umbels. <i>Umbelliferous</i>, bearing umbels.</p> + +<p><i>Umbellet</i> (<i>umbellula</i>), a secondary or partial umbel, <a href="#Page_76">76</a>.</p> + +<p><i>Umbilicate</i>, depressed in the centre, like the ends of an apple; with a +navel.</p> + +<p><i>Umbonate</i>, bossed; furnished with a low, rounded projection like a boss +(<i>umbo</i>).</p> + +<p><i>Umbraculiform</i>, umbrella-shaped.</p> + +<p><i>Unarmed</i>, destitute of spines, prickles, and the like.</p> + +<p><i>Uncial</i>, an inch (<i>uncia</i>) in length.</p> + +<p><i>Uncinate</i>, or <i>Uncate</i>, hook-shaped; hooked over at the end.</p> + +<p><i>Under-shrub</i>, partially shrubby, or a very low shrub.</p> + +<p><i>Undulate</i> or <i>Undate</i>, wavy, or wavy-margined, <a href="#Page_55">55</a>.</p> + +<p><i>Unequally pinnate</i>, pinnate with an odd number of leaflets, <a href="#Page_65">65</a>.</p> + +<p><i>Unguiculate</i>, furnished with a claw (<i>unguis</i>), <a href="#Page_91">91</a>.</p> + +<p><i>Uni-</i>, in compound words, one; as <i>Unicellular</i>, one-celled.</p> + +<p><i>Uniflorous</i>, one-flowered.</p> + +<p><i>Unifoliate</i>, one-leaved. <i>Unifoliolate</i>, of one leaflet, <a href="#Page_59">59</a>.</p> + +<p><i>Unijugate</i>, of one pair.</p> + +<p><i>Unilabiate</i>, one-lipped.</p> + +<p><i>Unilateral</i>, one-sided.</p> + +<p><i>Unilocular</i>, one-celled.</p> + +<p><i>Uniovulate</i>, having only one ovule.</p> + +<p><i>Uniserial</i>, in one horizontal row.</p> + +<p><i>Unisexual</i>, having stamens or pistils only, <a href="#Page_85">85</a>.</p> + +<p><i>Univalved</i>, a pod of only one piece after dehiscence.</p> + +<p><i>Unsymmetrical Flowers</i>, <a href="#Page_86">86</a>.</p> + +<p><i>Urceolate</i>, urn-shaped.</p> + +<p><i>Utricle</i>, a small thin-walled, one-seeded fruit, as of Goosefoot, <a href="#Page_121">121</a>.</p> + +<p><i>Utricular</i>, like a small bladder.</p> +</div> + +<div class="index"> +<p><i>Vaginate</i>, sheathed, surrounded by a sheath (<i>vagina</i>).</p> + +<p><i>Valve</i>, one of the pieces (or doors) into which a dehiscent pod, or any +similar body, splits, <a href="#Page_122">122</a>, <a href="#Page_123">123</a>.</p> + +<p><i>Valvate</i>, <i>Valvular</i>, opening by valves. <i>Valvate</i>, in æstivation, <a href="#Page_97">97</a>.</p> + +<p><i>Variety</i>, <a href="#Page_176">176</a>.</p> + +<p><i>Vascular</i>, containing vessels, or consisting of vessels or ducts, <a href="#Page_134">134</a>.</p> + +<p><i>Vascular Cryptogams</i>, <a href="#Page_156">156</a>.</p> + +<p><i>Vaulted</i>, arched; same as <i>fornicate</i>.</p> + +<p><i>Vegetable Life</i>, &c., <a href="#Page_128">128</a>. <i>Vegetable anatomy</i>, <a href="#Page_129">129</a>.</p> + +<p><i>Veins</i>, the small ribs or branches of the framework of leaves, &c., <a href="#Page_49">49</a>, +<a href="#Page_50">50</a>.</p> + +<p><i>Veined</i>, <i>Veiny</i>, furnished with evident veins. <i>Veinless</i>, destitute +of veins.</p> + +<p><i>Veinlets</i>, the smaller ramifications of veins, <a href="#Page_50">50</a>.</p> + +<p><i>Velate</i>, furnished with a veil.</p> + +<p><i>Velutinous</i>, velvety to the touch.</p> + +<p><i>Venation</i>, the veining of leaves, &c., <a href="#Page_50">50</a>.</p> + +<p><i>Venenate</i>, poisonous.</p> + +<p><i>Venose</i>, veiny; furnished with conspicuous veins.</p> + +<p><i>Ventral</i>, belonging to that side of a simple pistil, or other organ, +which looks towards the axis or centre of the flower; the opposite of +dorsal; as the</p> + +<p><i>Ventral Suture</i>, <a href="#Page_106">106</a>.</p> + +<p><i>Ventricose</i>, inflated or swelled out on one side.</p> + +<span class="pagenum"><a name="Page_226" id="Page_226">[Pg 226]</a></span><p><i>Venulose</i>, furnished with veinlets.</p> + +<p><i>Vermicular</i>, worm-like, shaped like worms.</p> + +<p><i>Vernal</i>, belonging to spring.</p> + +<p><i>Vernation</i>, the arrangement of the leaves in the bud, <a href="#Page_71">71</a>.</p> + +<p><i>Vernicose</i>, the surface appearing as if varnished.</p> + +<p><i>Verrucose</i>, warty; beset with little projections like warts.</p> + +<p><i>Versatile</i>, attached by one point, so that it may swing to and fro, +<a href="#Page_101">101</a>.</p> + +<p><i>Vertex</i>, same as <i>apex</i>.</p> + +<p><i>Vertical</i>, upright, perpendicular to the horizon, lengthwise.</p> + +<p><i>Verticil</i>, a whorl, <a href="#Page_68">68</a>. <i>Verticillate</i>, whorled, <a href="#Page_68">68</a>.</p> + +<p><i>Verticillaster</i>, a false whorl, formed of a pair of opposite cymes.</p> + +<p><i>Vesicular</i>, bladdery.</p> + +<p><i>Vespertine</i>, appearing or expanding at evening.</p> + +<p><i>Vessels</i>, ducts, &c., <a href="#Page_134">134</a>.</p> + +<p><i>Vexillary</i>, <i>Vexillar</i>, relating to the</p> + +<p><i>Vexillum</i>, the standard of a papilionaceous flower, <a href="#Page_92">92</a>.</p> + +<p><i>Villose</i>, shaggy with long and soft hairs (<i>Villosity</i>).</p> + +<p><i>Vimineous</i>, producing slender twigs, such as those used for +wicker-work.</p> + +<p><i>Vine</i>, in the American use, any trailing or climbing stem; as a +Grape-vine.</p> + +<p><i>Virescent</i>, <i>Viridescent</i>, greenish; turning green.</p> + +<p><i>Virgate</i>, wand-shape; as a long, straight, and slender twig.</p> + +<p><i>Viscous</i>, <i>Viscid</i>, having a glutinous surface.</p> + +<p><i>Vitta</i> (plural <i>vittæ</i>), the oil-tubes of the fruit of Umbelliferæ.</p> + +<p><i>Vitelline</i>, yellow, of the hue of yolk of egg.</p> + +<p><i>Viviparous</i>, sprouting or germinating while attached to the parent +plant.</p> + +<p><i>Voluble</i>, twining; as the stem of Hops and Beans, <a href="#Page_39">39</a>.</p> + +<p><i>Volute</i>, rolled up in any way.</p> +</div> + +<div class="index"> +<p><i>Wavy</i>, the surface or margin alternately convex and concave, <a href="#Page_55">55</a>.</p> + +<p><i>Waxy</i>, resembling beeswax in texture or appearance.</p> + +<p><i>Wedge-shaped</i>, broad above, tapering by straight lines to a narrow +base, <a href="#Page_53">53</a>.</p> + +<p><i>Wheel-shaped</i>, <a href="#Page_89">89</a>.</p> + +<p><i>Whorl</i>, an arrangement of leaves, &c., in circles around the stem.</p> + +<p><i>Whorled</i>, arranged in whorls, <a href="#Page_68">68</a>.</p> + +<p><i>Wing</i>, any membranous expansion. <i>Wings</i> of papilionaceous flowers, <a href="#Page_92">92</a>.</p> + +<p><i>Winged</i>, furnished with a wing; as the fruit of Ash and Elm, fig. <a href="#Fig300">300, +301</a>.</p> + +<p><i>Wood</i>, <a href="#Page_133">133</a>, <a href="#Page_142">142</a>. <i>Woody</i>, of the texture or consisting of wood.</p> + +<p><i>Woody Fibre</i>, or <i>Wood-Cells</i>, <a href="#Page_134">134</a>.</p> + +<p><i>Woolly</i>, clothed with long and entangled soft hairs.</p> + +<p><i>Work in plants</i>, <a href="#Page_149">149</a>, <a href="#Page_155">155</a>.</p> +</div> + +<div class="index"> +<p><i>Xanthos</i>, Greek for yellow, used in compounds; as <i>Xanthocarpus</i>, +yellow-fruited.</p> +</div> + +<div class="index"> +<p><i>Zygomorphous</i>, said of a flower which can be bisected only in one plane +into similar halves.</p> +</div> + + +<hr class="chap" /> +<h2>Transcriber's Notes.</h2> + +<p>Spelling variants where it wasn't possible to determine the author's +intent were left as is. These include: "backbone" and "back-bone;" +"Buttonwood" and "Button-wood;" "cross section" and "cross-section;" +"footstalk" and "foot-stalk;" "network" and "net-work;" "Pæony" and +"Peony;" "peapod" and "pea-pod," plus plurals; "penniveined" and +"penni-veined," plus capitalized versions; "Sapwood" and "Sap-wood;" +"Snowball" and "Snow-ball;" "Verticil" and "Verticel;" "Woodsorrel" and +"Wood-sorrel."</p> + +<p>Changed "Venation" to "Vernation" on page vi: "Vernation or +Præfoliation."</p> + +<p>Changed "Isoëtes" to "Isoetes" on page viii: "Quillworts (Isoetes)."</p> + +<p>Changed "liquifies" to "liquefies" on page 21: "as it liquefies."</p> + +<p>Changed "frame-work" to "framework" on page 52: "the framework or +skeleton."</p> + +<p>Changed "leafstalk" to "leaf-stalk" in three places: the caption to +figure 74; on page 71: "bent upon the leaf-stalk;" in the index entry +for "Petiolulate."</p> + +<p>Changed "Honey Locust" to "Honey-Locust" in the caption to figure 95: +"branching thorn of Honey-Locust."</p> + +<p>Page 47 has a reference to Fig. 89, which does not seem to follow +from the text. "Fig. 80" would make more sense, but the original was not +changed.</p> + +<p>Changed "Linnaeus" to "Linnæus" on page 50: "Linnæus called Nerves."</p> + +<p>Removed extra comma before closing parenthesis on page 51: +"the Maple (Fig. 20, 24)."</p> + +<p>Changed "Sugar-Maple" to "Sugar Maple" on page 58: "that of the Sugar +Maple."</p> + +<p>Changed "quadrifoliate" to "quadrifoliolate" on page 59: "trifoliolate, +quadrifoliolate, plurifoliolate."</p> + +<p>The reference to Fig. 167 on page 64 is probably intended to refer to +Fig. 157, but was not changed.</p> + +<p>The caption to figure 186 says that this is a two-ranked arrangement. +Although it's clear from the text and figure that this is actually a +three-ranked arrangement, the original text was preserved.</p> + +<p>Changed colon to semi-colon in the caption to figure 219: "a, filament; +b, anther."</p> + +<p>Changed comma to period after "Fig. 254" in its caption.</p> + +<p>Changed "Funnel-form" to "Funnelform" on page 90: "Funnel-shaped, or +Funnelform."</p> + +<p>Inserted closing parenthesis in caption to figure 304: "Of a Sage +(Salvia Texana)."</p> + +<p>Changed "Butter cup" to "Buttercup" on page 120: "fruit of the +Buttercup."</p> + +<p>Changed "carpophorse" to "carpophore" on page 121: "a slender axis or +carpophore."</p> + +<p>Changed "cocoanut" to "cocoa-nut" on page 122: "such as the cocoa-nut."</p> + +<p>Changed "Sepifragal" to "Septifragal" on page 123: "Septifragal +dehiscence."</p> + +<p>Inserted missing text "(Fig." on page 128: "Rice (Fig. 430a)."</p> + +<p>Perhaps the reference to unit 408 on page 136 should be to unit 402; +but it was not changed.</p> + +<p>Changed "White-Pine" to "White Pine" on page 148: "White Pine and +Basswood."</p> + +<p>Changed "discovary" to "discovery" on page 160: "discovery of a wholly +unsuspected kind."</p> + +<p>Changed "sporo-carps" to "sporocarps" in the caption to figure 520.</p> + +<p>Changed semi-colon to period after "microspores" in the caption to +figure 522.</p> + +<p>Changed "Lepreiurei" to "Leprieurei" in the caption to figure 561.</p> + +<p>Changed "Sun-flowers" to "Sunflowers" on page 187: "Sunflowers and +Thistles."</p> + +<p>Changed "♁" to "♂" on page 189: "To indicate sexes, ♂ +means staminate."</p> + +<p>Preserved the order of the index entries, even though they aren't +strictly alphabetical.</p> + +<p>Removed extra comma after "Amphitropous" in its index entry.</p> + +<p>The index references for "Anemophilous" and "Entomophilous" should +probably be 115, rather than 113, but were not changed.</p> + +<p>The index reference for "Antheridium" should probably be 159, rather +than 150, but was not changed.</p> + +<p>Changed "short-fluited" to "short-fruited" in the index entry for +"Brachy-."</p> + +<p>The index references for "Caulescent" and "Cauline" should probably be +38, rather than 36, but were not changed.</p> + +<p>The index reference for "Cladophylla" should probably be 61, rather +than 64, but was not changed.</p> + +<p>The index reference for "Dotted Ducts" should probably be 134, rather +than 148, but was not changed.</p> + +<p>Changed "trifoliate" to "trifoliolate" in the index entry for +"Foliolate."</p> + +<p>Changed "6" to "13" in the index entry for "Gemmule."</p> + +<p>The index references for "Hemitropous" and "Heterotropous" should +probably be 111, rather than 123, but were not changed.</p> + +<p>The index reference for "Hortus Siccus" was changed from 201 to 186.</p> + +<p>The index reference for "Incised" should probably be 55, rather +than 58, but was not changed.</p> + +<p>The index reference for "Jointed" is 64, and was not changed even though +it does not seem to make sense. Better candidates include 57 and 122.</p> + +<p>Changed the last index reference for "Ligule" from 57 to 67; +also the reference for "Ocreate."</p> + +<p>The index references for "Laticiferous" and "Milk-vessels" were changed +from 138 to 135.</p> + +<p>Changed comma to semi-colon after "86" in the index entry for "Naked."</p> + +<p>The index reference for "Panicle" should probably be 76, rather +than 81, but was not changed.</p> + +<p>The index references for "Pentandrous" and "Triandrous" should probably +be 100, rather than 112, but were not changed.</p> + +<p>Changed "leaf stalk" to "leaf-stalk" in its index entry, as well as the +index entry for "Petiole."</p> + +<p>The index reference for "Pinnule" should probably be 59, rather +than 66, but was not changed.</p> + +<p>The index reference for "Seed-vessel" should probably be 117, rather +than 127, but was not changed.</p> + +<p>The second index reference for "Starch" is 163, and was not changed even +though it does not seem to make sense.</p> + +<p>Changed "one" to "on" in the index entry for "Stipules": "appendages on +each side."</p> + +<p>The index reference to page 37 for "Suffruticose" doesn't seem to make +sense, but was left as is.</p> + +<p>The index reference for "Trigynous" should probably be 105, rather +than 116, but was not changed.</p> + +<p>The index reference for "Unequally Pinnate" should probably be 58, +rather than 65, but was not changed.</p> + + + + + + + + +<pre> + + + + + +End of the Project Gutenberg EBook of The Elements of Botany, by Asa Gray + +*** END OF THIS PROJECT GUTENBERG EBOOK THE ELEMENTS OF BOTANY *** + +***** This file should be named 33757-h.htm or 33757-h.zip ***** +This and all associated files of various formats will be found in: + https://www.gutenberg.org/3/3/7/5/33757/ + +Produced by Curtis Weyant, Stephen H. 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