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diff --git a/59074-0.txt b/59074-0.txt new file mode 100644 index 0000000..1ac5ec1 --- /dev/null +++ b/59074-0.txt @@ -0,0 +1,12188 @@ +*** START OF THE PROJECT GUTENBERG EBOOK 59074 *** + + + + + + + + + + + +BY THE SAME AUTHOR. + +The Foraminifera + +An Introduction to the Study of the Protozoa + +by + +FREDERICK CHAPMAN, A.L.S., F.R.M.S. + + +This book has been written with a view of meeting a demand which +has arisen for a concise account of the Foraminifera, suited to the +requirements of the student of Natural History and Palaeontology. + +With 14 plates and 42 illustrations in the Text. + + +DEMY 8vo. CLOTH, 10s. 6d. + +[Illustration: The Keystone Printing Co., 552-4 Lonsdale St., Melb.] + +[Illustration: =A FOSSIL CRINOID= + +(Helicocrinus plumosus), about 5/6 nat. size, in Silurian Mudstone, +Brunswick, Victoria. + + (_Spec. in Nat. Mus., Melbourne_). +] + + + + +Australasian Fossils + +A Students' Manual of Palaeontology + +By FREDERICK CHAPMAN, + +Palaeontologist to the National Museum, Melbourne. + +Formerly Assistant in the Geological Department of the Royal College of +Science, London. + +Assoc. Linnean Soc. [Lond.], F.R.M.S., etc. + +Author of "The Foraminifera," "A Monograph of the Silurian Bivalved +Mollusca of Victoria," "New or Little-known Victorian Fossils in the +National Museum," etc. + +With an Introduction by + +PROFESSOR E. W. SKEATS, D.Sc., F.G.S. + +GEORGE ROBERTSON & COMPANY PROPY. LTD., + +Melbourne, Sydney, Adelaide, Brisbane and London. + +1914. + + +To + +PROFESSOR JOHN WESLEY JUDD + + + this work is dedicated as a slight tribute of esteem, and in grateful + acknowledgement of kindly help and encouragement through many years. + + + + +CONTENTS. + + + Page + + Preface 10 + + Introduction by Professor E. W. Skeats, D.Sc., F.G.S. 13 + + + PART I.--GENERAL PRINCIPLES. + + Chap. I.--Nature and uses of Fossils 21 + + " II.--Classification of Fossil Animals and Plants 34 + + " III.--The Geological Epochs and Time-range of Fossils 41 + + " IV.--How Fossils are Found, and the Rocks They Form 51 + + + PART II.--SYSTEMATIC PALAEONTOLOGY. + + Chap. V.--Fossil Plants 82 + + " VI.--Fossil Foraminifera and Radiolaria 95 + + " VII.--Fossil Sponges, Corals and Graptolites 107 + + " VIII.--Fossil Starfishes, Sea-lilies and Sea-urchins 133 + + " IX.--Fossil Worms, Sea-mats and Lamp-shells 152 + + " X.--Fossil Shell-fish 174 + + " XI.--Fossil Trilobites, Crustacea and Insects 220 + + " XII.--Fossil Fishes, Amphibians, Reptiles, Birds and Mammals 257 + + + Appendix.--Notes on Collecting and Preserving Fossils 315 + + Index 321 + + + + +LIST OF ILLUSTRATIONS. + + + Fig. Page + + 1. Fossil Shells in clay 22 + + 2. Tracks, probably of Crustaceans 22 + + 3. Structure of Silicified Wood in tangential section: + _Araucarioxylon Daintreei_, Chapm. 24 + + 4. Portrait of William Smith 26 + + 5. Raised Beach: Brighton, England 28 + + 6. Raised Beach: Torquay, Victoria 28 + + 7. Marine Fossils in Volcanic Tuff: Summit of Snowdon 29 + + 8. Kitchen Middens: Torquay, Victoria 30 + + 9. Submerged Forest on the Cheshire Coast 30 + + 10. _Pecten murrayanus_, Tate. A fossil shell allied to + a living species 32 + + 11. Cliff section: Torquay, Victoria 42 + + 12. Diagram of superposition of Strata 42 + + 13. Diagram of the Range-in-time of Australasian Fossils 50 + + 14. _Diprotodon_ skeletons in situ: Lake Callabonna, S. Australia 51 + + 15. Bird remains on sand dunes: King Island, Bass Strait 52 + + 16. Impression of Bird's feather in Ironstone: Western Victoria 52 + + 17. A Fossil Turtle: _Notochelone costata_, Owen sp. 52 + + 18. A Ganoid Fish: _Pristisomus crassus_, A. S. Woodward 54 + + 19. A fossil Insect in amber (_Tipula sp._) 54 + + 20. A fossil Crustacean: _Thalassina emerii_, Bell 55 + + 21. An Ammonite: _Desmoceras flindersi_, McCoy sp. 55 + + 22. Belemnites: _Belemnites diptycha_, McCoy 56 + + 23. A Group of Lamp-shells: _Magellania flavescens_, Lam. sp. 56 + + 24. Zoarium of a living Polyzoan: _Retepora_ sp. 58 + + 25. A fossil Polyzoan: _Macropora clarkei_, T. Woods sp. 58 + + 26. Fossil Worm-tubes: (?) _Serpula_ 60 + + 27. A living Sea-urchin: _Strongylocentrotus erythrogrammus_, Val. 60 + + 28. A fossil Sea-urchin: _Linthia antiaustrails_, Tate 60 + + 29. A fossil Brittle-Star: _Ophioderma egertoni_, Brod. sp. 60 + + 30. A fossil Crinoid: _Taxocrinus simplex_, Phillips sp. 62 + + 31. Graptolites on Slate: _Tetragraptus fruticosus_, J. Hall sp. 62 + + 32. A Stromatoporoid: _Actinostroma_ 63 + + 33. Corals in Devonian Marble: _Favosites_ 64 + + 34. Siliceous Skeleton of a living Sponge: (?) _Chonelasma_ 64 + + 35. Spicules of a fossil Sponge: _Ecionema newberyi_, McCoy sp. 65 + + 36. Nummulites: _N. gizehensis_, Ehr. var. _champollioni_, De + la Harpe 65 + + 37. Cainozoic Radiolaria 66 + + 38. Radiolaria in Siliceous Limestone 67 + + 39. Travertin Limestone, with leaves of Beech (_Fagus_) 67 + + 40. Freshwater Limestone with shells (_Bulinus_) 68 + + 41. Hardened mudstone with Brachiopods (_Orthis_, etc.) 69 + + 42. Diatomaceous Earth 72 + + 43. _Lepidocyclina_ Limestone 73 + + 44. Coral in Limestone: _Favosites grandipora_, Eth. fil. 74 + + 45. Crinoidal Limestone 74 + + 46. Turritella Limestone 75 + + 47. Ostracodal Limestone 75 + + 48. _Halimeda_ Limestone 77 + + 49. Tasmanite: a Spore Coal 77 + + 50. Kerosene Shale 77 + + 51. Bone Bed 77 + + 52. Bone Breccia 79 + + 53. Cainozoic Ironstone with Leaves (_Banksia_) 80 + + 54. _Girvanella conferta_, Chapm., in Silurian Limestone 83 + + 55. Palaeozoic Plants 83 + + 56. Restoration of _Lepidodendron_ 84 + + 57. Stem of _Lepidodendron (Lepidophloios)_, showing leaf-scars 84 + + 58. Upper Palaeozoic Plants 85 + + 59. Map of Gondwana-Land 87 + + 60. Mesozoic Plants 88 + + 61. Cainozoic Plants 90 + + 62. Eucalyptus leaves from the Deep Leads 92 + + 63. Palaeozoic and Mesozoic Foraminifera 97 + + 64. _Lepidocyclina marginata_, Mich. sp. Sections of shell + showing structure 99 + + 65. Cainozoic Foraminifera 100 + + 66. Fossil Radiolaria 103 + + 67. Palaeozoic Sponges and Archaeocyathinae 108 + + 68. Cainozoic Sponges 111 + + 69. Silurian Corals 111 + + 70. Upper Palaeozoic Corals 116 + + 71. Cainozoic Corals 118 + + 72. Stromatoporoidea and Cladophora 121 + + 73. Lower Ordovician Graptolites 125 + + 74. Lower Ordovician Graptolites 125 + + 75. Upper Ordovician and Silurian Graptolites 127 + + 76. Fossil Crinoids 135 + + 77. Fossil Starfishes 140 + + 78. _Protaster brisingoides_, Gregory, in Silurian Sandstone 142 + + 79. _Gregoriura spryi_, Chapm., in Silurian Mudstone 143 + + 80. Cainozoic Sea-urchins 145 + + 81. Cainozoic Sea-urchins 147 + + 82. Fossil Worms 153 + + 83. Palaeozoic Polyzoa 156 + + 84. Cainozoic Polyzoa 157 + + 85. Lower Palaeozoic Brachiopods 159 + + 86. Silurian and Devonian Brachiopods 161 + + 87. Carbopermian Brachiopods 163 + + 88. Mesozoic Brachiopods 165 + + 89. Cainozoic Brachiopods 167 + + 90. Lower Palaeozoic Bivalves 176 + + 91. Palaeozoic Bivalves 179 + + 92. Carbopermian Bivalves 180 + + 93. Lower Mesozoic Bivalves 181 + + 94. Cretaceous Bivalves 183 + + 95. Cainozoic Bivalves 185 + + 96. Cainozoic Bivalves 186 + + 97. Fossil Scaphopods and Chitons 188 + + 98. Lower Palaeozoic Gasteropoda 192 + + 99. Silurian Gasteropoda 194 + + 100. Upper Palaeozoic Gasteropoda 195 + + 101. Mesozoic Gasteropoda 197 + + 102. Cainozoic Gasteropoda 199 + + 103. Cainozoic Gasteropoda 200 + + 104. Late Cainozoic and Pleistocene Gasteropoda 201 + + 105. Palaeozoic Cephalopoda 206 + + 106. Mesozoic and Cainozoic Cephalopoda 208 + + 107. Diagram restoration of an Australian Trilobite (_Dalmanites_) 224 + + 108. Cambrian Trilobites 226 + + 109. Older Silurian Trilobites 228 + + 110. Newer Silurian Trilobites 230 + + 111. Carboniferous Trilobites and a Phyllopod 232 + + 112. Silurian Ostracoda 236 + + 113. Upper Palaeozoic and Mesozoic Ostracoda 238 + + 114. Cainozoic Ostracoda 239 + + 115. Fossil Cirripedes 242 + + 116. Cirripedes. _Lepas anatifera_, Linn.: living goose + barnacle, and _L. pritchardi_, Hall: Cainozoic 242 + + 117. _Ceratiocaris papilio_, Salter 244 + + 118. Ordovician Phyllocarids 245 + + 119. Silurian Phyllocarids 245 + + 120. Fossil Crabs and Insects 247 + + 121. Silurian Eurypterids 249 + + 122. _Thyestes magnificus_, Chapm. 259 + + 123. _Gyracanthides murrayi_, A. S. Woodw. Restoration 260 + + 124. Teeth and Scales of Palaeozoic and Mesozoic Fishes 260 + + 125. _Cleithrolepis granulatus_, Egerton 263 + + 126. Tooth of _Ceratodus avus_, A. S. W., and phalangeal + of a carnivorous Deinosaur 264 + + 127. Scale of _Ceratodus ? avus_ 265 + + 128. The Queensland Lung-fish: _Neoceratodus forsteri_, Krefft 266 + + 129. _Leptolepis gregarius_, A. S. W. 266 + + 130. Cretaceous and Cainozoic Fish-teeth 268 + + 131. Cainozoic Fish remains 270 + + 132. _Bothriceps major_, A. S. W. 273 + + 133. _Ichthyosaurus australis_, McCoy 277 + + 134. Fossil Reptiles 278 + + 135. Impression of Bird's feather, magnified, Cainozoic: Victoria 281 + + 136. _Cnemiornis calcitrans_, Owen 284 + + 137. _Dinornis maximus_, Owen. Great Moa 284 + + 138. _Pachyornis elephantopus_, Owen 285 + + 139. Skeleton of _Sarcophilus ursinus_, Harris sp. 288 + + 140. Skull of fossil specimen of _Sarcophilus ursinus_ 288 + + 141. _Thylacinus major_, Owen. Hind part of mandible 289 + + 142. _Phascolomys pliocenus_, McCoy. Mandible 290 + + 143. Cainozoic Teeth and Otolith 291 + + 144. Skeleton of _Diprotodon australis_, Owen 291 + + 145. Right hind foot of _Diprotodon australis_ 292 + + 146. Restoration of _Diprotodon australis_ 292 + + 147. Skull and mandible of _Thylacoleo carnifex_, Owen 293 + + 148. _Wynyardia bassiana_, Spencer 294 + + 149. Tooth of _Scaldicetus macgeei_, Chapm. 297 + + 150. Impressions of footprints in dune sand-rock, Warrnambool 301 + + Map of Australia, showing chief fossiliferous localities. + + + + +PREFACE. + + +The more important discoveries of fossils in the southern hemisphere +have received, as a rule, very meagre notice in many of the text-books +of Geology and Palaeontology published in England, Germany and America, +and used by Australasian students. It is thought, therefore, that the +time has arrived when an attempt should be made to collect the main facts +bearing upon this subject, in order to present them from an _Australasian_ +standpoint. With this in view, references to fossils occurring in the +northern hemisphere are subordinated, seeing that these may be easily +obtained on reference to the accepted text-books in general use. + +The present work does not presume to furnish a complete record of +Australasian palaeontology, since that would mean the production of a much +more extensive and costly volume. Sufficient information is here given, +however, to form a groundwork for the student of this section of natural +science, and a guide to the collector of these "medals of creation." + +The systematic portion of this book has been arranged primarily from the +biological side, since Palaeontology is the "study of ancient life." +Taking each life-group, therefore, from the lowest to the highest types, +all the divisions represented by fossils are dealt with in turn, beginning +with their occurrence in the oldest rocks and ending with those in the +newest strata. + +If a commendation of the study of fossils, apart from its scientific +utility, were needed, it could be pointed out that palaeontology as a +branch of geology is, _par excellence_, an open-air study: and since +it requires as handmaids all the sister sciences, is a subject of +far-reaching interest. Microscopy and photography are of immense value +in certain branches of fossil research, the former in the examination of +the minute forms of mollusca, foraminifera and ostracoda, the latter in +the exact portraiture of specimens too intricate to copy with the brush, +or too evanescent to long retain, when out of their matrix, their clean +fresh surfaces. With geology or palaeontology as an objective, a country +walk may be a source of much enjoyment to its students, for "in their hand +is Nature like an open book"; and the specimens collected on a summer +excursion may be closely and profitably studied in the spare time of the +winter recess. + +The author sincerely trusts that students may share the same pleasure +which he has derived from the study of these relics of past life; and that +the present attempt to show their relationship both in geological time and +biological organisation, may be the means of inducing many to make further +advances in this fascinating subject. + +In the production of this work several friends and collaborators have +materially assisted, their aid considerably increasing its value. It is +therefore with grateful thanks that the author acknowledges the help and +encouragement given by Professor E. W. Skeats, D.Sc., who has not only +been good enough to write the Introductory passages, but who has carefully +gone over the MS. and made many helpful suggestions. Mr. W. S. Dun, +F.G.S., Palaeontologist to the Geological Survey Branch of the Department +of Mines, Sydney, has also rendered generous help in giving the benefit +of his full acquaintance of the palaeontology of his own State. To the +Trustees of the National Museum the author is under special obligations +for permission to photograph many unique fossil specimens in the Museum +collection, comprising Figs. 3, 16-18, 20-22, 28-31, 35, 39, 40, 45, 46, +51-54, 57, 62, 78, 79, 127, 133, 136, 147 and 148. The author's thanks +are also due to Dr. E. C. Stirling, M.D., M.A., F.R.S., for permission to +use Figs. 143, 144 and 145, whilst similar privileges have been accorded +by Prof. A. G. Seward, F.R.S., Dr. F. A. Bather, F.R.S., and Mr. C. L. +Barrett. Prof. T. W. Edgeworth David, F.R.S., has kindly cleared up +some doubtful points of stratigraphy and further increased the author's +indebtedness by the loan of a unique slide of Radiolaria figured on p. 69. +Mr. Eastwood Moore, to whom special thanks are due, has greatly added to +the pictorial side of this work by his skillful help in preparing many of +the illustrations for the press, as well as in the drawing of the several +maps. The grouped sets of fossils have been especially drawn for this work +by the author. They are either copied from authentic specimens or from +previously published drawings; references to the authorities being given +in the accompanying legends. Dr. T. S. Hall has kindly read the section +on Graptolites and Mammalia. For many helpful suggestions and the careful +reading of proofs, thanks are especially owing to Mr. W. E. G. Simons, Mr. +R. A. Keble, and to my wife. + + + + +INTRODUCTION. + + + Geological Department, + The University, Melbourne. + +William Smith, the Father of English Geology, used to apologize for the +study of palaeontology by claiming that "the search for a fossil is at +least as rational a proceeding as the pursuit of a hare." Those of us who +are accustomed to take the field, armed with a hammer, in the search for +"medals of creation" and from time to time have experienced the sporting +enjoyment of bringing to light a rare or perfect specimen are quite +prepared to support his claim. But the student of fossils needs the help +of a text book to guide him to the literature on the subject, to help +him with his identifications or to indicate that some of his finds are +new and hitherto undescribed. European and American workers have long +been provided with excellent books treating generally of fossils, but the +illustrations have been quite naturally taken mainly from forms occurring +in the Northern Hemisphere. Our own fossil forms both plants and animals +are numerous, interesting and in many cases peculiar, but the literature +concerning them is so widely scattered in various scientific publications +that a warm welcome should be given to this book of Mr. Chapman's, in +which the Australian evidence is brought together and summarised by one, +whose training, long experience, and personal research qualify him to +undertake the task. Especially will teachers and students of Geology and +Palaeontology value such an undertaking. Workers in other countries who +have only partial access to the Australian literature on the subject +should also find this a valuable book of reference. + +In the study of fossils we are concerned with the nature, evolution +and distribution of the former inhabitants of the earth. The study of +Palaeontology may be justified as a means of scientific discipline, for +the contributions the subject makes to the increase of natural knowledge +and the unfolding of panoramas of ancient life. It also provides perhaps +the most positive evidence in the story of evolution. So, too, the student +of the present day distribution of animals and plants finds the key to +many a problem in zoo-geography in the records of past migrations yielded +by the study of fossils in different lands. The stratigraphical geologist +is of course principally concerned with two important aspects of the study +of fossils. + +The masterly generalisation of William Smith that strata can be identified +by their fossil contents established by close study of the rocks and +fossils of the British Oolites has been confirmed generally by subsequent +work. The comparative study of the fossil contents of rocks in widely +separated areas has proved to be the most valuable means by which the +correlation of the rocks can be effected and their identity of age +established. In some cases the recognition of a single fossil species +in two areas separated, perhaps, by thousands of miles may suffice to +demonstrate that the rocks are of the same age. For example, a graptolite +such as _Phyllograptus typus_ is found in many parts of the world, but has +only a very restricted range in time. It has been found only in rocks of +Lower Ordovician age. Its occurrence in Wales and in the rocks of Bendigo +practically suffices to establish the identity in age of the rocks in +these widely separated areas. + +Generally, however, much closer study and a more detailed examination of a +large number of the fossils of a rock series are required before the age +of the rocks can be surely established and a safe correlation made with +distant localities. + +The stratigraphical generalisations to be made from the study of fossils +however must be qualified by certain considerations. Among these are the +fact that our knowledge of the life forms of a given geological period is +necessarily incomplete, that the differences in the fossil contents of +rocks may depend not only on differences of age but also in the conditions +under which the organisms lived and the rocks were accumulated, and that +forms of life originating in one area do not spread themselves immediately +over the earth but migrate at velocities depending on their mode of life +and the presence or absence of barriers to their progress. + +Our incomplete knowledge of the forms living in remote geological periods +arises partly from the fact that some forms had no permanent skeleton +and were therefore incapable of preservation, partly to the obliteration +of the skeletons of organisms through subsequent earth movements in the +rocks or through the solvent action of water. Many land forms, too, +probably disintegrated on the surface before deposits were formed over +the area. Apart from these causes which determine that a full knowledge +of the fossils from ancient rocks in particular, will never be acquired, +our knowledge is incomplete by reason either of difficulty of access to +certain areas or incomplete search. As a result of later discoveries +earlier conclusions based on incomplete evidence as to the age of a rock +series, have not infrequently been modified. + +The study of the present distribution of animals and plants over the +earth is a help in the attempt to decide how far the fossil differences +in the sets of rocks are due to differences in the ages of the rocks or +to differences in the conditions under which the organisms lived. The +present, in this, as in many other geological problems, is the key to the +past. + +We know, for instance, that differences of climate largely control the +geographical distribution of land animals and especially of land plants, +and for that reason among others, fossil plants are generally less +trustworthy guides to geological age than fossil animals. + +In the distribution of marine animals at the present day we find that +organisms of simple structure are generally more wide-spread and less +susceptible to changes in their environment than are the more complex +organisms with specialised structures. Hence we find, for instance, a +fossil species of the Foraminifera may persist unchanged through several +geological periods, while a species of fossil fish has in general not only +a short range in time but often a restricted geographical extent. If we +consider the marine organisms found at the present day we find a number +of free-swimming forms very widely distributed, while a large number are +restricted either by reason of climate or of depth. Certain organisms +are only to be found between high and low tide levels, others between +low tide level and a depth of thirty fathoms, while many quite different +forms live in deeper waters. If we confine our attention to shallow-water +marine forms we note that certain forms are at the present day restricted +to waters of a certain temperature. We find, therefore, a contrast between +arctic and tropical faunas, while other types characterize temperate +latitudes. Climatic and bathymetrical differences at the present day +therefore lead to distinct differences in the distribution of certain +organisms, while other forms, less sensitive to these factors, range +widely and may be almost universally distributed. Similar conditions +obtained in past geological times, and therefore in attempting to +correlate the rocks of one area with those of another those fossils which +are most wide-spread are often found to be the most valuable. + +Attention should also be paid to the conditions under which the deposits +accumulated, since it is clear that rocks may be formed at the same time +in different areas and yet contain many distinct fossils by reason of +climatic or bathymetrical differences. Among living marine organisms we +find certain forms restricted to sandy or muddy sea-bottoms and others +to clear water, and these changes in the conditions of deposition +of sediment have played their part in past geological periods in +determining differences in the fossil faunas of rocks which were laid +down simultaneously. We not infrequently find mudstones passing laterally +into limestones, and this lithological change is always accompanied by +a more or less notable change in the fossil contents of the two rock +types. Such facts emphasize the close connection between stratigraphy +and palaeontology, and indicate that the successful tracing out of the +geological history of any area is only possible when the evidence of the +stratigrapher is reinforced by that provided by the palaeontologist. The +fact that species of animals and plants which have been developed in a +particular area do not spread all over the world at once but migrate +very slowly led Huxley many years ago to put forward his hypothesis of +"homotaxis." He agreed that when the order of succession of rocks and +fossils has been made out in one area, this order and succession will be +found to be generally similar in other areas. The deposits in two such +contrasted areas are homotaxial, that is, show a similarity of order, +but, he claimed, are not necessarily synchronous in their formation. In +whatever parts of the world Carboniferous, Devonian and Silurian fossils +may be found, the rocks with Carboniferous fossils will be found to +overlie those with Devonian, and these in their turn rest upon those +containing Silurian fossils. And yet Huxley maintained that if, say, +Africa was the area in which faunas and floras originated, the migration +of a Silurian fauna and flora might take place so slowly that by the time +it reached Britain the succeeding Devonian forms had developed in Africa, +and when it reached North America, Devonian forms had reached Britain and +Carboniferous forms had developed in Africa. If this were so a Devonian +fauna and flora in Britain may have been contemporaneous with Silurian +life in North America and with a Carboniferous fauna and flora in Africa. + +This could only be true if the time taken for the migration of faunas +and floras was so great as to transcend the boundaries between great +geological periods. This does not appear to be the case, and Huxley's idea +in its extreme form has been generally abandoned. At the same time certain +anomalies in the range in time of individual genera have been noted, and +may possibly be explained on such lines. For instance, among the group of +the graptolites, in Britain the genus _Bryograptus_ occurs only in the +Upper Cambrian and the genus _Leptograptus_ only in the Upper Ordovician +rocks. In Victoria these two genera, together with typical Lower +Ordovician forms, may be found near Lancefield preserved on a single slab +of shale. In the same way, in a single quarry in Triassic rocks in New +South Wales, a number of fossil fish have been found and described, some +of which have been compared to Jurassic, others to Permian, and others to +Carboniferous forms in the Northern Hemisphere. + +Another point which the palaeontologist may occasionally find evidence for +is the existence of "biological asylums," areas which by means of land or +other barriers may be for a long period separated from the main stream +of evolution. We know that the present fauna and flora of Australia +is largely of archaic aspect, as it includes a number of types which +elsewhere have long ago become extinct or were never developed. This +appears to be due to the long isolation of Australia and, as Professor +Gregory happily puts it--its "development in a biological backwater." We +have some evidence that similar asylums have existed in past geological +periods, with the result that in certain areas where uniform conditions +prevailed for a long time or where isolation from competition prevented +rapid evolution, some organisms which became extinct in other areas, +persisted unchanged in the "asylum" into a younger geological period. + +The broad generalizations that rocks may be identified by their fossil +contents and that the testimony of the rocks demonstrates the general +order of evolution from simple to complex forms, have only been placed +on a surer footing by long continued investigations. The modifications +produced by conditions of deposit, of climate and of natural barriers +to migration, while introducing complexities into the problems of +Palaeontology, are every year becoming better known; and when considered +in connection with the variations in the characters of the rocks, provide +valuable and interesting evidence towards the solution of the ultimate +problems of geology and palaeontology, which include the tracing out of +the evolution of the history of the earth from the most remote geological +period to that point at which the geologist hands over his story to the +archaeologist, the historian, and the geographer. + + ERNEST W. SKEATS. + + + + +PART I. + +GENERAL PRINCIPLES. + + + + +CHAPTER I. + +NATURE AND USES OF FOSSILS. + + +=Scope of Geology.--= + +The science of GEOLOGY, of which PALAEONTOLOGY or the study of fossils, +forms a part, is concerned with the nature and structure of the earth, the +physical forces that have shaped it, and the organic agencies that have +helped to build it. + + +=Nature of Fossils.--= + +The remains of animals and plants that formerly existed in the different +periods of the history of the earth are spoken of as fossils. They are +found, more or less plentifully, in such common rocks as clays, shales, +sandstones, and limestones, all of which are comprised in the great series +of Sedimentary Rocks (Fig. 1). + +According to the surroundings of the organisms, whether they existed on +land, in rivers, lakes, estuaries, or the sea, they are spoken of as +belonging to terrestrial, fluviatile, lacustrine, estuarine, or marine +deposits. + +[Illustration: =Fig. 1.--Fossil Shells Embedded in Sandy Clay.= + +About 3/4 nat. size. Of Cainozoic or Tertiary Age (Kalimnan Series). +Grange Burn, near Hamilton, Victoria. + + (_F.C. Coll._) + +(G = Glycimeris. L = Limopsis. N = Natica).] + +[Illustration: =Fig. 2.--Tracks probably of Crustaceans (Phyllocarids).= + +About 3/4 nat. size. Impression of a Slab of Upper Ordovician Shale. +Diggers' Rest, Victoria. + + (_F.C. Coll._) +] + +The name fossil, from the Latin 'fodere' to dig,--'fossilis,' dug out,--is +applied to the remains of any animals or plants which have been buried +either in sediments laid down in water, in materials gathered together +by the wind on land as sand-dunes, in beds of volcanic ash, or in cave +earths. But not only remains of organisms are thus called fossils, for the +name is also applied to structures only indirectly connected with once +living objects, such as rain-prints, ripple-marks, sun-cracks, and tracks +or impressions of worms and insects (Fig. 2). + + +=Preservation of Fossils.--= + +In ordinary terms, fossils are the durable parts of animals and plants +which have resisted complete decay by being covered over with the deposits +above-named. It is due, then, to the fact that they have been kept from +the action of the air, with its destructive bacteria, that we are able to +still find these relics of life in the past. + + +=Petrifaction of Fossils.--= + +When organisms are covered by a tenacious mud, they sometimes undergo no +further change. Very often, however, moisture containing mineral matter +such as carbonate of lime or silica, percolates through the stratum which +contains the fossils, and then they not only have their pores filled with +the mineral, but their actual substance may also undergo a molecular +change, whereby the original composition of the shell or the hard part is +entirely altered. This tends almost invariably to harden the fossils still +further, which change of condition is called petrifaction, or the making +into stone. + +[Illustration: =Fig. 3. Thin Slice of Petrified or Silicified Wood in +Tangential Section.= + +Araucarioxylon Daintreei, Chapm. = Dadoxylon australe, Arber; × 28. +Carbopermian: Newcastle, New South Wales. + + (_Nat. Mus. Coll._) +] + + +=Structure Preserved.--= + +Petrifaction does not necessarily destroy the structure of a fossil. For +example, a piece of wood, which originally consisted of carbon, hydrogen, +and nitrogen, may be entirely replaced by flint or silica: and yet the +original structure of the wood may be so perfectly preserved that when +a thin slice of the petrifaction is examined under a high power of the +microscope, the tissues with their component cells are seen and easily +recognised (Fig. 3). + + +=Early Observers.--= + +Remains of animals buried in the rocks were known from the earliest times, +and frequent references to these were made by the ancient Greek and Roman +philosophers. + + +Xenophanes.-- + +Xenophanes, who lived B.C. 535, wrote of shells, fishes and seals which +had become dried in mud, and were found inland and on the tops of the +highest mountains. The presence of these buried shells and bones was +ascribed by the ancients to a plastic force latent in the earth itself, +while in some cases they were regarded as freaks of nature. + + +Leonardo da Vinci.-- + +In the sixteenth and seventeenth centuries Italian observers came to +the fore in clearly demonstrating the true nature of fossils. This was +no doubt due in part to the fact that the Italian coast affords a rich +field of observation in this particular branch of science. The celebrated +painter Leonardo da Vinci (early part of the sixteenth century), who +carried out some engineering works in connection with canals in the north +of Italy, showed that the mud brought down by rivers had penetrated into +the interior of shells at a time when they were still at the bottom of the +sea near the coast. + + +Steno.-- + +In 1669, Steno, a Danish physician residing in Italy, wrote a work on +organic petrifactions which are found enclosed in solid rocks, and showed +by his dissection of a shark which had been recently captured and by a +comparison of its teeth with those found fossil in the cliffs, that they +were identical. The same author also pointed out the resemblance between +the shells discovered in the Italian strata and those living on the +adjacent shores. It was not until the close of the eighteenth century, +however, that the study of fossil remains received a decided impetus. +It is curious to note that many of these later authors maintained the +occurrence of a universal flood to account for the presence of fossil +shells and bones on the dry land. + +[Illustration: =Fig. 4.--William Smith (1769-1839.)= + +"The Father of English Geology," at the age of 69. + + (_From Brit. Mus. Cat._) +] + + +=Fossils an Index to Age.--= + +A large part of the credit of showing how fossils are restricted to +certain strata, and help to fix the succession and age of the beds, is due +to the English geologist and surveyor, William Smith (Fig. 4). "The Father +of English Geology," as he has been called, published two works[1] in the +early part of last century, in which he expressed his view of the value of +fossils to the geologist and surveyor, and showed that there was a regular +law of superposition of one bed upon another, and that strata could be +identified at distant localities by their included fossils. Upon this +foundation the work of later geologists has been firmly established; and +students of strata and of fossils work hand in hand. + +[Footnote 1: "Strata identified by Organised Fossils," 1816-1819; and +"Stratigraphical System of Organised Fossils," 1817.] + + +=Stratigraphy.--= + +That branch of geology which discusses the nature and relations of the +various sediments of the earth's crust, and the form in which they were +laid down, is called Stratigraphy. From it we learn that in bygone times +many of those places that are now occupied by dry land have been, often +more than once, covered by the sea; and thus Tennyson's lines are forcibly +brought to mind-- + + "There where the long street roars hath been + The stillness of the central sea." + + +=Elevated Sea-beds.--= + +A striking illustration in proof of this emergence of the land from +the sea is the occurrence of marine shells similar to those now found +living in the sea, in sea-cliffs sometimes many hundreds of feet above +sea-level. When these upraised beds consist of shingle or sand with +shore-loving shells, as limpets and mussels, they are spoken of as Raised +Beaches. Elevated beaches are often found maintaining the same level along +coast-lines for many miles, like those recorded by Darwin at Chili and +Peru, or in the south of England (Fig. 5). They also occur intermittently +along the Victorian coast, especially around the indents, where they have +survived the wear and tear of tides along the coast line (Fig. 6). They +are also a common feature, as a capping, on many coral islands which have +undergone elevation. + +[Illustration: =Fig. 5.--A Raised Beach at Black Rock, Brighton, England.= + + (_Original_). +] + +[Illustration: =Fig. 6.--Raised Beach (a) and Native Middens (b)= + +Torquay, Victoria. + + (Original). +] + +[Illustration: =Fig. 7.--Marine Fossils (Orthis flabellulum, Sowerby.)= + +About nat. size. In Volcanic Tuff of Ordovician Age. From the Summit of +Snowdon, North Wales, at an elevation of 3571 feet above sea level. + + (_F.C. Coll._) +] + + +=Sea-beds far from the Present Coast.--= + +Marine beds of deeper water origin may be found not only close to the +coast-line, but frequently on the tops of inland hills some miles from +the sea-coast. Their included sea-shells and other organic remains are +often found covered by fine sediment forming extensive beds; and they +may frequently occur in the position in which they lived and died (Fig. +7). Although it is well known that sea-birds carry shell-fish for some +distance inland, yet this would not account for more than a few isolated +examples. + + +=Raised Beaches as Distinct from Middens.--= + +Again, it may be argued that the primitive inhabitants of countries +bordering the coast were in the habit of piling up the empty shells of +the edible molluscs used by them for food: but these "kitchen middens" +are easily distinguished from fossil deposits like shelly beaches, by the +absence of stratified layers; and, further, by the shells being confined +to edible species, as the Cockle (_Cardium_), the Blood-cockle (_Arca_), +the Mussel (_Mytilus_), and the Oyster (_Ostrea_) (Fig. 8). + +[Illustration: =Fig. 8.--Remains of Edible Shell Fish= +(Kitchen-midden--native, mirrn-yong) + +in Sand Dunes near Spring Creek, Torquay, Victoria. + + (_Original_). +] + +[Illustration: =Fig. 9.--Part of a Submerged Forest= + +seen at low water on the Cheshire coast at Leasowe, England. + + (_From Seward's "Fossil Plants"_) +] + + +=Submerged Forests.--= + +Evidence of change in the coast-line is shown by the occurrence of +submerged forest-land, known as "fossil forests," which consist of the +stumps of trees still embedded in the black, loamy soil. Such forests, +when of comparatively recent age, are found near the existing coast-line, +and may sometimes extend for a considerable distance out to sea (Fig. 9). + +From the foregoing we learn that:-- + + _1.--Fossils afford data of the various Changes that have taken place + in past times in the Relative Positions of Land and Water._ + +=Changes of Climate in the Past.--= + +At the present day we find special groups of animals (fauna), and plants +(flora), restricted to tropical climates; and others, conversely, to the +arctic regions. Cycads and tree-ferns, for example, seem to flourish best +in warm or sub-tropical countries: yet in past times they were abundant +in northern Europe in what are now temperate and arctic regions, as in +Yorkshire, Spitzbergen, and Northern Siberia, where indeed at one time +they formed the principal flora. + +The rein-deer and musk-sheep, now to be found only in the arctic regions, +once lived in the South of England, France and Germany. The dwarf willow +(_Salix polaris_) and an arctic moss (_Hypnum turgescens_), now restricted +to the same cold region, occur fossil in the South of England. + +In Southern Australia and in New Zealand, the marine shells which lived +during the earlier and middle Tertiary times belong to genera and species +which are indicative of a warmer climate than that now prevailing; this +ancient fauna being like that met with in dredging around the northern +coasts of Australia (Fig. 10.) + +[Illustration: =Fig. 10.--A Fossil Shell (Pecten murrayanus, Tate).= + +Of Oligocene to Lower Pliocene Age in Southern Australia; closely allied +to, if not identical with, a species living off the coast of Queensland. +About nat. size. + + (_F.C. Coll._) +] + +From the above evidence we may say that:-- + + _2.--Fossils teach us that in Former Times the Climate of certain + parts of the earth's surface was Different from that now existing._ + +=Fossils as Guides to Age of Strata.--= + +In passing from fossil deposits of fairly recent origin to those of older +date, we find the proportion of living species gradually diminish, being +replaced by forms now extinct. After this the genera themselves are +replaced by more ancient types, and if we penetrate still deeper into +the series of geological strata, even families and orders of animals and +plants give place to others entirely unknown at the present day. + +From this we conclude that:-- + + _3.--Fossil Types, or Guide Fossils, are of great value in indicating + the Relative Age of Geological Formations._ + +=Gradual Evolution of Life-forms from Lower to Higher Types.--= + +As a general rule the various types of animals and plants become simpler +in organisation as we descend the geological scale. For example, in the +oldest rocks the animals are confined to the groups of Foraminifera, +Sponges, Corals, Graptolites, Shell-fish and Trilobites, all back-boneless +animals: whilst it was not until the Devonian period that the primitive +fishes appeared as a well-defined group; and in the next formation, the +Carboniferous Series, the first traces of the Batrachians (Frog-like +animals) and Reptiles are found. Birds do not appear, so far as their +remains are known, until near the close of the Jurassic; whilst Mammals +are sparsely represented by Monotremes and Marsupials in the Triassic and +Jurassic, becoming more abundant in Cainozoic times, and by the Eutheria +(Higher Mammals) from the commencement of the Eocene period. + +It is clear from the above and other facts in the geological distribution +of animal types that:-- + + _4.--The Geological Record supports in the main the Doctrine of + Evolution from Simpler to more Complex types; and fossils throw much + light upon the Ancestry of Animals and Plants now found Living._ + + + + +CHAPTER II. + +THE CLASSIFICATION OF FOSSIL ANIMALS AND PLANTS. + + +An elementary knowledge of the principles underlying the classification of +animals and plants is essential to the beginner in the study of fossils. + + +=The Naming of Animals.--= + +In order to make a clearly understood reference to an animal, or the +remains of one, it is as necessary to give it a name as it is in the case +of a person or a place. Before the time of Linnaeus (1707-1778), it was +the custom to refer, for example, to a shell, in Latin[2] as "the little +spiral shell, with cross markings and tubercles, like a ram's horn;" or to +a worm as "the rounded worm with an elevated back." Improvements in this +cumbersome method of naming were made by several of the earlier authors +by shortening the description; but no strict rule was established until +the tenth edition of Linnaeus' "Systema Naturae" (1758), when that author +instituted his binomial nomenclature by giving each form enumerated both +a generic and specific name. In plain words, this method takes certain +life-forms closely related, but differing in minute particulars, and +places them together in a genus or kindred group. Thus the true dogs +belong to the genus _Canis_, but since this group also includes wolves, +jackals, and foxes, the various canine animals are respectively designated +by a specific name; thus the dog (_Canis familiaris_), the dingo (_C. +dingo_), the wolf (_C. lupus_), the jackal (_C. aureus_), and the fox (_C. +vulpes_). The generic name is placed first. Allied genera are grouped +in families, (for example, Canidae), these into orders (ex. Carnivora), +the orders into classes (ex. Mammalia), and the classes into phyla or +subkingdoms (ex. Vertebrata). + +[Footnote 2: The Latin description was used more commonly than it is at +present, as a universal scientific language.] + +Plants are classified in much the same way, with the exception that +families and orders are, by some authors, regarded as of equal value, or +even reversed in value; and instead of the term phylum the name series is +used. + + +Classification of the Animal Kingdom. + + NAME OF PHYLUM. | FORMS FOUND FOSSIL + ----------------------+------------------------------------------------- + I.--PROTOZOA | Foraminifera, Radiolaria. + | + II.--COELENTERATA | Sponges, Corals, Stromatoporoids, Graptolites. + | + III.--ECHINODERMATA | Crinoids, Starfishes, Brittle-stars, Sea-urchins. + | + IV.--VERMES | Worms (tube-making and burrowing kinds). + | + V.--MOLLUSCOIDEA | Polyzoa or Sea-mats, Brachiopods or Lamp-shells. + | + VI.--MOLLUSCA | Shell-fish: as Bivalves, Tusk-shells, + | Chitons or Mail-shells, Gasteropods or + | Snails, Pteropods or Sea-butterflies; + | Cuttle-fishes. + | + VII.--ARTHROPODA | Joint-footed animals: as Trilobites, Cyprids, + | Crabs and Lobsters, Centipedes, Spiders + | and Insects. + | + VIII.--VERTEBRATA | Fishes, Amphibians, Reptiles, Birds and Mammals. + + +Classification of Animal Kingdom. + +The first seven groups of the above classification are back-boneless +animals or Invertebrata; the eighth division alone comprising the animals +with a vertebra or backbone. + + +=Characters of the Several Phyla.--= + +In the first group are placed those animals which, when living, consist +of only one cell, or a series of similar cells, but where the cells were +never combined to form tissues having special functions, as in the higher +groups. + +PROTOZOA.-- + +The _Amoeba_ of freshwater ponds is an example of such, but owing to its +skin or cortex being soft, and its consequent inability to be preserved, +it does not concern us here. There are, however, certain marine animals +of this simple type of the Protozoa which secrete carbonate of lime to +form a chambered shell (Foraminifera); or silica to form a netted and +concentrically coated shell held together with radial rods (Radiolaria); +and both of these types are found abundantly as fossils. They are mainly +microscopic, except in the case of the nummulites and a few other kinds of +foraminifera, which are occasionally as large as a crown piece. + +COELENTERATA.-- + +The second group, the Coelenterata, shows a decided advance in +organisation, for the body is multicellular, and provided with a +body-cavity which serves for circulation and digestion. The important +divisions of this group, in which the organisms have hard parts capable of +being fossilised, are the limy and flinty Sponges, the Corals, and allied +groups, as well as the delicate Graptolites which often cover the surface +of the older slates with their serrated, linear forms, resembling pieces +of fret-saws. + +ECHINODERMATA.-- + +The third group, Echinodermata, comprises the Sea-lilies (Crinoids), +Starfishes and Sea-urchins, besides a few other less important types; and +all these mentioned are found living at the present day. Their bodies are +arranged in a radial manner, the skin being strengthened by spicules and +hardened by limy deposits ultimately forming plates. They have a digestive +canal and a circulatory system, and are thus one remove higher than the +preceding group. + +VERMES.-- + +The fourth group, Vermes (Worms), are animals with a bilateral or +two-sided body, which is sometimes divided into segments, but without +jointed appendages. Those which concern the student of fossils are the +tube-making worms, the errant or wandering worms which form casts like the +lob-worm, and the burrowing kinds whose crypts or dwellings become filled +with solid material derived from the surrounding mud. + +MOLLUSCOIDEA.-- + +Group five, the Molluscoidea, contains two types; the Flustras or Sea-mats +(Polyzoa) and the Lamp-shells (Brachiopoda). They are at first sight +totally unlike; for the first-named are colonies of compound animals, +and the second are simple, and enclosed between two valves. They show in +common, however, a bilateral symmetry. The mouth is furnished with fine +tentacles, or with spirally rolled hair-like or ciliated processes. + +MOLLUSCA.-- + +The sixth group, the Mollusca, includes all shell-fish. They are +soft-bodied, bilaterally symmetrical animals, without definite segments. +The shells, on account of being formed of carbonate of lime on an organic +basis, are often found preserved in fossiliferous strata. + +ARTHROPODA.-- + +The seventh group, the Arthropoda, or joint-footed animals, are +distinguished by their segmented, lateral limbs, and by having a body +composed of a series of segments or somites. The body and appendages are +usually protected by a horny covering, the 'exoskeleton.' The group of the +Trilobites played an important part in the first era of the formation of +the earth's crust; whilst the other groups were more sparsely represented +in earlier geological times, but became more and more predominant until +the present day. + +VERTEBRATA.-- + +The great group of the Vertebrata comes last, with its chief +characteristic of the backbone structure, which advances in complexity +from the Fishes to the Higher Mammals. + +=A Simplified Classification of the Vegetable Kingdom.= + + SERIES. | FORMS FOUND FOSSIL. + ---------------------+---------------------------------- + I.--THALLOPHYTA |Sea-weeds: as Corallines and + | Calcareous Algae. + | + II.--BRYOPHYTA |Mosses, Liverworts. + | + III.--PTERIDOPHYTA |Fern-like plants, as Horse-tails, + | Club-mosses and true Ferns. + | + IV.--PTERIDOSPERMEAE|Oldest Seed-bearing plants, + | with fern-like foliage. + | + V.--GYMNOSPERMEAE |Plants with naked seeds, as Cycads + | (Fern-palms), Ginkgo + | (Maiden-hair Tree), and + | Conifers (Pine trees). + | + VI.--ANGIOSPERMEAE |Flowering plants, as Grasses, + | Lilies and all ordinary trees + | and plants. + + +=Characters of the Plant Series.= + +THALLOPHYTA.-- + +The first series, the Thallophytes, are simple unicellular plants, and +occupy the same position in the vegetable kingdom as the Protozoa do in +the animal kingdom. Fossil remains of these organisms seem to be fairly +well distributed throughout the entire geological series, but, owing to +the soft structure of the fronds in most of the types, it is often a +matter of doubt whether we are dealing with a true thallophyte or not. +Many of the so-called sea-weeds (fucoids) may be only trails or markings +left by other organisms, as shell-fish and crustaceans. + +BRYOPHYTA.-- + +The second series, the Bryophytes or moss plants, are represented in the +fossil state by a few unimportant examples. + +PTERIDOPHYTA.-- + +The third series, the Pteridophytes, includes the Ferns found from +the Devonian up to the present day, Horse-tails and allied forms, +like _Equisetites_, and the Club-mosses and _Lepidodendron_ of the +Carboniferous period in various parts of the world. + +PTERIDOSPERMEAE.-- + +The fourth series, the Pteridospermeae, comprises some of the earliest +seed-bearing plants, as _Alethopteris_ and _Neuropteris_. They occur in +rocks of Upper Palaeozoic age as far as known. + +GYMNOSPERMEAE. + +The fifth series, the Gymnospermeae, contains the most important types of +plants found fossil, especially those of the primary and secondary rocks: +they were more abundant, with the exception of the Coniferae, in the +earlier than in the more recent geological periods. + +ANGIOSPERMEAE.-- + +The sixth series, the Angiospermeae, comprises all the Flowering Trees +and Plants forming the bulk of the flora now living, and is divided +into the kinds having single or double seed-leaves (Monocotyledones the +Dicotyledones respectively). This important group came into existence +towards the close of the Cretaceous period simultaneously with the higher +mammals, and increased in abundance until modern times. + + + + +CHAPTER III. + +THE GEOLOGICAL EPOCHS: AND THE TIME RANGE OF FOSSILS. + + +=Superposition of Strata.--= + +Fossils are chiefly found in rocks which have been formed of sediments +laid down in water, such as sandstone, shale and most limestones. These +rocks, broadly speaking, have been deposited in a horizontal position, +though really slightly inclined from shore to deep-water. One layer has +been formed above another, so that the oldest layer is at the bottom, and +the newest at the top, of the series (Fig. 11). Let us, for instance, +examine a cliff showing three layers: the lower, a sandstone, we will +Call A; the intermediate, a shale or clay bed, B; and the uppermost, a +limestone or marl, C (Fig. 12). In forming a conclusion about the relative +ages of the beds, we shall find that A is always older than B, and B than +C, provided no disturbance of the strata has taken place. For instance, +the beds once horizontally deposited may have been curved and folded +over, or even broken and thrust out of place, within limited areas; but +occurrences like these are extremely rare. Moreover, an examination of the +surrounding country, or of deep cuttings in the neighbourhood, will tell +us if there is any probability of this inversion of strata having taken +place. + +[Illustration: =Fig. 11.--Horizontal Layers of Fossiliferous Clays and +Sands.= + +In Sea Cliff, Torquay Coast, Victoria, looking towards Bird Rock. + + (_Original_). +] + +[Illustration: =Fig. 12.--Cliff-Section to Show Superposition of Strata.= + +A = Sandstone. B = Shale. C = Limestone.] + +This law of superposition holds good throughout the mass of sedimentary +rocks forming the crust of the earth. + +(1). Thus, the position of the strata shows the relative ages of the beds. + + +=Differences in Fossil Faunas.--= + +Turning once again to our ideal cliff section, if we examine the fossils +obtained from bed A, we shall find them differing in the number of kinds +or species common to the other beds above and below. Thus, there will +be more species alike in beds A and B or in B and C. In other words the +faunas of A and B are more nearly related than those of A and C. This is +explained by the fact that there is a gradual change in specific forms as +we pass through the time series of strata from below upwards; so that the +nearer one collecting platform is to another, as a rule, the stronger is +the community of species. + + +=Guide Fossils.--= + +Certain kinds of fossils are typical of particular formations. They are +known as guide fossils, and by their occurrence help us to gain some idea +of the approximate age of rocks widely separated by ocean and continent. +Thus we find fossils typical of the Middle Devonian rocks in Europe, which +also occur in parts of Australia, and we therefore conclude that the +Australian rocks containing those particular fossils belong to the same +formation, and are nearly of the same age. + +(2). The included fossils, therefore, give evidence of the age of the +beds. + + +=Value of Lithological Evidence.--= + +The test of age by rock-structure has a more restricted use, but is of +value when taken in conjunction with the sequence of the strata and the +character of their included fossils. + +To explain both the valuable and the uncertain elements of this last +method as a determinant of age, we may cite, for instance, the Upper +Ordovician slates of Victoria and New South Wales as an example of uniform +rock formation; whilst the yellow mudstones and the grey limestones of +the Upper Silurian (Yeringian series) of the same states, are instances +of diverse lithological structures in strata of similar age. A reference +in the latter case to the assemblages of fossils found therein, speedily +settles the question. + +(3). Hence, the structure and composition of the rocks (lithology), gives +only partial evidence in regard to age. + + +=Strata Vertically Arranged.--= + +The Stratigraphical Series of fossiliferous sediments comprises bedded +rocks from all parts of the world, which geologists arrange in a vertical +column according to age. + +A general computation of such a column for the fossiliferous rocks of +Europe gives a thickness of about 14 miles. This is equivalent to a mass +of strata lying edgewise from Melbourne to Ringwood. The Australian +sediments form a much thicker pile of rocks, for they can hardly fall +short of 37 miles, or nearly the distance from Melbourne to Healesville. + +This vertical column of strata was formed during three great eras of time. +The oldest is called the Primary or Palaeozoic ("ancient life"), in which +the animals and plants are of primitive types. This is followed by the +Secondary or Mesozoic ("middle life"), in which the animals and plants are +intermediate in character between the Palaeozoic and the later, Cainozoic. +The third era is the Tertiary or Cainozoic ("recent life"), in which the +animals and plants are most nearly allied to living forms. These great +periods are further subdivided into epochs, as the Silurian epoch; and +these again into stages, as the Yeringian stage. + + +Vertical Column of Fossiliferous Strata, Australia. + + ERA. | EPOCHS IN | EQUIVALENT STRATA + | EUROPE. | IN AUSTRALIA. + -------------+---------------+------------------------------- + | HOLOCENE | Dunes, Beaches, and Shell-beds + | | now forming. + | | + | PLEISTOCENE | Raised Beaches, River Terraces, + | | Swamp Deposits + | | with Diprotodon, Cave + | | Breccias, Helix Sandstone. + | | + CAINOZOIC | PLIOCENE | Upper.--Estuarine beds of + or | | bores in the Murray basin, + TERTIARY | | Marine beds of + (Note 1). | | Limestone Creek, Glenelg + | | River, Vic. (Werrikooian). + | | + | | Lower.--Kalimnan red + | | sands (terrestrial) and + | | shell marls (marine) of + | | Victoria, Deep Leads + | | (fluviatile) in part, Upper + | | Aldingan of South + | | Australia. + -------------+---------------+------------------------------- + CAINOZOIC | MIOCENE | Deep Leads in part: Leaf-beds + or | | of Bacchus Marsh, + TERTIARY | | Dalton and Gunning. + (Continued). | | Janjukian Series of C. + | | Otway, Spring Creek, and + | | Table Cape. Batesford + | | Limestone. Polyzoal + | | Rock of Mt. Gambier and + | | the Nullarbor Plains. + | | Older Cainozoic of Murray + | | basin, Lower Aldingan + | | Series of S. Australia, + | | Corio Bay and + | | Bairnsdale Series. + | | + | OLIGOCENE | Shelly clays and leaf-beds + | | of the Balcombian Series + | | at Mornington; also + | | Shell-marls and clays + | | with Brown Coal, Altona + | | Bay, and lower beds at + | | Muddy Creek, W. Vict. + | | + | EOCENE | Probably no representatives. + -------------+---------------+------------------------------- + | | + MESOZOIC | CRETACEOUS | Upper.--Leaf-beds of Croydon, + or | | Q. Desert Sandstone, + SECONDARY | | Q. Radiolarian Rock, N. + | | Territory. Gin-gin Chalk, + | | W.A. + | | + | | Lower.--Rolling Downs + | | Formn., Q. Lake Eyre + | | beds, S.A. + | | + | JURASSIC | Marine.--Geraldton, W.A. + | | + | | Freshwater.--Carbonaceous + | | sandstone of S. + | | Gippsland, the Wannon, + | | C. Otway and Barrabool + | | Hills. Ipswich Series, Q. + | | Mesozoic of Tasmania, + | | Talbragar beds, N.S.W. + | | + | TRIASSIC | Upper leaf-beds at Bald + | | Hill, Bacchus Marsh, Vict. + | | Hawkesbury Series (Parramatta + | | Shales, Hawkesbury + | | Sandstone, Narrabeen + | | beds), N.S.W. Burrum + | | Beds, Q. + -------------+---------------+------------------------------ + PALAEOZOIC | PERMIAN and | Carbopermian (Note 2), + or | CARBONIFEROUS,| Coal Measures of New + PRIMARY | UPPER | South Wales, W. Australia, + | | Queensland (Gympie + | | Series) and Tasmania. + | | Gangamopteris beds of + | | Bacchus Marsh, Vict. + | | Upper Carboniferous of + | | Clarence Town, N.S.W. + | CARBONIFEROUS,| Fish and Plant beds, + | LOWER | Mansfield, Vict. Grampian + | | sandstone; Avon + | | River sandstone, Vict. + | | (?) Star beds, Queensland. + | | Lepidodendron + | | beds of Kimberley, W.A. + | | (Note 3). + | DEVONIAN | Upper.--Sandstones of Iguana + | | Creek, with plant remains. + | | Lepidodendron + | | beds with Lingula, Nyrang + | | Creek, N.S. Wales. + | | Middle.--Fossiliferous marbles + | | and mudstones of + | | Buchan, Bindi and Tabberabbera, + | | Vict. Rocks + | | of the Murrumbidgee, + | | N.S. Wales, and of Burdekin, + | | Queensland. + | SILURIAN | Upper.--(Yeringian stage).--Lilydale, + | | Loyola, Thomson + | | River, and Waratah + | | Bay, Vict.; Bowning and + | | Yass (in part), N.S. + | | Wales; Queensland. + | | Lower (Melbournian + | | stage).--Melbourne, + | | Heathcote, Vict.; Bowning + | | and Yass (in part), + | | N.S. Wales. Gordon R. + | | Limestone. + | ORDOVICIAN, | Slates (graptolitic).--Victoria + | UPPER and | and New South + | LOWER | Wales. (?) Gordon River + | | Limestone, Tas., in part + | | (Note 4). Larapintine + | | series of Central Australia. + | CAMBRIAN | Mudstones and limestones + | | of Tasmania, + | | South Australia, Victoria + | | and W. Australia. + | PRE-CAMBRIAN | Fossiliferous rocks doubtful; + | | chiefly represented + | | by schistose and other + | | metamorphic rocks. + +1.--The classification of the Cainozoics as employed here is virtually +the same as given by McCoy in connection with his work for the Victorian +Geological Survey. The writer has obtained further evidence to support +these conclusions from special studies in the groups of the cetacea, +mollusca and the protozoa. The alternative classification of the +cainozoics as given by one or two later authors, introducing the useful +local terminology of Hall and Pritchard for the various stages or assises +is as follows:-- + + TATE AND DENNANT. | HALL AND PRITCHARD. + Stages. | Stages. + | + Werrikooian Pleistocene | Werrikooian Pliocene. + Pliocene | + | + Kalimnan Miocene | Kalimnan Miocene. + | + Janjukian (?) Oligocene | Balcombian Eocene. + | + Balcombian Eocene | Janjukian + | and + Aldingan Eocene | Aldingan Eocene. + (lower beds | in part + at that loc.) | + +2.--Or Permo-carboniferous. As the series is held by some authorities to +partake of the faunas of both epochs, it is preferable to use the shorter +word, which moreover gives the natural sequence. There is, however, strong +evidence in favour of using the term Permian for this important series. + +3.--Mr. W. S. Dun regards the _Lepidodendron_ beds of W. Australia, New +South Wales and Queensland as of Upper Devonian age. There is no doubt, +from a broad view of the whole question as to the respective age of +these beds in Australia, that the one series is continuous, and probably +represents the Upper Devonian and the Lower Carboniferous of the northern +hemisphere. + +4.--These limestones contain a fauna of brachiopods and corals which, at +present, seems to point to the series as intermediate between the older +Silurian and the Upper Ordovician. + +Vertical Column of Fossiliferous Strata, New Zealand. + + | EPOCHS IN | EQUIVALENT STRATA + ERA. | EUROPE. | IN NEW ZEALAND. + ------------+------------------+---------------------- + | HOLOCENE | River Alluvium. Beach + | | Sands and Gravel. + | | + CAINOZOIC | PLEISTOCENE | Raised Beaches. Older Gravel + or | | Drifts. + TERTIARY | | Moraines. Boulder Clays. + | | + | PLIOCENE | Upper.--Petane series. } + | | Lower.--Waitotara } Wanganui + | | and Awatere series. } system. + | | + | MIOCENE | Oamaru series. + | | + | OLIGOCENE | Waimangaroa series. + ------------+------------------+--------------------------- + | CRETACEOUS | Waipara series (of Hutton). + | | + MESOZOIC | JURASSIC | Mataura and Putataka + or | | series. + SECONDARY | | + | TRIASSIC | Wairoa, Otapiri and Kaihiku + | | series. + ------------+------------------+----------------------------- + | PERMIAN | Aorangi (unfossiliferous) + | | series. + | | + | (?)CARBONIFEROUS | Maitai series (with Spirifer + | | and Productus.) + | | + | | (?)Te Anau series (unfossiliferous). + PALAEOZOIC | | + or | SILURIAN | Wangapeka series. + PRIMARY | | + | ORDOVICIAN | Kakanui series (with Lower + | | Ordovician graptolite + | | facies). + | | + | CAMBRIAN | Unfossiliferous. Metamorphic + | | schists of the Maniototo + | | series. + +1.--Based for the most part, but with some slight modifications, on Prof. +J. Park's classification in "Geology of New Zealand," 1910. + +[Illustration: =Fig. 13.= + +Range-in-Time of Fossils in Australasian Sedimentary Rocks. + +_E.M., del._] +] + +[Illustration: =Fig. 14.--Skeleton of Diprotodon australis, Owen.= + +Uncovered in Morass at Lake Callabonna, South Australia. + + (_By permission of Dr. E. C. Stirling_). +] + + + + +CHAPTER IV. + +HOW FOSSILS ARE FOUND: AND THE ROCKS THEY FORM. + + +As already noticed, it is the hard parts of buried animals and plants that +are generally preserved. We will now consider the groups of organisms, +one by one, and note the particular parts of each which we may reasonably +expect to find in the fossil state. + +MAMMALS.--The bones and teeth: as the _Diprotodon_ remains of Lake +Callabonna in South Australia (Fig. 14), of West Melbourne Swamp, +Victoria, and the Darling Downs, Queensland. Rarely the skin, as in the +carcases of the frozen Mammoth of the tundras of Northern Siberia; or the +dried remains of the _Grypotherium_ of South American caves. + +[Illustration: =Fig. 15.--Bird Bones.= + +Exposed on Sand-blow at Seal Bay, King Island. + + (_Photo by C. L. Barrett_). +] + +[Illustration: =Fig. 16.--Impression of a Bird's Feather in Ironstone.= + +About 2/3 nat. size. Of Cainozoic (? Janjukian) Age. Redruth, Victoria. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 17.--Notochelone costata, Owen sp. (Anterior portion +of carapace.)= + +About 1/4 nat. size. A Marine Turtle from the Lower Cretaceous of Flinders +River, Queensland. + + (_Nat. Mus. Coll._) +] + +BIRDS:--Bones: as the Moa bones of New Zealand and the Emu bones of the +King Island sand-dunes (Fig. 15). Very rarely the impressions of the +feathers of birds are found, as in the ironstone occurring in the Wannon +district of Victoria (Fig. 16), and others in fine clays and marls on +the continent of Europe and in England. Fossil eggs of sea-birds are +occasionally found in coastal sand-dunes of Holocene age. + +REPTILES.--Skeletons of fossil turtles (_Notochelone_) are found in +Queensland (Fig. 17). Whole skeletons and the dermal armour (spines and +bony plates) of the gigantic, specialised reptiles are found in Europe, +North America, and in other parts of the world. + +FISHES.--Whole skeletons are sometimes found in sand and clay rocks, as +in the Trias of Gosford, New South Wales (Fig. 18), and in the Jurassic +of South Gippsland. The ganoid or enamel-scaled fishes are common fossils +in the Devonian and Jurassic, notably in Germany, Scotland and Canada: +and they also occur in the sandy mudstone of the Lower Carboniferous of +Mansfield, Victoria. + +INSECTS.--Notwithstanding their fragility, insects are often well +preserved as fossils, for the reason that their skin and wings consist +of the horny substance called chitin. The Tertiary marls of Europe are +very prolific in insect remains (Fig. 19). From the Miocene beds of +Florissant, Colorado, U.S.A., several hundred species of insects have been +described. + +[Illustration: =Fig. 18.= + +=A Fossil Fish with Ganoid Scales (Pristisomus crassus, A.S. Woodw.).= + +About 1/2 nat. size. Trias (Hawkesbury Series), of Gosford, New South +Wales. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig 19.--A Fossil Insect (Tipula sp.) in Amber.= + +Nat. size. Oligocene beds; Baltic Prussia. + + (_F.C. Coll._) +] + +[Illustration: =Fig. 20.--A Fossil Lobster (Thalassina emerii, Bell).= + +Slightly reduced. From the Pleistocene of Port Darwin, Northern Territory. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 21.--An Ammonite (Desmoceras flindersi, McCoy sp.)= + +Half nat. size. Showing complex sutures. L. Cretaceous: Marathon, Flinders +River, Queensland. + + (_Nat. Mus. Coll._) +] + +CRUSTACEA.--The outer crust, or exoskeleton, of these animals is often +hard, being formed of a compound of carbonate and phosphate of lime on +an organic, chitinous base. The earliest forms of this group were the +trilobites, commencing in Cambrian times, and of which there is a good +representative series in Australian rocks. Remains of crabs and lobsters +are found in the various Cainozoic deposits in Australia (Fig. 20), and +also in the Jurassic in other parts of the world. + +MOLLUSCA.--The Cuttle-fish group (Cephalopoda, "head-footed"), is well +represented by the Nautilus-like, but straight _Orthoceras_ shells +commencing in Ordovician times, and, in later periods, by the beautiful, +coiled Ammonites (Fig. 21). The true cuttle-fishes possess an internal +bone, the sepiostaire, which one may see at the present day drifted on to +the sand at high-water mark on the sea-shore. The rod-like Belemnites are +of this nature, and occur abundantly in the Australian Cretaceous rocks of +South Australia and Queensland (Fig. 22). + +[Illustration: =Fig. 22. Belemnites (Belemnites diptycha, McCoy).= + +1/3 nat. size. Lower Cretaceous. Central South Australia. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 23.--A Group of Lamp Shells (Magellania flavescens, +Lam. sp.)= + +Attached to a Polyzoan. + +About 1/3 nat. size. Dredged from Westernport, Victoria. + + (_C.J. Gabriel Coll._) +] + +Elephant-tusk shells (Scaphopoda) are frequent in our Tertiary beds: they +are also sparingly found in the Cretaceous, and some doubtful remains +occur in the Palaeozoic strata of Australia. + +The shells of the ordinary mollusca, such as the snails, whelks, mussels, +and scallops, are abundant in almost all geological strata from the +earliest periods. Their calcareous shells form a covering which, after the +decay of the animal within, are from their nature among the most easily +preserved of fossil remains. There is hardly an estuary bed, lake-deposit, +or sea-bottom, but contains a more or less abundant assemblage of these +shell-fish remains, or testacea as they were formerly called ("testa," a +shell or potsherd). We see, therefore, the importance of this group of +fossils for purposes of comparison of one fauna with another (_antea_, +Fig. 1). + +The chitons or mail-shells, by their jointed nature, consisting of a +series of pent-roof-shaped valves united by ligamental tissue, are nearly +always represented in the fossil state by separate valves. Fossil examples +of this group occur in Australia both in Palaeozoic rocks and, more +numerously, in the Cainozoic series. + +[Illustration: =Fig. 24.--Zoarium of a Living Polyzoan. (Retepora)= + +2/3 nat. size. + +Flinders, Victoria. + + (_F.C. Coll._) +] + +[Illustration: =Fig. 25.--A Fossil Polyzoan (Macropora clarkei, T. Woods, + sp.)= + +About 1/2 nat. size. Cainozoic (Balcombian). Muddy Creek, Victoria. + + (_F.C. Coll._) +] + + +MOLLUSCOIDEA.--The Brachiopods or Lamp-shells consist generally of two +calcareous valves as in the true mollusca (Fig. 23), but are sometimes of +horny texture. Like the previous class, they are also easily preserved +as fossils. They possess bent, loop-like or spiral arms, called brachia, +and by the movement of fine ciliated (hair-like) processes on their +outer edges conduct small food particles to the mouth. The brachia are +supported by shelly processes, to which are attached, in the Spirifers, +delicate spirally coiled ribbons. These internal structures are often +beautifully preserved, even though they are so delicate, from the fact +that on the death of the animal the commissure or opening round the valves +is so tightly closed as to prevent the coarse mud from penetrating while +permitting the finer silt, and more rarely mineral matter in solution, to +pass, and subsequently to be deposited within the cavity. At the Murray +River cliffs in South Australia, a bed of Cainozoic limestone contains +many of these brachiopod shells in a unique condition, for the hollow +valves have been filled in with a clear crystal of selenite or gypsum, +through which may be seen the loop or brachial support preserved in its +entirety. + +The Sea-mats or Polyzoa, represented by _Retepora_ (the Lace-coral) +(Fig. 24) and _Flustra_ (the Sea-mat) of the present sea-shore, have a +calcareous skeleton, or zoarium, which is easily preserved as a fossil. +Polyzoa are very abundant in the Cainozoic beds of Australia, New Zealand, +and elsewhere (Fig. 25). In the Mesozoic series, on the other hand, they +are not so well represented; but in Europe and North America they play an +important part in forming the Cretaceous and some Jurassic strata by the +abundance of their remains. + +WORMS (VERMES).--The hard, calcareous tubes of Sea-worms, the Polychaeta +("many bristles") are often found in fossiliferous deposits, and sometimes +form large masses, due to their gregarious habits of life; they also occur +attached to shells such as oysters (Fig. 26). The burrows of the wandering +worms are found in Silurian strata in Australia; and the sedentary forms +likewise occur from the Devonian upwards. + +ECHINODERMATA.--Sea-urchins (Echinoidea) possess a hard, calcareous, +many-plated test or covering and, when living are covered with spines +(Fig. 27). Both the tests and spines are found fossil, the former +sometimes whole when the sediment has been quietly thrown down upon them; +but more frequently, as in the Shepherd's crown type (_Cidaris_), are +found in disjointed plates, owing to the fact that current action, going +on during entombment has caused the plates to separate. The spines are +very rarely found attached to the test, more frequently being scattered +through the marl or sandy clay in which the sea-urchins are buried. The +best conditions for the preservation of this group is a marly limestone +deposit, in which case the process of fossilisation would be tranquil +(Fig. 28). + +[Illustration: =Fig. 26.--Fossil Worm Tubes (? Serpula.)= + +Attached to a Pecten. + +Slightly Enlarged. Cainozoic (Balcombian). Muddy Creek, Hamilton, Victoria. + + (_F. C. Coll._) +] + +[Illustration: =Fig. 27.= + +=A Regular Sea-Urchin (Strongylocentrotus erythrogrammus, Val.)= + +About 2/3 nat. size. Showing Spines attached. Living. Victoria. + + (_F. C. Coll._) +] + +[Illustration: =Fig. 28.--A Fossil Sea-Urchin (Linthia antiaustralis, +Tate).= + +Test denuded of Spines. + +About 2/3 nat. size. Cainozoic (Janjukian): Curlewis, Victoria. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 29.--Ophioderma egertoni, Broderip, sp.= + +About 1/2 nat. size. A Brittle Star from the Lias of Seaton, Devon, +England. + + (_Nat. Mus. Coll._) +] + +The true Starfishes (Asteroidea), are either covered with calcareous +plates, or the skin is hardened by rough tubercles; and these more lasting +portions are preserved in rocks of all ages. The shape of the animal is +also often preserved in an exquisite manner in beds of fine mud or clay. + +The Brittle-stars (Ophiuroidea) have their body covered with hard, +calcareous plates. Their remains are found in rocks as old as the +Ordovician in Bohemia but their history in Australia begins with the +Silurian period (Fig. 29). From thence onward they are occasionally found +in successive strata in various parts of the world. + +The bag-like echinoderms (Cystidea) form a rare group, restricted to +Palaeozoic strata. The plates of the sack, or theca, and those of the +slender arms are calcareous, and are capable of being preserved in the +fossil state. A few doubtful remains of this group occur in Australia. + +The bud-shaped echinoderms (Blastoidea) also occur chiefly in Devonian and +Carboniferous strata. This is also a rare group, and is represented by +several forms found only in New South Wales and Queensland. + +The well known and beautiful fossil forms, the Stone-lilies (Crinoidea) +have a very extended geological history, beginning in the Cambrian; whilst +a few species are living in the ocean at the present day. The many-jointed +skeleton lends itself well to fossilisation, and remains of the crinoids +are common in Australia mainly in Palaeozoic strata (Fig. 30). In Europe +they are found abundantly also in Jurassic strata, especially in the Lias. + +[Illustration: =Fig. 30.= + +=A Fossil Crinoid (Taxocrinus simplex, Phillips sp.)= + +About 1/2 nat. size. + +Wenlock Limestone (Silurian), Dudley, England. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 31.--Graptolites on Slate (Tetragraptus fruticosus, +J. Hall, sp.)= + +Nat. Size. Lower Ordovician. Bendigo, Victoria. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 32.= + +=Polished Vertical Section of a Stromatoporoid. (Actinostroma).= + +Nat. size. Middle Devonian. South Devon, England. + + (_F.C. Coll._) +] + +HYDROZOA.--The Graptolites ("stone-writing") have a chitinous skin +(periderm) to the body or hydrosome, which is capable of preservation to +a remarkable degree; for their most delicate structures are preserved on +the surfaces of the fine black mud deposits which subsequently became +hardened into slates. In Australia graptolites occur from the base of the +Ordovician to the top of the Silurian (Fig. 31). + +Another section of the Hydrozoa is the Stromatoporoidea. These are +essentially calcareous, and their structure reminds one of a dense coral. +The polyps build their tiers of cells (coenosteum) in a regular manner, +and seem to have played the same part in the building of ancient reefs in +Silurian, Devonian and Carboniferous times as the Millepora at the present +day (Fig. 32). + +[Illustration: =Fig. 33.--Fossil Corals (Favosites).= + +Photograph of a Polished Slab, 2/3 nat. size. In Devonian Limestone, +Buchan, Victoria.] + +[Illustration: =Fig. 34.--Siliceous Skeleton of a Living Hexactinellid +Sponge.= + +Probably Chonelasma. + +× 4. Mauritius. (Viewed in Two Directions.) + + (_F.C. Coll._) +] + +ANTHOZOA.--The true Corals have a stony skeleton, and this is capable of +easy preservation as a fossil. There is hardly any fossiliferous stratum +of importance which has not its representative corals. In Australia their +remains are especially abundant in the Silurian, Devonian (Fig. 33), and +Carboniferous formations, and again in the Oligocene and Miocene. + +SPONGES.--The framework of the sponge may consist either of flinty, +calcareous, or horny material (Fig. 34). The two former kinds are well +represented in our Australian rocks, the first appearing in the Lower +Ordovician associated with graptolites, and again in the Cretaceous and +Tertiary rocks (Fig. 35); whilst the calcareous sponges are found in +Silurian strata, near Yass, and again in the Cainozoic beds of Flinders, +Curlewis and Mornington in Victoria. + +[Illustration: =Fig. 35.= + +=Spicules of a Siliceous Sponge (Ecionema newberyi, McCoy sp.)= + +Highly magnified. Cainozoic Shell-Marl. + +Altona Bay Coal-Shaft.] + +[Illustration: =Fig. 36.= + +=Nummulites (N. gizehensis Ehr. var. champollioni, de la Harpe).= + +About nat. size. Middle Eocene Limestone. Cyrene, Northern Africa. + + (_Coll. by Dr. J. W. Gregory_). +] + +PROTOZOA.--The important and widely-distributed group of the Foraminifera +("hole-bearers") belonging to the lowest phylum, the Protozoa, generally +possess a calcareous shell. The tests range in size from tiny specks of +the fiftieth of an inch in diameter, to the giant Nummulite, equalling a +five shilling piece in size (Fig. 36). Their varied and beautiful forms +are very attractive, but their great interest lies in their multifarious +distribution in all kinds of sediments: they are also of importance +because certain of the more complex forms indicate distinct life zones, +being restricted to particular strata occurring in widely-separated areas. + +[Illustration: =Fig. 37.--Siliceous Skeletons of Radiolaria.= + +× 58. Late Cainozoic Age. Bissex Hill, Barbados, West Indies. + + (_F.C. Coll._) +] + +Members of the allied order of the Radiolaria have a flinty shell (Fig. +37); and these organisms are often found building up siliceous rocks such +as cherts (Fig. 38). + +PLANTS.--The harder portions of plants which are found in the fossil +state are,--the wood, the coarser vascular (vessel-bearing) tissue of the +leaves, and the harder parts of fruits and seeds. + +Fossil wood is of frequent occurrence in Palaeozoic, Mesozoic and +Cainozoic strata in Australia, as, for instance, the wood of the trees +called _Araucarioxylon_ and _Dadoxylon_ in the Coal measures of New South +Wales (see _antea_, Fig. 3). + +[Illustration: =Fig. 38.--Radiolaria in Siliceous Limestone.= + +× 40. Middle Devonian: Tamworth, New South Wales. + + (_From Prof. David's Collection_). +] + +[Illustration: =Fig. 39.--Travertin Limestone with Leaves of Beech +(Fagus).= + +Nat. size. Pleistocene: near Hobart, Tasmania. + + (_Nat. Mus. Coll._) +] + +Fossil leaves frequently occur in pipe-clay beds, as at Berwick, Victoria, +and in travertine from near Hobart, Tasmania (Fig. 39). Fossil fruits are +found in abundance in the ancient river gravels at several hundreds of +feet below the surface, in the "deep leads" of Haddon, Victoria, and other +localities in New South Wales, Queensland and Tasmania. + +[Illustration: =Fig. 40--Freshwater Limestone with Shells (Bulinus).= + +About 4/5 nat. size. Mount Arapiles, Western Victoria. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 41.--Fossiliferous Mudstone of Silurian (Yeringian) +Age.= + +With Brachiopods. About 2/3 nat. size. Near Lilydale, Victoria. + + (_F.C. Coll._) +] + + +FOSSILIFEROUS ROCKS. + + +Section I.--ARGILLACEOUS ROCKS. + +Under this head are placed the muds, clays, mudstones, shales and slates. +MUDS are usually of a silty nature, that is, containing a variable +proportion of sand (quartz) grains. Such are the estuarine muds of +Pleistocene and Recent age, containing brackish water foraminifera and +ostracoda, and those shells of the mollusca usually found associated with +brackish conditions. Lacustrine mud can be distinguished by the included +freshwater shells, as _Limnaea_, _Coxiella_ (brackish), _Cyclas_ and +_Bulinus_, as well as the freshwater ostracoda or cyprids (Fig. 40). + +CLAYS are tenacious mud deposits, having the general composition of a +hydrous silicate of alumina with some iron. When a clay deposit tends to +split into leaves or laminae, either through moderate pressure or by the +included fossil remains occupying distinct planes in the rock, they are +called SHALES. + +Clays and Shales of marine origin are often crowded with the remains +of mollusca. The shells are sometimes associated with leaves and other +vegetable remains, if forming part of an alternating series of freshwater +and marine conditions. An example of this type of sediments is seen in the +Mornington beds of the Balcombian series in Victoria. + +MUDSTONE is a term applied to a hardened clay deposit derived from the +alteration of an impure limestone, and is more often found in the older +series of rocks. Mudstones are frequently crowded with fossils, but +owing to chemical changes within the rock, the calcareous organisms are +as a rule represented by casts and moulds. At times these so faithfully +represent the surface and cavities of the organism that they are almost +equivalent to a well preserved fossil (Fig. 41). + +SLATE.--When shale is subjected to great pressure, a plane of regular +splitting called cleavage is induced, which is rarely parallel to the +bedding plane or surface spread out on the original sea-floor: the +cleavage more often taking place at an appreciable angle to the bedding +plane. The graptolitic rocks of Victoria are either shales or slates, +according to the absence or development of this cleavage structure in the +rock. + + +Section II.--SILICEOUS ROCKS. + +In this group are comprised all granular quartzose sediments, and organic +rocks of flinty composition. + +SANDSTONES.--Although the base of this type of rock is formed of quartz +sand, it often contains fossils. Owing to its porous nature, percolation +of water containing dissolved CO_{2} tends to bring about the solution +of the calcareous shells, with the result that only casts of the shells +remain. + +FLINTS and CHERTS.--These are found in the form of nodules and bands in +other strata, principally in limestone. In Europe, flint is usually found +in the Chalk formation, whilst chert is found in the Lower Greensands, +the Jurassics, the Carboniferous Limestone and in Cambrian rocks. In +Australia, flint occurs in the Miocene or Polyzoal-rock formation of Mount +Gambier, Cape Liptrap and the Mallee borings. Flint is distinguished +from chert by its being black in the mass, often with a white crust, +and translucent in thin flakes; chert being more or less granular in +texture and sub-opaque in the mass. Both kinds appear to be formed as +a pseudomorph or replacement of a portion of the limestone stratum by +silica, probably introduced in solution as a soluble alkaline silicate. +Both flint and chert often contain fossil shells and other organic +remains, such as radiolaria and sponge-spicules, which can be easily seen +with a lens in thin flakes struck off by the hammer. + +DIATOMITE is essentially composed of the tiny frustules or flinty cases +of diatoms (unicellular algae), usually admixed with some spicules of the +freshwater sponge, _Spongilla_. It generally forms a layer at the bottom +of a lake bed (Fig. 42). + +[Illustration: =Fig. 42.--Diatomaceous Earth. (Post-Tertiary).= + +Containing freshwater forms, as Pinnularia, Cocconeis and Synedra. × 150. +Talbot, Victoria.] + + +Section III.--CALCAREOUS ROCKS. + +LIMESTONES FORMED BY ORGANISMS.--Organic limestones constitute by far +the most important group of fossiliferous rocks. Rocks of this class are +composed either wholly of carbonate of lime, or contain other mineral +matter also, in varying proportion. Many kinds of limestones owe their +origin directly to the agency of animals or plants, which extracted the +calcareous matter from the water in which they lived in order to build +their hard external cases, as for example the sea-urchins; or their +internal skeletons, as the stony corals. The accumulated remains of these +organisms are generally compacted by a crystalline cement to form a +coherent rock. + +The chief groups of animals and plants forming such limestone rocks are:-- + +(a) _FORAMINIFERA._--Example. Foraminiferal limestone as the Nummulitic +limestone of the Pyramids of Egypt, or the _Lepidocyclina_ limestone of +Batesford, near Geelong, Victoria (Fig. 43). + +[Illustration: =Fig. 43.= + +=Limestone composed of Polyzoa and Foraminifera (Lepidocyclina).= + +× 6. Cainozoic (Janjukian). Batesford, near Geelong, Victoria. + + (_F.C. Coll._) +] + +(b) _CORALS._--Ex. "Madrepore limestone," or Devonian marble, with +_Pachypora_. Also the Lilydale limestone, with _Favosites_, of Silurian +age, Victoria (Fig. 44). + +[Illustration: =Fig. 44.--A Fossil Coral (Favosites grandipora).= + +2/3 nat. size. From the Silurian of Lilydale, Victoria. + + (_F.C. Coll._). +] + +[Illustration: =Fig. 45.--Polished Slab of Marble formed of Joints of +Crinoids.= + +About 2/3 nat. size. Silurian. + +Toongabbie, Gippsland, Victoria. + + (_Nat. Mus. Coll._) +] + +(c) _STONE-LILIES._--Ex. Crinoidal or Entrochial limestone, Silurian, +Toongabbie, Victoria (Fig. 45). Also the Carboniferous or Mountain +limestone, Derbyshire, England. + +(d) _WORM-TUBES.-_-Ex. Serpulite limestone of Hanover, Germany. _Ditrupa_ +limestone of Torquay and Wormbete Creek, Victoria. + +(e) _POLYZOA._---Ex. Polyzoal limestone, as the so-called Coralline Crag +of Suffolk, England; and the Polyzoal Rock of Mount Gambier, S. Australia. + +(f) _BRACHIOPODA._--Ex. Brachiopod limestone of Silurian age, Dudley, +England. _Orthis_ limestone of Cambrian age, Dolodrook River, N. E. +Gippsland. + +(g) _MOLLUSCA._--Ex. Shell limestone, as the _Turritella_ bed of Table +Cape, Tasmania, and of Camperdown, Victoria (Fig. 46), or the Purbeck +Marble of Swanage, Dorset, England. + +[Illustration: =Fig. 46.--Turritella Limestone.= + +(T. acricula, Tate); 3/4 nat. size. Cainozoic. + +Lake Bullen Merri, near Camperdown, Victoria.] + +[Illustration: =Fig. 47.--Limestone composed of the Valves of an Ostracod +(Cypridea).= + +Upper Jurassic. × 9. + +Swanage, Dorset, England.] + +(h) _OSTRACODA._--Ex. Cypridiferous limestone, formed of the minute valves +of the bivalved ostracoda, as that of Durlston, Dorset, England (Fig. 47). + +(i) _CADDIS FLY LARVAE._--Ex. Indusial limestone, formed of tubular cases +constructed by the larvae of the Caddis fly (_Phryganea_). Occurs at +Durckheim, Rhine District, Germany. + +(j) _RED SEAWEEDS._--Ex. Nullipore limestone, formed by the stony thallus +(frond) of the calcareous sea-weed _Lithothamnion_, as in the Leithakalk, +a common building stone of Vienna. + +(k) _GREEN SEAWEEDS._--Ex. _Halimeda_ limestone, forming large masses of +rock in the late Cainozoic reefs of the New Hebrides (Fig. 48). + +(l) (?) _BLUE-GREEN SEAWEEDS._--Ex. _Girvanella_ limestone, forming the +Peagrit of Jurassic age, of Gloucester, England. + + +Section IV.--CARBONACEOUS and MISCELLANEOUS ROCKS. + +COALS and KEROSENE SHALES (Cannel Coal).--These carbonaceous rocks are +formed in much the same way as the deposits in estuaries and lagoon +swamps. They result from the sometimes vast aggregation of vegetable +material (leaves, wood and fruits), brought down by flooded rivers from +the surrounding country, which form a deposit in a swampy or brackish +area near the coast, or in an estuary. Layer upon layer is thus formed, +alternating with fine mud. The latter effectually seals up the organic +layers and renders their change into a carbonaceous deposit more certain. + +When shale occurs between the coal-layers it is spoken of as the +under-clay, which in most cases is the ancient sub-soil related to the +coal-layer immediately above. It is in the shales that the best examples +of fossil ferns and other plant-remains are often found. The coal itself +is composed of a partially decomposed mass of vegetation which has become +hardened and bedded by pressure and gradual drying. + +Spore coals are found in thick deposits in some English mines, as at +Burnley in Yorkshire. They result from the accumulation of the spores of +giant club-mosses which flourished in the coal-period. They are generally +referred to under the head of Cannel Coals. The "white coal" or Tasmanite +of the Mersey Basin in Tasmania is an example of an impure spore coal with +a sandy matrix (Fig. 49). + +[Illustration: =Fig. 48.= + +=Rock composed of the calcareous joints of Halimeda (a green sea-weed).= + +About 2/3 nat. size. Late Cainozoic. Reef-Rock. Malekula, New Hebrides. + + (_Coll. by Dr. D. Mawson._) +] + +[Illustration: =Fig. 49.--Thin Slices of "White Coal" or "Tasmanite," +showing crushed Megaspores.= + +× 28. Carbopermian. Latrobe, Tasmania. + + (_F. C. Coll._) +] + +[Illustration: =Fig. 50.--Thin Slice of "Kerosene Shale."= + +× 28. Carbopermian. Hartley, New South Wales. + + (_F. C. Coll._) +] + +[Illustration: =Fig. 51.--Bone Bed, with Fish and Reptilian Remains.= + +About 1/2 nat. size. (Rhaetic). Aust Cliff, Gloucestershire, England. + + (_Nat. Mus. Coll._) +] + +The Kerosene Shale of New South Wales is related to the Torbanite of +Scotland and Central France. It occurs in lenticular beds between the +bituminous coal. It is a very important deposit, commercially speaking, +for it yields kerosene oil, and is also used for the manufacture of gas. +The rock is composed of myriads of little cell-bodies, referred to as +_Reinschia_, and first supposed to be allied to the freshwater alga, +_Volvox_; but this has lately been questioned, and an alternative view is +that they may be the megaspores of club-mosses (Fig. 50). + +The coals of Jurassic age in Australia are derived from the remains of +coniferous trees and ferns; and some beautiful examples of these plants +may often be found in the hardened clay or shale associated with the coal +seams. + +The Brown Coals of Cainozoic or Tertiary age in Australia are still but +little advanced from the early stage, lignite. The leaves found in them +are more or less like the present types of the flora. The wood is found +to be of the Cypress type (_Cupressinoxylon_). In New Zealand, however, +important deposits of coal of a more bituminous nature occur in the +Oligocene of Westport and the Grey River Valley, in the Nelson District. + +BONE BEDS.--The bones and excreta of fish and reptiles form considerable +deposits in some of the sedimentary formations; especially those partly +under the influence of land or swamp conditions. They constitute a kind of +conglomerate in which are found bone-fragments and teeth (Fig. 51). These +bone-beds are usually rich in phosphates, and are consequently valuable +as a source of manure. The Miocene bone-bed with fish teeth at Florida, +U.S.A., is a notable example. The nodule bed of the Victorian Cainozoics +contains an assemblage of bones of cetaceans (whales, etc.). + +[Illustration: =Fig. 52.--Bone Breccia, with remains of Marsupials.= + +About 3/4 nat. size. Pleistocene. + +Limeburners Point, Geelong, Victoria. + + (_Nat. Mus. Coll._) +] + +BONE BRECCIAS.--These are usually formed of the remains of the larger +mammals, and consist of a consolidated mass of fragments of bones and +teeth embedded in a calcareous matrix. Bone-breccias are of frequent +occurrence on the floors of caves which had formerly been the resort of +carnivorous animals, and into which they dragged their prey. The surface +water percolating through the overlying calcareous strata dissolved a +certain amount of lime, and this was re-deposited on the animal remains +lying scattered over the cave floor. A deposit so formed constitutes a +stalagmite or floor encrustation. As examples of bone-breccias we may +refer to the limestone at Limeburners Point, Geelong (Fig. 52); and the +stalagmitic deposits of the Buchan Caves. + +IRONSTONE.--Rocks formed almost entirely of limonite (hydrated peroxide of +iron) are often due to the agency of unicellular plants known as diatoms, +which separate the iron from water, and deposit it as hydrous peroxide +of iron within their siliceous skeletons. In Norway and Sweden there are +large and important deposits of bog iron-ore, which have presumably been +formed in the beds of lakes. + +[Illustration: =Fig. 53.= + +=Cainozoic Ironstone with Leaves (Banksia ? marginata, Cavanilles).= + +Slightly enlarged. Below Wannon Falls, Redruth, Victoria.] + +Clay ironstone nodules (sphaerosiderite) have generally been formed +as accretions around some decaying organic body. Many clay ironstone +nodules, when broken open, reveal a fossil within, such as a coprolitic +body, fern frond, fir-cone, shell or fish. + +Oolitic ironstones are composed of minute granules which may have +originally been calcareous grains, formed by a primitive plant or alga, +but since replaced by iron oxide or carbonate. + +The Tertiary ironstone of western Victoria is found to contain leaves, +which were washed into lakes and swamps (Fig. 53); and the ferruginous +groundmass may have been originally due to the presence of diatoms, though +this yet remains to be proved. + + + + +PART II.--SYSTEMATIC PALAEONTOLOGY. + + + + +CHAPTER V. + +FOSSIL PLANTS. + + +=Cambrian Plants.--= + +The oldest Australian plant-remains belong to the genus _Girvanella_. +This curious little tubular unicellular organism, once thought to be a +foraminifer, shows most affinity with the blue-green algae (Cyanophyceae), +an important type of plant even now forming calcareous deposits such +as the calcareous grains on the shores of the Salt Lake, Utah, and the +pea-grit of the Carlsbad hot springs. _Girvanella problematica_ occurs +in the Lower Cambrian limestones of South Australia, at Ardrossan and +elsewhere. + + +=Silurian Plants.--= + +Amongst Silurian plants may be mentioned the doubtful sea-weeds known as +_Bythotrephis_. Their branch-like impressions are fairly common in the +mudstones of Silurian age found in and around Melbourne. They generally +occur in association with shallow-water marine shells and crustacea of +that period. + +The genus _Girvanella_ before mentioned is also found in the Silurian +(Yeringian) of Lilydale and the Tyers River limestone, Victoria (Fig. 54). + +[Illustration: =Fig. 54.--Section through pellet of Girvanella conferta=, +Chapm. + +× 35. From the Silurian (Yeringian) Limestone of Tyers River, Gippsland, +Victoria. + + (_Nat. Mus. Coll._) +] + +_Haliserites_ is a primitive plant of the type of the club-mosses so +common in the rocks of the Carboniferous period. This genus is found in +some abundance in the Yeringian stage of the Silurian in Gippsland (Fig. +55). + +[Illustration: =Fig. 55.--PALAEOZOIC PLANTS.= + +(Approximate dimensions in fractions). + + A--Bythotrephis tenuis, J. Hall. Silurian. Victoria. + + B--Haliserites Dechenianus, Göppert. Silurian. Victoria. + + C--Cordaites australis, McCoy. Upper Devonian. Victoria. + + D--Sphenopteris iguanensis, McCoy. Upper Devonian. Victoria. + + E--Glossopteris Browniana, Brongniart. Carbopermian. N.S.W. + +] + +[Illustration: =Fig. 56.= + +=Restoration of Lepidodendron elegans.= + + (_After Grand'Eury._) +] + +[Illustration: =Fig. 57.= + +=Lepidodendron australe, McCoy.= + +Portion of Stem showing Leaf-cushions. + +Slightly reduced. + +Carboniferous. + +Manilla River, Co. Darling, N.S.W. + + (_Nat. Mus. Coll._) +] + + +=Devonian and Carboniferous Plants.--= + +[Illustration: =Fig. 58.--UPPER PALAEOZOIC PLANTS.= + +A--Rhacopteris inaequilatera, Göppert sp. Up. Carboniferous. Stroud, New +South Wales. + + (_After Feistmantel_). + +B--Gangamopteris spatulata, McCoy. Carbopermian. Bacchus Marsh, Victoria.] + +Plant-life was not abundant, however, until Upper Devonian and +Carboniferous times. In the rocks of these periods we meet with the large +strap-shaped leaves of _Cordaites_ and a fern, _Sphenopteris_, in the +first-named series; and the widely distributed _Lepidodendron_ with its +handsome lozenge-scarred stems in the later series (Fig. 56). _Cordaites_ +has been found in Victoria in the Iguana Creek beds (Upper Devonian), +and it also probably occurs at the same horizon at Nungatta, New South +Wales. _Lepidodendron_ occurs in the Lower Carboniferous sandstone of +Victoria and Queensland (Fig. 57): in New South Wales it is found at Mt. +Lambie, Goonoo, Tamworth and Copeland in beds generally regarded as Upper +Devonian. Both of these plants are typical of Carboniferous (Coal Measure) +beds in Europe and North America. The fern _Rhacopteris_ is characteristic +of Upper Carboniferous shales and sandstones near Stroud, and other +localities in New South Wales as well as in Queensland (Fig. 58). These +beds yield a few inferior seams of coal. _Girvanella_ is again seen in the +oolitic limestones of Carboniferous age in Queensland and New South Wales. + + +=Carbopermian Plants.--= + +The higher division of the Australian Carboniferous usually spoken of +as the Permo-carboniferous, and here designated the Carbopermian (see +Footnote 2, page 48), is typified by a sudden accession of plant forms, +chiefly belonging to ferns of the _Glossopteris_ type. The lingulate +or tongue-shaped fronds of this genus, with their characteristic +reticulate venation, are often found entirely covering the slabs of +shale intercalated with the coal seams of New South Wales; and it is +also a common fossil in Tasmania and Western Australia. The allied form, +_Gangamopteris_, which is distinguished from _Glossopteris_ by having no +definite midrib, is found in beds of the same age in Victoria, New South +Wales, and Tasmania. These plant remains are also found in India, South +Africa, South America and the Falkland Islands. This wide distribution +of such ancient ferns indicates that those now isolated land-surfaces +were once connected, forming an extensive continent named by Prof. Suess +"Gondwana-Land," from the Gondwana district in India (Fig. 59). + +[Illustration: + + _E. M. del._ (_After J. W. Gregory_). + + =Fig. 59.--Map of the World in the Upper Carboniferous Era.= +] + + +=Triassic Plants.--= + +The widely distributed pinnate fern known as _Thinnfeldia_ is first found +in the Trias; in the Narrabeen shales near Manly, and the Hawksbury +sandstone at Mount Victoria, New South Wales. It is also a common +fossil of the Jurassic of South Gippsland, and other parts of Victoria. +The grass-like leaves of _Phoenicopsis_ are frequently met with in +Triassic strata, as in the upper series at Bald Hill, Bacchus Marsh, and +also in Tasmania. The large Banana-palm-like leaves of _Taeniopteris_ +(_Macrotaeniopteris_) are common to the Triassic and Lower Jurassic beds +of India: they are also met with in New Zealand, and in the upper beds at +Bald Hill, Bacchus Marsh. + +[Illustration: =Fig. 60.--MESOZOIC PLANTS.= + + A--Thinnfeldia odontopteroides. Morris sp. Trias. N.S. Wales. + + B--Cladophlebis denticulata, Brongn. sp. var. australis, Morr. + Jurassic, Victoria. + + C--Taeniopteris spatulata, McClell. var. Daintreei, McCoy. Jurassic, + Victoria. + + D--Brachyphyllum gippslandicum, McCoy. Jurassic, Victoria. + + E--Ginkgo robusta, McCoy. Jurassic, Victoria. + +] + + +=Jurassic Plants.--= + +The Jurassic flora of Australasia is very prolific in plant forms. +These range from liverworts and horse-tails to ferns and conifers. The +commonest ferns were _Cladophlebis_, _Sphenopteris_, _Thinnfeldia_ +and _Taeniopteris_. The conifers are represented by _Araucarites_ +(cone-scales, leaves and fruits), _Palissya_ and _Brachyphyllum_ (Fig. +60). The _Ginkgo_ or Maiden-hair tree, which is still living in China +and Japan, and also as a cultivated plant, was extremely abundant in +Jurassic times, accompanied by the related genus, _Baiera_, having more +deeply incised leaves; both genera occur in the Jurassic of S. Gippsland, +Victoria, and in Queensland. The Jurassic flora of Australasia is in many +respects like that of the Yorkshire coast near Scarborough. In New Zealand +this flora is represented in the Mataura series, in which there are many +forms identical with those of the Australian Jurassic, and even of India. + + +=Cretaceous Plants.--= + +An upper Cretaceous fern, (?) _Didymosorus gleichenioides_, is found in +the sandstones of the Croydon Gold-field, North Queensland. + + +=Plants of the Cainozoic.--Balcombian Stage.--= + +The older part of the Cainozoic series in Australasia may be referred +to the Oligocene. These are marine beds with occasional, thick seams of +lignite, and sometimes of pipe-clay with leaves, the evidence of river +influence in the immediate neighbourhood. The fossil wood in the lignite +beds appears to be a _Cupressinoxylon_ or Cypress wood. Leaves referable +to plants living at the present day are also found in certain clays, +as at Mornington, containing _Eucalyptus precoriacea_ and a species of +_Podocarpus_. + +[Illustration: =Fig. 61.--CAINOZOIC PLANTS.= + + A--Cinnamomum polymorphoides, McCoy. Cainozoic. Victoria. + + B--Laurus werribeensis, McCoy. Cainozoic. Victoria. + + C--Banksia Campbelli, Ettingsh. Cainozoic. Vegetable Creek, N.S.W. + + D--Fagus Risdoniana, Ettingsh. Cainozoic. Tasmania. + + E--Spondylostrobus Smythi, Mueller. Cainozoic. (Deep Leads), Victoria. + +] + + +=Miocene Leaf-beds.--Janjukian Stage.--= + +Later Cainozoic deposits, evidently accumulated in lakes, and sometimes +ferruginous, may be referred to the Miocene. They are comparable in +age with the Janjukian marine beds of Spring Creek and Waurn Ponds in +Victoria. These occur far inland and occupy distinct basins, as at the +Wannon, Bacchus Marsh (Maddingley), and Pitfield Plains. Leaf-beds of +this age occur also on the Otway coast, Victoria, containing the genera +_Coprosmaephyllum_, _Persoonia_ and _Phyllocladus_. In all probability the +Dalton and Gunning leaf-beds of New South Wales belong here. Examples of +the genera found in beds of this age are _Eucalyptus_ (a species near _E. +amygdalina_), _Banksia_ or Native Honeysuckle, _Cinnamomum_ or Cinnamon, +_Laurus_ or Laurel, and _Fagus_ (_Notofagus_) or Beech (Fig. 61). In the +leaf-beds covered by the older basalt on the Dargo High Plains, Gippsland, +leaves of the _Ginkgo Murrayana_ occur. + +In South Australia several occurrences of leaf beds have been recorded, +containing similar species to those found in the Cainozoic of Dalton and +Vegetable Creek, New South Wales. For example, _Magnolia Brownii_ occurs +at Lake Frome, _Bombax Sturtii_ and _Eucalyptus Mitchelli_ at Elizabeth +River, and _Apocynophyllum Mackinlayi_ at Arcoona. + + +=Fruits of the "Deep Leads."--= + +The Deep Leads of Victoria, New South Wales and Tasmania probably begin +to date from the period just named, for they seem to be contemporaneous +with the "Older Gold Drift" of Victoria; a deposit sometimes containing +a marine fauna of Janjukian age. This upland river system persisted +into Lower Pliocene times, and their buried silts yield many fruits, of +types not now found in Australia, such as _Platycoila_, _Penteune_ and +_Pleioclinis_, along with _Cupressus_ (_Spondylostrobus_) and _Eucalyptus_ +of the existing flora (Fig. 62). + + +=Pleistocene Plants.--= + +The Pleistocene volcanic tuffs of Mount Gambier have been shown to contain +fronds of the living _Pteris_ (_Pteridium_) _aquilina_ or Bracken fern, +and a _Banksia_ in every way comparable with _B. marginata_, a species of +the Native Honeysuckle still living in the same district. + +[Illustration: =Fig. 62.--Leaves of a Fossil Eucalyptus. (E. pluti, +McCoy).= + +About 3/4 nat. size. From the Cainozoic Deep Leads, Daylesford, Victoria. + + (_Nat. Mus. Coll._) +] + +The siliceous valves of freshwater diatoms constitute the infusorial +earths of Victoria, Queensland, New South Wales and New Zealand. The +commonest genera met with are _Melosira_, _Navicula_, _Cymbella_ (or +_Cocconema_), _Synedra_, _Tabellaria_, _Stauroneis_ and _Gomphonema_. +They are, generally speaking, of Pleistocene age, as they are often found +filling hollows in the newer basalt flows. In Victoria diatomaceous +earths are found at Talbot (See Fig. 42), Sebastopol and Lancefield; in +Queensland, at Pine Creek; in New South Wales, at Cooma, Barraba, and the +Richmond River; and in New Zealand at Pakaraka, Bay of Islands. In the +latter country there is also a marine diatomaceous rock in the Oamaru +Series, of Miocene age. + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + _Girvanella problematica_, Nicholson and Etheridge. Cambrian: S. + Australia. + + _Bythotrephis tenuis_, J. Hall. Silurian: Victoria. + + _Haliserites Dechenianus_, Göppert sp. Silurian and Devonian: Victoria. + + _Cordaites australis_, McCoy. Upper Devonian: Victoria. + + _Lepidodendron australe_, McCoy. Lower Carboniferous: Victoria and + Queensland. Up. Devonian: New South Wales. + + _Rhacopteris inaequilatera_, Göppert sp. Carboniferous: New South + Wales. + + _Glossopteris Browniana_, Brongniart. Carbopermian: New South Wales, + Queensland, Tasmania and W. Australia. + + _Gangamopteris spatulata_, McCoy. Carbopermian: Victoria, New South + Wales and Tasmania. + + _Thinnfeldia odontopteroides_, Morris sp. Triassic: New South Wales. + Jurassic: Victoria, Queensland and Tasmania. + + _Cladophlebis denticulata._, Brongn. sp., var. australis, Morris. + Jurassic: Queensland, New South Wales, Victoria, Tasmania and New + Zealand. + + _Taeniopteris spatulata_, McClelland. Jurassic: Queensland, New South + Wales, Victoria, and Tasmania. + + (?) _Didymosorus gleichenioides_, Etheridge fil. Upper Cretaceous: + Queensland. + + _Eucalyptus precoriacea_, Deane. Oligocene: Victoria. + + _Eucalyptus_, _Banksia_, _Cinnamomum_, _Laurus_ and _Fagus_. Miocene: + Victoria, New South Wales and Tasmania. + + _Spondylostrobus Smythi_, von Mueller. (Fruits and wood). Lower + Pliocene: Victoria and Tasmania. + + _Pteris_ (_Pteridium_) _aquilina_, Linné, and _Banksia_ cf. + _marginata_, Cavanilles. Pleistocene: Victoria and South + Australia. + + + * * * * * + + +LITERATURE. + + + Girvanella.--Etheridge, R. jnr. Trans. R. Soc. S. Australia, vol. + XIII. 1890, pp. 19, 20. Etheridge, R. and Card, G. Geol. Surv. + Queensland, Bull. No. 12, 1900, pp. 26, 27, 32. Chapman, F. Rep. + Austr. Assoc. Adv. Sci., Adelaide Meeting (1907), 1908, p. 337. + + Devonian Ferns and Cordaites.--McCoy, F. Prod. Pal. Vict. Dec. V., + 1876, p. 21. Dun, W. S. Rec. Geol. Surv. New South Wales, vol. V. + pt. 3, 1897, p. 117. + + Lepidodendron.--McCoy, F. Prod. Pal. Vict., Dec. I. 1874, p. 37. + Etheridge, R. jnr. Rec. Geol. Surv, New South Wales, vol. II., + pt. 3, 1891, p. 119. Idem, Geol. and Pal. Queensland, 1892, p. + 196. + + Carboniferous Fungi.--Etheridge, R. jnr. Geol. Surv. W.A., Bull, No. + 10, 1903, pp. 25-31. + + Carboniferous Ferns.--Dun, W. S. Rec. Geol. Surv. New South Wales, + vol. VIII. pt. 2, 1905, pp. 157-161, pls. XXII. and XXIII. + + Glossopteris.--Feistmantel, O. Mem. Geol. Surv. New South Wales, Pal. + No. 3, 1890. Arber, N. Cat. Foss. Plants, Glossopteris Flora, + Brit. Mus., 1905. + + Gangamopteris.--McCoy, F. Prod. Pal. Vict., Dec. II. 1875, p. 11. + + Jurassic Plants.--McCoy, F. Prod. Pal. Vic., Dec. II. 1875, p. 15. + Woods, T. Proc. Linn. Soc. New South Wales, vol. VIII. pt. I. + 1883, p. 37. Etheridge, R. jnr. Geol. Pal. Queensland, 1892, p. + 314. Dun, W. S. (Taeniopteris), Rep. Austr. Asso. Adv. Sci., + Sydney, 1898, pp. 384-400. Seward, A. C. Rec. Geol. Surv. Vic., + vol. I. pt. 3, 1904; Chapman, F. Ibid., vol II. pt. 4, 1908; vol. + III., pt. 1, 1909. Dun, W. S. Rec. Geol. Surv. New South Wales, + vol. VIII. pt. 4, 1909, p. 311. + + Older Cainozoic Plants.--McCoy, F. Prod. Pal. Vic., Dec. IV. 1876, p. + 31. Ettingshausen, C. von. Mem. Geol. Surv. New South Wales, Pal. + No. 2, 1888. Idem, Trans. New Zealand Inst., vol. XXIII. (1890), + 1891, p. 237. Deane, H. Rec. Geol. Surv. Vict., vol. I. pt. 1, + 1902, pp. 15, 20. + + Lower Pliocene Deep Leads.--McCoy, F. Prod. Pal. Vict., Dec. IV. 1876, + p. 29. Mueller, F. von. Geol. Surv. Vic., New Veg. Foss., 1874 + and 1883. + + Pleistocene and other Diatom Earths.--Card, G. W. and Dun, W. S., Rec. + Geol. Surv. New South Wales, vol. V. pt. 3, 1897, p. 128. + + + + +CHAPTER VI. + +FOSSIL FORAMINIFERA AND RADIOLARIA. + + +=Protozoans, Their Structure.--= + +The animals forming the sub-kingdom PROTOZOA ("lowliest animals"), are +unicellular (one-celled), as distinguished from all the succeeding higher +groups, which are known as the METAZOA ("animals beyond"). The former +group, Protozoa, have all their functions performed by means of a simple +cell, any additions to the cell-unit merely forming a repetitional +or aggregated cell-structure. A familiar example of such occurs in +pond-life, in the Amoeba, a form which is not found fossilised on account +of the absence of any hard parts or covering capable of preservation. +Foraminifera and Radiolaria, however, have such hard parts, and are +frequently found fossilised. + + +=Foraminifera: Their Habitats.--= + +The _FORAMINIFERA_ are a group which, although essentially one-celled, +have the protoplasmic body often numerously segmented. The shell or test +formed upon, and enclosing the jelly-like sarcode, may consist either +of carbonate of lime, cemented sand-grains, or a sub-calcareous or +chitinous (horny) covering. The Foraminifera, with very few exceptions, +as _Mikrogromia_, _Lieberkuehnia_, and some forms of _Gromia_, are all +marine in habit. Some genera, however, as _Miliolina_, _Rotalia_ and +_Nonionina_, affect brackish water conditions. + +Since Foraminifera are of so lowly a grade in the animal kingdom, we may +naturally expect to find their remains in the oldest known rocks that show +any evidence of life. They are, indeed, first seen in rocks of Cambrian +age, although they have not yet been detected there in Australian strata. + + +=Cambrian Foraminifera.--= + +In parts of Siberia and in the Baltic Provinces, both Cambrian and +Ordovician rocks contain numerous glauconite casts of Foraminifera, +generally of the _Globigerina_ type of shell. In England some Middle +Cambrian rocks of Shropshire are filled with tiny exquisitely preserved +spiral shells belonging to the genus _Spirillina_, in which all the +characters of the test are seen as clearly as in the specimens picked out +of shore-sand at the present day. + + +=Silurian Foraminifera.--= + +The Silurian rocks in all countries are very poor in foraminiferal shells, +only occasional examples being found. In rocks of this age at Lilydale, +Victoria, the genus _Ammodiscus_, with fine sandy, coiled tests, is found +in the Cave Hill Limestone. + +So far as known, hardly any forms of this group occur in Devonian strata, +although some ill-defined shells have been found in the Eifel, Germany. + + +=Carboniferous Foraminifera.--= + +The Carboniferous rocks in many parts of the world yield an abundant +foraminiferal fauna. Such, for instance, are the _Saccammina_ and +_Endothyra_ Limestones of the North of England and the North of Ireland. +The Australian rocks of this age have not afforded any examples of the +group, since they are mainly of estuarine or freshwater origin. + +[Illustration: =Fig. 63.--PALAEOZOIC and MESOZOIC FORAMINIFERA.= + + A--Nubecularia stephensi, Howchin. Carbopermian. N.S.W. + + B--Frondicularia woodwardi, Howchin. Carbopermian. N.S.W. + + C--Geinitzina triangularis, Chapman and Howchin. Carbopermian. N.S.W. + + D--Valvulina plicata, Brady. Carbopermian. West Australia. + + E--Vaginulina intumescens, Reuss. Jurassic. West Australia. + + F--Flabellina dilatata, Wisniowski. Jurassic. West Australia. + + G--Marginulina solida, Terquem. Jurassic. West Australia. + + H--Frondicularia gaultina, Reuss. Cretaceous. West Australia. + +] + + +=Carbopermian Foraminifera.--= + +In Australia, as at Pokolbin, New South Wales, in the Mersey River +district, Tasmania, and in the Irwin River district, Western Australia, +the Permian rocks, or "Permo-carboniferous" as they are generally called, +often contain beds of impure limestone crowded with the chalky white +tests of _Nubecularia_: other interesting genera occur at the first named +locality as _Pelosina_, _Hyperammina_, _Haplophragmium_, _Placopsilina_, +_Lituola_, _Thurammina_, _Ammodiscus_, _Stacheia_, _Monogenerina_, +_Valvulina_, _Bulimina_, (?)_Pleurostomella_, _Lagena_, _Nodosaria_, +_Frondicularia_, _Geinitzina_, _Lunucammina_, _Marginulina_, _Vaginulina_, +_Anomalina_ and _Truncatulina_. The sandy matrix of certain _Glossopteris_ +leaf-beds in the Collie Coal measures in W. Australia have yielded +some dwarfed examples belonging to the genera _Bulimina_, _Endothyra_, +_Valvulina_, _Truncatulina_ and _Pulvinulina_; whilst in the Irwin River +district similar beds contain _Nodosaria_ and _Frondicularia_ (Fig. 63). + + +=Triassic Foraminifera.--= + +The Triassic and Rhaetic clays of Europe occasionally show traces of +foraminiferal shells, probably of estuarine habitat, as do the Wianamatta +beds of New South Wales, which also belong to the Triassic epoch. The +Australian representatives are placed in the genera _Nubecularia_, +_Haplophragmium_, _Endothyra_, _Discorbina_, _Truncatulina_, and +_Pulvinulina_. These shells are diminutive even for foraminifera, and +their starved condition indicates uncongenial environment. + + +=Jurassic Foraminifera.--= + +The Jurassic limestones of Western Australia, at Geraldton, contain many +species of Foraminifera, principally belonging to the spirally coiled and +slipper-shaped _Cristellariae_. Other genera present are _Haplophragmium_, +_Textularia_, _Bulimina_, _Flabellina_, _Marginulina_, _Vaginulina_, +_Polymorphina_, _Discorbina_, and _Truncatulina_. + + +=Cretaceous Foraminifera.--= + +In the Lower Cretaceous rocks known as the Rolling Downs Formation in +Queensland, shells of the Foraminifera are found in some abundance +at Wollumbilla. They are represented chiefly by _Cristellaria_ and +_Polymorphina_. + +[Illustration: =Fig. 64.--Structure in Lepidocyclina.= + + A--Vertical section through test of Lepidocyclina marginata, + Michelotti sp.: showing the equatorial chambers (eq. c) and the + lateral chambers (l.c.) + + B--Section through the median disc, showing the hexagonal and ogive + chambers. × 18. + + Cainozoic (Janjukian). Batesford, near Geelong, Victoria. + + (_F.C. Coll._) +] + + +=Cainozoic Foraminifera.--= + +The Cainozoic strata in all parts of the world are very rich in +Foraminifera, and the genera, and even many species are similar to those +now found living. Certain types, however, had a restricted range, and are +therefore useful as indicators of age. Such are the Nummulites and the +_Orbitoides_ of the Eocene and the Oligocene of Europe, India and the West +Indies; and the _Lepidocyclinae_ of the Miocene of Europe, India, Japan +and Australia (Fig. 64). + +The genus _Lepidocyclina_ is typically represented in the Batesford beds +near Geelong, Victoria by _L. tournoueri_, a fossil of the Burdigalian +stage (Middle Miocene) in Europe, as well as by _L. marginata_. A +limestone with large, well-preserved tests of the same genus, and +belonging to a slightly lower horizon in the Miocene has lately been +discovered in Papua. + +[Illustration: =Fig. 65.--CAINOZOIC FORAMINIFERA.= + + A--Miliolina vulgaris, d'Orb. sp. Oligocene-Recent. Vict. and S.A. + + B--Textularia gibbosa, d'Orb. Oligocene and Miocene. Vict. & S.A. + + C--Nodosaria affinis, d'Orb. Oligocene. Victoria. + + D--Polymorphina elegantissima. P. and J. Oligocene-Recent. Vict. and + S.A. + + E--Truncatulina ungeriana, d'Orb. sp. Oligocene-Recent. Vict. & S.A. + + F--Amphistegina vulgaris, d'Orb. Oligocene-L. Pliocene. Vict. & S.A. + +] + +Some of the commoner Foraminifera found in the Cainozoic beds of Southern +Australia are--_Miliolina vulgaris_, _Textularia gibbosa_, _Nodosaria +affinis_, _Polymorphina elegantissima_, _Truncatulina ungeriana_ and +_Amphistegina lessonii_ (Fig. 65). The first-named has a chalky or +porcellanous shell; the second a sandy test; the third and fourth glassy +or hyaline shells with excessively fine tubules; the fifth a glassy shell +with numerous surface punctations due to coarser tubules than usual in +the shell-walls; whilst the last-named has a smooth, lenticular shell, +also hyaline, and occurring in such abundance as often to constitute a +foraminiferal rock in itself. + + +=Pleistocene Foraminifera.--= + +The estuarine deposits of Pleistocene age in southern Australia often +contain innumerable shells of _Miliolina_, _Rotalia_ and _Polystomella_. +One thin seam of sandy clay struck by the bores in the Victorian Mallee +consists almost entirely of the shells of the shallow-water and estuarine +species, _Rotalia beccarii_. + + * * * * * + + +=Radiolaria: Their Structure.--= + +The organisms belonging to the order _RADIOLARIA_ are microscopic, and +they are all of marine habitat. The body of a radiolarian consists of a +central mass of protoplasm enclosed in a membranous capsule, and contains +the nuclei, vacuoles, granules and fat globules; whilst outside is a +jelly-like portion which throws off pseudopodia or thin radiating threads. +The skeleton of Radiolaria is either chitinous or composed of clear, +glassy silica, and is often of exquisitely ornamental and regular form. + + +=Habitat.--= + +These tiny organisms generally live in the open ocean at various depths, +and sinking to the bottom, sometimes as deep as 2,000 to 4,000 fathoms, +they form an ooze or mud. + + +=Subdivisions.--= + +Radiolaria are divided into the four legions or orders,--Acantharia, +Spumellaria, Nasselaria and Phaeodaria: only the second and third groups +are found fossil. The Spumellarians are spherical, ellipsoidal, or +disc-shaped, and the Nasselarians conical or helmet-shaped. + + +=Cambrian Radiolaria.--= + +Certain cherts or hard, siliceous rocks of the palaeozoic era are often +crowded with the remains of Radiolaria, giving the rock a spotted +appearance. (See _antea_, Fig. 38). Some of the genera thus found are +identical with those living at the present day, whilst others are peculiar +to those old sediments. In Australia, remains of their siliceous shells +have been found in cherts of Lower Cambrian age near Adelaide. These have +been provisionally referred to the genera _Carposphaera_ and _Cenellipsis_ +(Fig. 66). + + +=Ordovician Radiolaria.--= + +Radiolaria have been detected in the Lower Ordovician rocks of Victoria, +in beds associated with the Graptolite slates of this series. In New South +Wales Radiolarian remains are found in the cherts and slates of Upper +Ordovician age at Cooma and Mandurama. + + +=Silurian Radiolaria.--= + +The Silurian black cherts of the Jenolan Caves in New South Wales contain +casts of Radiolaria. + + +=Devonian Radiolaria.--= + +The Lower Devonian red jaspers of Bingera and Barraba in New South Wales +have afforded some casts of Radiolaria, resembling _Carposphaera_ and +_Cenosphaera_. + +[Illustration: =Fig. 66.--FOSSIL RADIOLARIA.= + + A--Aff. Carposphaera (after David and Howchin). Cambrian. Brighton, + S.A. + + B--Cenosphaera affinis, Hinde. Mid. Devonian. Tamworth, N.S.W. + + C--Amphibrachium truncatum, Hinde. Up. Cretaceous. Pt. Darwin. + + D--Dictyomitra triangularis, Hinde. Up. Cretaceous. Pt. Darwin. + +] + +The large number of fifty-three species have been found in the radiolarian +rocks of Middle Devonian age at Tamworth in New South Wales (Fig. 66). +These have been referred to twenty-nine genera comprising amongst +others, _Cenosphaera_, _Xiphosphaera_, _Staurolonche_, _Heliosphaera_, +_Acanthosphaera_ and _Spongodiscus_. + + +=Cretaceous Radiolaria.--= + +Although certain silicified rocks in the Jurassic in Europe have furnished +a large series of Radiolaria, the Australian marine limestones of this age +have not yielded any of their remains up to the present. They have been +found, however, in the Lower Cretaceous of Queensland, and in the (?)Upper +Cretaceous of Port Darwin, N. Australia. The Radiolaria from the latter +locality belong to the sub-orders Prunoidea, Discoidea and Cyrtoidea +(Fig. 66). The rock which contains these minute fossils is stated to be +eaten by the natives for medicinal purposes. As its composition is almost +pure silica, its efficacy in such cases must be more imaginary than real. + + +=Cainozoic Radiolaria.--= + +Cainozoic rocks of Pliocene age, composed entirely of Radiolaria, occur at +Barbados in the West Indies. No Cainozoic Radiolaria, however, have been +found either in Australia or New Zealand up to the present time. + + * * * * * + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + +FORAMINIFERA. + + _Nubecularia stephensi_, Howchin. Carbopermian: Tasmania and New South + Wales. + + _Frondicularia woodwardi_, Howchin. Carbopermian: W. Australia and New + South Wales. + + _Geinitzina triangularis_, Chapm. & Howchin. Carbopermian: New South + Wales. + + _Pulvinulina insignis_, Chapman. Trias (Wianamatta Series): New South + Wales. + + _Marginulina solida_, Terquem. Jurassic: W. Australia. + + _Flabellina dilatata_, Wisniowski. Jurassic: W. Australia. + + _Vaginulina striata_, d'Orbigny. Lower Cretaceous: Queensland. + + _Truncatulina lobatula_, W. and J. sp. Lower Cretaceous: Queensland. + + _Miliolina vulgaris_, d'Orb. sp. Cainozoic: Victoria and S. Australia. + + _Textularia gibbosa_, d'Orb. Cainozoic: Victoria and S. Australia. + + _Nodosaria affinis_, d'Orb. Cainozoic: Victoria and S. Australia. + + _Polymorphina elegantissima_, Parker and Jones. Cainozoic: Victoria, + Tasmania, and S. Australia. + + _Truncatulina ungeriana_, d'Orb. sp. Cainozoic: Victoria, King Island, + and S. Australia. + + _Amphistegina lessonii_, d'Orb. Cainozoic: Victoria and S. Australia. + + _Lepidocyclina martini_, Schlumberger. Cainozoic (Balcombian and + Janjukian): Victoria. + + _L. tournoueri_, Lemoine and Douvillé. Cainozoic (Junjukian): + Victoria. + + _Cycloclypeus pustulosus_, Chapman. Cainozoic (Janjukian): Victoria. + + _Fabularia howchini_, Schlumberger. Cainozoic (Kalimnan): Victoria. + + _Hauerina intermedia_, Howchin. Cainozoic (Kalimnan): Victoria. + + _Rotalia beccarii_, Linné sp. Pleistocene: Victoria and S. Australia. + + _Polystomella striatopunctata_, Fichtel and Moll sp. Pleistocene: + Victoria and S. Australia. + + +RADIOLARIA. + + (?) _Carposphaera_ sp. Lower Cambrian: South Australia. + + (?) _Cenellipsis_ sp. Lower Cambrian: South Australia. + + _Cenosphaera affinis_, Hinde. Devonian: New South Wales. + + _Staurolonche davidi_, Hinde. Devonian: New South Wales. + + _Amphibrachium truncatum_, Hinde. Upper Cretaceous: Northern Territory. + + _Dictyomitra triangularis_, Hinde. Upper Cretaceous: Northern + Territory. + + + * * * * * + + +LITERATURE. + + +FORAMINIFERA. + + Carbopermian.--Howchin, W. Trans. Roy. Soc. S. Austr., vol. XIX. 1895; + pp. 194-198. Chapman, F. and Howchin, W. Mem. Geol. Surv. New + South Wales, Pal. No. 14, 1905. Chapman, F. Bull. Geol. Surv. W. + Austr., No. 27, 1907, pp. 15-18. + + Trias.--Chapman, F. Rec. Geol. Surv. New South Wales, vol. VIII. pt. + 4, 1909, pp. 336-339. + + Jurassic.--Chapman, F. Proc. Roy. Soc. Vict., vol. XVI. (N.S.), pt. + II., 1904, pp. 186-199. + + Cretaceous.--Moore, C. Quart. Journ. Geol. Soc., vol. XXVI. 1870, pp. + 239 and 242. Howchin, W. Trans. Roy. Soc. S. Austr., vol. VIII. + 1886, pp. 79-93. Idem, ibid., vol. XIX., 1895, pp. 198-200. Idem, + Bull. Geol. Surv. W. Austr., No. 27, 1907, pp. 38-43. + + Cainozoic.--Howchin, W. Trans. Roy. Soc. S. Austr., vol. XII. 1889, + pp. 1-20. Idem, ibid., vol. XIV. 1891, pp. 350-356. Jensen, H. + I. Proc. Linn. Soc. New South Wales, vol. XXIX. pt. 4, 1905, pp. + 829-831. Goddard, E. J. and Jensen, H. I. ibid., vol. XXXII. pt. + 2, 1907, pp. 308-318. Chapman, F. Journ. Linn. Soc. Lond. Zool., + vol. XXX. 1907, pp. 10-35. + + General.--Howchin, W. Rep. Austr. Assoc. Adv. Sci., Adelaide Meeting, + 1893, pp. 348-373. + + +RADIOLARIA. + + Lower Cambrian.--David, T. W. E. and Howchin, W. Proc. Linn. Soc. New + South Wales, vol. XXI. 1897, p. 571. + + Devonian.--David, T. W. E. Proc. Linn. Soc. New South Wales, vol. XXI. + 1897, pp. 553-570. Hinde, G. J. Quart. Journ. Geol. Soc., vol. + LV. 1890, pp. 38-64. + + Upper Cretaceous.--Hinde, G. J. Quart. Journ. Geol. Soc., vol. XLIX. + 1893, pp. 221-226. + + + + +CHAPTER VII. + +FOSSIL SPONGES, CORALS AND GRAPTOLITES. + + +_SPONGES._ + + +=Characteristics of Sponges.--= + +The Sponges are sometimes placed by themselves as a separate phylum, the +Porifera. With the exception of a few freshwater genera, they are of +marine habit and to be found at all depths between low tide (littoral) +and deep water (abyssal). Sponges are either fixed or lie loosely on the +sea-floor. They possess no organs of locomotion, and have no distinct axis +or lateral appendages. They exist by setting up currents in the water +whereby the latter is circulated through the system, carrying with it +numerous food particles, their tissues being at the same time oxygenated. +Their framework, in the siliceous and calcareous sponges, is strengthened +by a mineral skeleton, wholly or partially capable of preservation as a +fossil. + + +=Cambrian and Ordovician Sponges.--= + +The oldest rocks in Australia containing the remains of Sponges are the +Cambrian limestones of South Australia, at Ardrossan and elsewhere. +Some of these sponge-remains are referred to the genus _Protospongia_, +a member of the Hexactinellid group having 6-rayed skeletal elements. +When complete, the _Protospongia_ has a cup- or funnel-shaped body, +composed of large and small modified spicules, which form quadrate areas, +often seen in isolated or aggregated patches on the weathered surface of +the rock. _Protospongia_ also occurs in the Lower Ordovician slates and +shales of Lancefield (_P. oblonga_), and Bendigo (_P. reticulata_ and _P. +cruciformis_), in Victoria (Fig. 67 A). At St. David's, in South Wales, +the genus is found in rocks of Middle Cambrian age. The South Australian +limestones in which _Protospongia_ occurs are usually placed in the Lower +Cambrian. + +[Illustration: =Fig. 67.--PALAEOZOIC SPONGES, &c.= + + A--Protospongia reticulata, T. S. Hall. Low. Ordovician. Bendigo. + + B--Receptaculites fergusoni, Chapm. Silurian. Wombat Creek, Vict. + + C--R. australis, Salter. (Section of wall, etched, after Eth. & Dun) + Mid. Devonian. Co. Murray, N.S.W. + + D--Protopharetra scoulari, Eth. fil. Cambrian. S.A. + +] + +Another genus of Sponges, _Hyalostelia_, whose affinities are not very +clear, occurs in the South Australian Cambrian at Curramulka. This type +is represented by the long, slightly bent, rod-like spicules of the +root-tuft, and the skeletal spicules with six rays, one of which is much +elongated. + +_Stephanella maccoyi_ is a Monactinellid sponge, found in the Lower +Ordovician (Bendigo Series) of Bendigo, Victoria. + + +=Silurian Sponges.--= + +Numerous Sponges of Silurian age are found in the neighbourhood of Yass, +New South Wales, which belong to the Lithistid group, having irregular, +knotty and branching spicules. These sponges resemble certain fossil +fruits, generally like diminutive melons; their peculiar spicular +structure, however, is usually visible on the outside of the fossil, +especially in weathered specimens. The commonest genus is _Carpospongia_. + + +=Receptaculites: Silurian to Carboniferous.--= + +In Upper Silurian, Devonian, and Carboniferous times the curious saucer- +or funnel-shaped bodies known as _Receptaculites_ must have been fairly +abundant in Australia, judging by their frequent occurrence as fossils. +They are found as impressions or moulds and casts in some of the mudstones +and limestones of Silurian age in Victoria, as at Loyola and Wombat +Creek, in west and north-east Gippsland respectively. In the Devonian +limestones of New South Wales they occur at Fernbrook, near Mudgee, at the +Goodradigbee River, and at Cavan, near Yass; also in beds of the same age +in Victoria, at Bindi, and Buchan (Fig. 67, B.C.). _Receptaculites_ also +occur in the Star Beds of Upper Devonian or Lower Carboniferous age in +Queensland, at Mount Wyatt. It will thus be seen that this genus has an +extensive geological range. + + +=Carbopermian Sponges.--= + +A Monactinellid Sponge, provisionally referred to _Lasiocladia_, has been +described from the Gympie beds of the Rockhampton District, Queensland. +_Lasiocladia_, as well as the Hexactinellid Sponge _Hyalostelia_, occurs +in the Carbopermian of New South Wales. + + +=Cretaceous Sponges.--= + +No sponge-remains seem to occur above the Carbopermian in Australia +until we reach the Cretaceous rocks. In the Lower Cretaceous series in +Queensland a doubtful member of the Hexactinellid group is found, namely, +_Purisiphonia clarkei_. In the Upper Cretaceous of the Darling Downs +District pyritized Sponges occur which have been referred to the genus +_Siphonia_, a member of the Lithistid group, well known in the Cretaceous +of Europe. + + +=Cainozoic Sponges.--= + +A white siliceous clay, supposed to be from a "Deep Lead," in the +Norseman district in Western Australia, has proved to consist almost +entirely of siliceous sponge-spicules, belonging to the Monactinellid, +the Tetractinellid, the Lithistid, and the Hexactinellid groups (Fig. 69 +A, B). The reference of the deposit to a "deep lead" or alluvial deposit +presents a difficulty, since these sponge-spicules represent moderately +deep water marine forms. This deposit resembles in some respects the +spicule-bearing rock of Oamaru, New Zealand, which is of Miocene age. + +[Illustration: =Fig. 68.--CAINOZOIC SPONGES.= + + A--Latrunculia sp. (after Hinde). Cainozoic. Deep Lead, Norseman, W.A. + + B--Geodia sp. (after Hinde). Cainozoic. Deep Lead, Norseman, W.A. + + C--Ecionema newberyi. McCoy sp. Cainozoic. Boggy Creek, Gippsland, + Vict. + + D--Plectroninia halli, Hinde. Cainozoic (Janjukian). Moorabool, Vict. + + E--Tretocalia pezica, Hinde. Cainozoic. Flinders, Vict. + +] + +[Illustration: =Fig. 69.--SILURIAN CORALS.= + + A--Cyathophyllum approximans, Chapm. Silurian (Yer.). Gippsland, Vict. + + B--Favosites grandipora, Eth. fil. Silurian (Yer.). Lilydale, Vict. + + C--Favosites grandipora, vertical section. Ditto. + + D--F. grandipora, transverse section. Ditto. + + E--Pleurodictyum megastomum, Dun. Lilydale, Vict. + + F--Halysites peristephesicus, Eth. fil. Silurian. N.S. Wales. + + G--Heliolites interstincta, Wahl sp Vict. (transv. sect). Silurian.. + +] + +In the Cainozoic beds of southern Australia Sponges with calcareous +skeletons are not at all uncommon. The majority of these belong to the +Lithonine section of the Calcispongiae, in which the spicules are regular, +and not fixed together. Living examples of these sponges, closely related +to the fossils, have been dredged from the Japanese Sea. The fossils +are found mainly in the Janjukian, at Curlewis, in the Moorabool River +limestones, and in the polyzoal rock of Flinders, all in Victoria. They +belong to the genera _Bactronella_, _Plectroninia_ and _Tretocalia_ (Fig. +68, D and E). Some diminutive forms also occur in the older series, the +Balcombian, at Mornington, namely, _Bactronella parvula_. At Boggy Creek, +near Sale, in Victoria, a Tetractinellid Sponge, _Ecionema newberyi_, is +found in the Janjukian marls; spicules of this form have also been noted +from the clays of the Altona Bay coal-shaft (Fig. 68 C). + + * * * * * + +The _ARCHAEOCYATHINAE_: an ancient class of organisms related both to the +Sponges and the Corals. + + +=Archaeocyathinae in Cambrian Strata.--= + +These curious remains have been lately made the subject of detailed +research, and it is now concluded that they form a group probably +ancestral both to the sponges and the corals. They are calcareous, and +generally cup-shaped or conical, often furnished at the pointed base +with roots or strands for attachment to the surrounding reef. They have +two walls, both the inner and the outer being perforated like sponges. +As in the corals, they are divided by transverse septa and these are +also perforated. Certain of the genera as _Protopharetra_ (Fig. 67 D), +_Coscinocyathus_, and _Archaeocyathina_, are common to the Cambrian +of Sardinia and South Australia, whilst other genera of the class are +also found in Siberia, China, Canada and the United States. A species +of _Protopharetra_ was recently detected in a pebble derived from +the Cambrian limestone in the Antarctic, as far south as 85 deg. An +_Archaeocyathina_ limestone has also been found in situ from Shackleton's +farthest south. + + +_CORALS_ (Class Anthozoa). + + +=Rugose Corals.--= + +Many of the older types of Corals from the Palaeozoic rocks belong to the +Tetracoralla (septa in multiples of four), or Rugosa (i.e., with wrinkled +exterior). + + +=Ordovician Corals.--= + +In Great Britain and North America Rugose Corals are found as early as +Ordovician times, represented by _Streptelasma_, _Petraia_, etc. In +Australia they seem to first make their appearance in the Silurian period. + + +=Silurian Corals.--= + +In rocks of Silurian age in Australia we find genera like _Cyathophyllum_ +(with single cups or compound coralla), _Diphyphyllum_, _Tryplasma_ +and _Rhizophyllum_, the first-named often being very abundant. The +compound corallum of _Cyathophyllum approximans_ presents a very handsome +appearance when cut transversely and polished. This coral is found in +the Newer Silurian limestone in Victoria; it shows an alliance with _C. +mitchelli_ of the Middle Devonian of the Murrumbidgee River, New South +Wales (Fig. 69 A). + + +=Silurian Hexacoralla.--= + +It is, however, to the next group, the Hexacoralla, with septa in +multiples of six, twelve, and twenty-four, that we turn for the most +varied and abundant types of Corals in Silurian times. The genus +_Favosites_ (Honey-comb Coral) is extremely abundant in Australian +limestones (Fig. 69 B, C), such as those of Lilydale, Walhalla, and +Waratah Bay in Victoria, and of Hatton's Corner and other localities near +Yass, in New South Wales. _Pleurodictyum_ is also a familiar type in the +Australian Silurian, being one of the commonest corals in the Yeringian +stage; although, strange to say, in Germany and N. America, it is typical +of Devonian strata (Fig. 69 E). _Pleurodictyum_ had a curious habit of +growing, barnacle fashion, on the side of the column of the crinoids +or sea-lilies which flourished in those times. _Syringopora_, with its +funnel-shaped tabulae or floor partitions, is typical of many Australian +limestones, as those from Lilydale, Victoria, and the Delegate River, New +South Wales. _Halysites_ (Chain Coral), with its neat strings of tubular +and tabulated corallites joined together by their edges, is another +striking Coral of the Silurian period (Fig. 69 F). This and the earlier +mentioned _Syringopora_, is by some authors regarded as belonging to the +Alcyonarian Corals (typically with eight tentacles). _Halysites_ is known +from the limestones of the Mitta Mitta River, N.E. Gippsland, Victoria; +from the Molong and Canobolas districts in New South Wales; from the +Gordon River limestone in Tasmania; and from Chillagoe in Queensland. +Abroad it is a well known type of Coral in the Wenlockian of Gotland +in Scandinavia, and Shropshire in England, as well as in the Niagara +Limestone of the United States. + + +=Silurian Octocoralla.--= + +Perhaps the most important of the Octocoralla is _Heliolites_ +("Sunstone"), which is closely allied to the Blue Coral, _Heliopora_, a +frequent constituent of our modern coral reefs. The genus _Heliolites_ +has a massive, calcareous corallum, bearing two kinds of pores or tubes, +large (autopores) containing complete polyps, and small (siphonopores) +containing the coenosarc or flesh of the colony. Both kinds of tubes are +closely divided by tabulae, whilst the former are septate. _Heliolites_ is +of frequent occurrence in the Silurian limestones of New South Wales and +Victoria (Fig. 69 G). + + +=Devonian Corals.--= + +The Middle Devonian beds of Australia are chiefly limestones, such as +the Buchan limestone, Victoria; the Burdekin Series, Queensland; and +the Tamworth limestone of New South Wales. These rocks, as a rule, are +very fossiliferous, and the chief constituent fossils are the Rugose and +Perforate Corals. _Campophyllum gregorii_ is a common form in the Buchan +limestone (Fig. 70 A), as well as some large mushroom-shaped _Favosites_, +as _F. gothlandica_ and _F. multitabulata_. Other genera which may +be mentioned as common to the Australian Middle Devonian rocks are, +_Cyathophyllum_, _Sanidophyllum_ and _Spongophyllum_, _Heliolites_ is +also found in limestones of this age in New South Wales and Queensland. + +[Illustration: =Fig. 70.--UPPER PALAEOZIC CORALS.= + + A--Campophyllum gregorii, Eth. fil. Mid. Devonian. Buchan, Vict. + + B--Pachypora meridionalis, Nich. & Eth. fil. Mid Devonian. Queens. + + C--Aulopora repens, Kn. & W. (after Hinde). Devonian. Kimberley + district, W.A. + + D--Zaphrentis culleni, Eth. fil. Carboniferous. New South Wales. + + E--Trachypora wilkinsoni, Eth. fil. Carbopermian (Up. Marine Ser.) New + South Wales. + + F--Stenopora crinita, Lonsdale. Carbopermian (Up. Mar. Ser.) N.S.W. + +] + +In the Burdekin Series (Middle Devonian) in Queensland we also find +_Cystiphyllum_, _Favosites gothlandica_, and _Pachypora meridionalis_ +(Fig. 70 B), whilst in beds of the same age at Rough Range in Western +Australia are found _Aulopora repens_ (Fig. 70 C), and another species of +_Pachypora_, namely, _P. tumida_. + + +=Carbopermian Corals.--= + +The only true Carboniferous marine fauna occurring in Australia, appears +to be that of the Star Beds in Queensland, but so far no corals have been +found. The so-called Carboniferous of Western Australia may be regarded +as Carbopermian or even of Permian age. The marine Carbopermian beds of +New South Wales contain several genera of Corals belonging to the group +Rugosa, as _Zaphrentis_ (Fig. 70 D), _Lophophyllum_, and _Campophyllum_. +Of the Tabulate corals may be mentioned _Trachypora wilkinsoni_, very +typical of the Upper Marine Series (Fig. 70 E) and _Cladochonus_. + +In the Gympie beds of the same system in Queensland occur the following +rugose corals, _Zaphrentis profunda_ and a species of _Cyathophyllum_. + +In the Carbopermian of Western Australia the rugose corals are represented +by _Amplexus_, _Cyathophyllum_, and _Plerophyllum_, which occur in rocks +on the Gascoyne River. + +The imperfectly understood group of the Monticuliporoids, by some authors +placed with the Polyzoa (Order Trepostomata), are well represented in +Australia by the genus _Stenopora_ (Fig. 70 F). The corallum is a massive +colony of long tubes set side by side and turned outwards, the polyp +moving upwards in growth and cutting off the lower part of the tube by +platforms like those in the tabulate corals. Some of the species of +_Stenopora_, like _S. tasmaniensis_, of New South Wales and Tasmania, +are found alike in the Lower and Upper Marine Series. _S. australis_ is +confined to the Bowen River Coal-field of Queensland. _Stenopora_ often +attains a large size, the corallum reaching over a foot in length. + +Neither Jurassic or Cretaceous Corals have been found in Australasia, +although elsewhere as in Europe and India, the representatives of modern +corals are found in some abundance. + + +=Cainozoic Corals.--= + +In Tertiary times the marine areas of southern Australia were the home +of many typical solitary Corals of the group of the Hexacoralla. In the +Balcombian beds of Mornington, Victoria, for instance, we have genera +such as _Flabellum_, _Placotrochus_, _Sphenotrochus_, _Ceratotrochus_, +_Conosmilia_, _Trematotrochus_, _Notophyllia_ and _Balanophyllia_ (Fig. +71). + +[Illustration: =Fig. 71.--CAINOZOIC CORALS.= + + A--Flabellum victoriae, Duncan. Balcombian. Mornington, Vict. + + B--Placotrochus deltoideus, Dunc. Balcombian. Muddy Creek, Hamilton, + Vic. + + C--Balanophyllia seminuda, Dunc. Balcombian. Muddy Creek, Hamilton, + Vic. + + D--Stephanotrochus tatei, Dennant. Janjukian. Torquay, near Geelong, + Vict. + + E--Thamnastraea sera, Duncan. Janjukian. Table Cape, Tas. + + F--Graphularia senescens. Tate sp. Janjukian. Waurn Ponds, near + Geelong, Vic. + + G--Trematotrochus clarkii, Dennant. Kalimnan. Gippsland Lakes, Vic. + +] + +Corals especially characteristic of the Janjukian Series are _Paracyathus +tasmanicus_, _Stephanotrochus tatei_, _Montlivaltia variformis_, +_Thamnastraea sera_ and _Dendrophyllia epithecata_. The stony axis of the +Sea-pen, _Graphularia senescens_, a member of the Octocoralla, is also +typical of this stage, and are called "square-bones" by the quarrymen at +Waurn Ponds, near Geelong, where these fossils occur. + +The Kalimnan Corals are not so abundantly represented as in the foregoing +stages, but certain species of _Flabellum_ and _Trematotrochus_, as _F. +curtum_ and _T. clarkii_, are peculiar to those beds. Several of the +Janjukian Corals persist into Kalimnan times, some dating as far back as +the Balcombian, as _Sphenotrochus emarciatus_. The Sea-pen, _Graphularia +senescens_ is again found at this higher horizon, at Beaumaris; it +probably represents a varietal form, the axis being smaller and more +slender. + +Other examples of the Octocoralla are seen in _Mopsea_, two species of +which are found in the Janjukian at Cape Otway; the deeper beds of the +Mallee; and the Mount Gambier Series. + +A species of the Astraeidae (Star-corals) of the reef-forming section, +_Plesiastraea st.vincenti_, is found in the Kalimnan of Hallett's Cove, +South Australia. + + +_HYDROZOA._ + +The few animals of this group met with in fossil faunas are represented by +the living _Millepora_ (abundant as a coral reef organism), _Hydractinia_ +(parasitic on shells, etc.), and _Sertularia_ (Sea-firs). + + +=Milleporids and Stylasterids.--= + +Although so abundant at the present time, the genus _Millepora_ does +not date back beyond the Pleistocene. The Eocene genus _Axopora_ is +supposed to belong here, but is not Australian. Of the Stylasterids one +example is seen in _Deontopora_, represented by the branchlets of _D. +mooraboolensis_, from the Janjukian limestone of the Moorabool Valley, +near Geelong. + + +=Hydractinia.--= + +_Hydractinia_ dates from the Upper Cretaceous rocks in England, and in +Australia its encrusting polypidom is found attached to shells in the +polyzoal limestone of Mount Gambier (Miocene). + + +Stromatoporoids. + +An important group of reef-builders in Palaeozoic times was the organism +known as _Stromatopora_, and its allies. The structures of these hydroid +polyps resemble successional and repetitional stages of a form like +_Hydractinia_. As in that genus it always commenced to grow upon a base +of attachment such as a shell, increasing by successive layers, until the +organic colony often reached an enormous size, and formed great mounds +and reefs (see _antea_, Fig. 32). The stromatoporoid structure was formed +by a layer of polyp cells separated by vertical partitions, upon which +layer after layer was added until a great vertical thickness was attained. +This limestone-making group first appeared in the Silurian, and probably +reached its maximum development in Middle Devonian times, when it almost +disappeared, except to be represented in Carbopermian strata by a few +diminutive forms. + +[Illustration: =Fig. 72.--STROMATOPOROIDEA and CLADOPHORA.= + + A--Actinostroma clathratum, Nich. Devonian. Rough Range, W.A. + + B--Actinostroma clathratum, Nich. Devonian. Rough Range, W.A. Vertical + section. + + (_After G. J. Hinde_). + + C--Callograptus sp. Up. Ordovician. San Remo, Vict. + + (_After T. S. Hall_). + + D--Ptilograptus sp. Up. Ordovician. San Remo, Vict. + + (_After T. S. Hall_). + + E--Dictyonema pulchellum, T. S. Hall. L. Ordov. Lancefield, Vict. + + F--Dictyonema macgillivrayi, T. S. Hall. L. Ordov. Lancefield, Vict. + +] + + +=Silurian Stromatoporoids.--= + +In the Silurian limestones of Victoria (Lilydale, Waratah Bay, Walhalla +and Loyola), and New South Wales (near Yass), Stromatoporoids belonging to +the genera _Clathrodictyon_ (probably _C. regulare_), _Stromatopora_ and +_Idiostroma_ occur. _Stromatoporella_ has been recorded from the Silurian +rocks of the Jenolan Caves, New South Wales. + + +=Devonian Stromatoporids.--= + +The Middle Devonian strata of Bindi, Victoria, yield large, massive +examples of _Actinostroma_. This genus is distinguished from the closely +allied _Clathrodictyon_ by its vertical pillars passing through several +laminae in succession. Rocks of the same age in Queensland contain +_Stromatopora_, whilst in Western Australia the Rough Range Limestone +has been shown to contain _Actinostroma clathratum_ (Fig. 72 A, B) and +_Stromatoporella eifeliensis_. + + +Cladophora. + + +=Palaeozoic Cladophora.--= + +Some branching and dendroid forms of Hydrozoa probably related to +the modern Calyptoblastea ("covered buds"), such as _Sertularia_ and +_Campanularia_, are included in the Cladophora ("Branch bearers"). They +existed from Cambrian to Devonian times, and consist of slender, forking +branches sometimes connected by transverse processes or dissepiments, the +branches bearing on one or both sides little cups or hydrothecae which +evidently contained the polyps, and others of modified form, perhaps for +the purpose of reproduction. The outer layer, called the periderm was of +chitinous material. They were probably attached to the sea-floor like the +Sertularians (Sea-firs). + + +=Dictyonema and Allies.--= + +Remains of the above group are represented in the Australian rocks by +several species of _Dictyonema_ (Fig. 72 E, F) occurring in the Lower +Ordovician of Lancefield, and in similar or older shales near Mansfield. +Some of these species are of large size, _D. grande_ measuring nearly a +foot in width. The genera _Callograptus_, _Ptilograptus_ (Fig. 72 C, D) +and _Dendrograptus_ are also sparsely represented in the Upper Ordovician +of Victoria, the two former from San Remo, the latter from Bulla. + + +Graptolites (Graptolitoidea).-- + + +=Value of Graptolites to Stratigraphist.--= + +The Graptolites were so named by Linnaeus from their resemblances to +writing on the slates in which their compressed remains are found. They +form a very important group of Palaeozoic fossils in all parts of the +world where these rocks occur, and are well represented in Australasia. +The species of the various Graptolite genera are often restricted to +particular beds, and hence they are of great value as indicators of +certain horizons or layers in the black, grey or variously coloured slates +and shales of Lower Ordovician to Silurian times. By their aid a stratum +or set of strata can be traced across country for long distances, and the +typical species can be correlated even with those in the older slates and +shales of Great Britain and North America. + + +=Nature of Graptolites.--= + +The Graptolites were compound animals, consisting of a number of polyps +inserted in cups or thecae which budded out in a line from the primary +sicula or conical chamber, which chamber was probably attached to floating +sea-weed, either by a fine thread (nema), or a disc-like expansion. This +budding of the polyp-bearing thecae gives to the polypary or colony the +appearance of a fret-saw, with the teeth directed away from the sicula. + +The habit of the earlier graptolites was to branch repeatedly, as +in _Clonograptus_, or to show a compound leaf-like structure as in +_Phyllograptus_. Later on the many-branched forms had their branches +reduced until, as in _Didymograptus_, there were only two branches. +Sometimes the branches opened out to direct the thecae upwards, the better +to procure their food supply. In _Diplograptus_ the thecae turned upwards +and acquired a support by the formation of a medium rod (virgula), often +ending in a disc or float. In Silurian times _Monograptus_ prevailed, a +genus having only a single row of thecae supported by a straight or curved +virgula. In _Retiolites_ the polypary opened out by means of a net-work +of fine strands, rendering it better able to float, at the same time +retaining its original strength. + + +=Lower Ordovician Graptolites, Victoria.--= + +The Lower Ordovician slates and shales of Victoria have been successfully +divided into several distinct series by means of the Graptolites. These, +commencing at the oldest, are:-- + +(1) Lancefield Series. Characterised by _Bryograptus clarki_, _B. +victoriae_, _Didymograptus pritchardi_, _D. taylori_ and _Tetragraptus +decipiens_. Other forms less restricted are, _Clonograptus magnificus_ +(measuring over a yard in breadth), _C. flexilis_, _C. rigidus_, +_Leptograptus antiquus_ and _Tetragraptus approximatus_ (Fig. 73). + +(2) Bendigo Series. Characterised by _Tetragraptus fruticosus_, _T. +pendens_, _Trichograptus fergusoni_ and _Goniograptus thureaui_. This +series also contains _Tetragraptus serra_ (ranging into Darriwill +Series), _T. bryonoides_, _T. quadribrachiatus_, _T. approximatus_ (base +of the series), _Phyllograptus typus_, _Dichograptus octobrachiatus_, +_Goniograptus macer_ and many _Didymograpti_, including _D. bifidus_ (Fig. +74). + +[Illustration: =Fig. 73.--LOWER ORDOVICIAN GRAPTOLITES.= + + A--Bryograptus clarki, T. S. Hall. L. Ordovician. Lancefield, Vict. + + B--Tetragraptus fruticosus, J. Hall sp. L. Ordovician. Lancefield. + + C--Phyllograptus typus, J. Hall. L. Ordovician. Lancefield. + + D--Goniograptus macer, T. S. Hall. L. Ordovician. Lancefield. + + E--Didymograptus caduceus, Salter. L. Ordovician. Lancefield. + + F--Trigonograptus wilkinsoni, T. S. Hall. L. Ordov. Darriwill, Vict. + +] + +[Illustration: =Fig. 74.--LOWER ORDOVICIAN GRAPTOLITES.= + + A--Loganograptus logani, J. Hall sp. L. Ordov. Newham, Vict. + + B--Tetragraptus approximatus, Nich. L. Ordovician. Canada and Victoria. + + (_After Nicholson_) + + C--Tetragraptus serra, Brongn. sp. L. Ordovician. Lancefield, Vict. + + D--Didymograptus bifidus, J. Hall. L. Ordovician. Guildford, Vict. + +] + +(3) Castlemaine Series. Characterised by _Didymograptus bifidus_, _D. +caduceus_ and _Loganograptus logani_. _Phyllograptus_ persists from the +Bendigo Series. It also contains _Tetragraptus serra_, _T. bryonoides_, +_T. quadribrachiatus_, _Goniograptus macer_ and several _Didymograpti_. + +(4) Darriwill Series. Characterised by _Trigonograptus wilkinsoni_. +Also contain _Diplograptus_, _Glossograptus_ and _Lasiograptus_, whilst +_Didymograptus_ is rare. + + +=Lower Ordovician Graptolites, New Zealand.--= + +In New Zealand Lower Ordovician Graptolites are found in the Kakanui +Series, at Nelson, north-west of South Island. Some of the commoner forms +are _Didymograptus extensus_, _D. caduceus_, _Loganograptus logani_, +_Phyllograptus typus_, _Tetragraptus similis_ and _T. quadribrachiatus_. + +Graptolites agreeing closely with those of the Lancefield Series of +Victoria occur near Preservation Inlet in the extreme South-west, and have +been identified as _Clonograptus rigidus_, _Bryograptus victoriae_ and +_Tetragraptus decipiens_. + + +=Upper Ordovician Graptolites, Victoria.--= + +The Upper Ordovician rocks of Victoria, as at Wombat Creek and Mount +Wellington in Gippsland, and at Diggers' Rest near Sunbury, contain the +double branched forms like _Dicranograptus ramosus_, _Dicellograptus +elegans_ and _D. sextans_; the sigmoidal form _Stephanograptus gracilis_; +and the diprionidian (biserial) forms as _Diplograptus tardus_, +_Climacograptus bicornis_, _Cryptograptus tricornis_, _Glossograptus +hermani_ and _Lasiograptus margaritatus_ (Fig. 75). + +[Illustration: =Fig. 75.--UPPER ORDOVICIAN and SILURIAN GRAPTOLITES.= + + A--Dicranograptus ramosus, J. Hall sp. Up. Ordovician. Victoria. + + B--Dicellograptus elegans, Carruthers sp. Up. Ordovician. Victoria. + + C--Diplograptus carnei, T. S. Hall. Up. Ordovician. N. S. Wales. + + D--Climacograptus bicornis, J. Hall. Up. Ordovician. Victoria. + + E--Glossograptus hermani, T. S. Hall. Up. Ordovician. Victoria. + + F--Retiolites australis, McCoy. Silurian. Keilor, Victoria. + + G--Monograptus dubius, Suess. Silurian. Wood's Point, Victoria. + +] + + +=Upper Ordovician Graptolites, New South Wales.--= + +In New South Wales, at Tallong, the Upper Ordovician Graptolites are well +represented by such forms as _Dicellograptus elegans_, _Dicranograptus +nicholsoni_, _Diplograptus carnei_, _D. foliaceus_, _Cryptograptus +tricornis_ and _Glossograptus quadrimucronatus_, etc. Other localities in +New South Wales for this Graptolite fauna are Stockyard Creek, Currowang, +Tingaringi, Lawson, and Mandurama. + + +=Tasmania.--= + +From Tasmania a _Diplograptus_ has been recorded, but the particular +horizon and locality are uncertain. + + +=Silurian Graptolites, Victoria.--= + +In the Silurian shales at Keilor, in Victoria, _Monograptus_ is a common +genus, and _Cyrtograptus_ and _Retiolites australis_ (Fig. 75 F) also +occur. Several species of _Monograptus_ have also been found at South +Yarra and Studley Park. At the latter place and Walhalla _Monograptus +dubius_, which is a Wenlock and Ludlow fossil in Britain, has been found +in some abundance (Fig. 75 G). + + * * * * * + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + +SPONGES. + + _Protospongia_ sp. Cambrian: S. Australia. + + _Hyalostelia_ sp. Cambrian: S. Australia. + + _Protospongia oblonga_, Hall. L. Ordovician: Victoria. + + _Stephanella maccoyi_, Hall. L. Ordovician: Victoria. + + _Carpospongia_ sp. Silurian: Yass, New South Wales. + + _Receptaculites fergusoni_, Chapman. Silurian: Victoria. + + _Receptaculites australis_, Salter sp. Devonian: Victoria and New + South Wales. Carboniferous: Queensland. + + (?) _Lasiocladia hindei_, Eth. fil. Carbopermian: Queensland. + + _Purisiphonia clarkei_, Bowerbank. Lower Cretaceous: Queensland. + + _Geodia_ sp. Cainozoic: W. Australia. + + _Tethya_ sp. Cainozoic: W. Australia. + + _Ecionema newberyi_, McCoy sp. Cainozoic: Victoria. + + _Plectroninia halli_, Hinde. Cainozoic (Janjukian): Victoria. + + _Tretocalia pezica_, Hinde. Cainozoic (Janjukian): Victoria. + + +ARCHAEOCYATHINAE. + + _Protopharetra scoulari_, Etheridge, fil. Cambrian: S. Australia. + + _Coscinocyathus australis_, Taylor. Cambrian: S. Australia. + + _Archaeocyathina ajax_, Taylor. Cambrian: S. Australia. + + +CORALS. + + _Cyathophyllum approximans_, Chapman. Silurian: Victoria. + + _Tryplasma liliiformis_, Etheridge, fil. Silurian: New South Wales. + + _Favosites grandipora_, Etheridge fil. Silurian: Victoria. + + _Pleurodictyum megastomum_, Dun. Silurian: Victoria. + + _Halysites peristephicus_, Etheridge, fil. Silurian: New South Wales. + + _Heliolites interstincta_, Linné sp. Silurian: Victoria. + + _Campophyllum gregorii_, Eth. fil. Middle Devonian: Victoria and + Queensland. + + _Cystiphyllum australasicum_, Eth. fil. Middle Devonian: New South + Wales and Queensland. + + _Favosites multitabulata_, Eth. fil. Middle Devonian: Victoria and New + South Wales. + + _Pachypora meridionalis_, Eth. fil. Middle Devonian: Queensland. + + _Zaphrentis culleni_, Eth. fil. Carboniferous: New South Wales. + + _Lophophyllum corniculum_, de Koninck. Carboniferous: New South Wales. + + _Zaphrentis profunda_, Eth. fil. Carbopermian: Queensland. + + _Campophyllum columnare_, Eth. fil. Carbopermian: New South Wales. + + _Trachypora wilkinsoni_, Eth. fil. Carbopermian: New South Wales. + + _Stenopora tasmaniensis_, Lonsdale. Carbopermian: Tasmania and New + South Wales. + + _Flabellum gambierense_, Duncan. Cainozoic: Victoria, S. Australia and + Tasmania. + + _Placotrochus deltoideus_, Duncan. Cainozoic: Victoria, S. Australia + and Tasmania. + + _Sphenotrochus emarciatus_, Duncan. Cainozoic: Victoria, S. Australia, + and Tasmania. + + _Ceratotrochus exilis_, Dennant. Cainozoic: Victoria. + + _Conosmilia elegans_, Duncan. Cainozoic: Victoria. + + _Balanophyllia armata_, Duncan. Cainozoic: Victoria. + + _Thamnastraea sera_, Duncan. Cainozoic: Victoria and Tasmania. + + _Graphularia senescens_, Tate sp. Cainozoic: Victoria and S. Australia. + + +HYDROZOA. + + _Clathrodictyon_ (?) _regulare_, Rosen sp. Silurian: Victoria. + + _Actinostroma clathratum_, Nicholson. Devonian: W. Australia. + + _Stromatoporella eifeliensis_, Nich. Devonian: W. Australia. + + _Dictyonema pulchella_, T. S. Hall. Lower Ordovician: Victoria. + + _Ptilograptus_ sp. L. Ordovician: Victoria. + + _Callograptus_ sp. Lower Ordovician: Victoria. + + +GRAPTOLITES. + + _Bryograptus victoriae_, T. S. Hall. Lower Ordovician (Lancefield + Series): Victoria. + + _Tetragraptus fruticosus_, J. Hall. L. Ordovician (Bendigo Series): + Victoria. + + _Didymograptus caduceus_, Salter. L. Ordovician (Castlemaine Series): + Victoria. Also New Zealand. + + _Didymograptus bifidus_, J. Hall. L. Ordovician (Castlemaine Series): + Victoria. Also New Zealand. + + _Trigonograptus wilkinsoni_, T. S. Hall. L. Ordovician (Darriwill + Series): Victoria. + + _Dicranograptus ramosus_, J. Hall sp. Upper Ordovician: Victoria. + + _Monograptus dubius_, Suess. Silurian: Victoria. + + _Retiolites australis_, McCoy. Silurian: Victoria. + + + * * * * * + + +LITERATURE. + + +SPONGES. + + Cambrian.--Tate, R. Trans. R. Soc. S. Austr., vol. XV. (N.S.), 1892, + p. 188. + + Ordovician.--Hall, T. S. Proc. R. Soc. Vict., vol. I. pt. I. 1889, pp. + 60, 61 (_Protospongia_). Idem, ibid., vol. XI. (N.S.), pt. II. + 1899, pp. 152-155 (_Protospongia and Stephanella_). + + Silurian to Carboniferous.--Salter, J. W. Canad. Org. Rem. Dec. I. + 1859, p. 47. Etheridge, R. jnr. and Dun, W. S. Rec. Geol. Surv. + New South Wales, vol. VI. 1898, pp. 62-75. Chapman, F. Proc. R. + Soc. Vict. vol. XVIII. (N.S.), pt. 1, 1905, pp. 5-15. + + Carbopermian.--Etheridge, R. jnr., in Geol. and Pal. Q., 1892, p. 199. + + Cretaceous.--Bowerbank, J. S. Proc. Zool. Soc. Lond., 1869, p. 342. + Etheridge, R. jnr. in Geol. and Pal. Queensland, 1892, pp. 438, + 439 (_Purisiphonia_). + + Cainozoic.--McCoy, F. Prod. Pal. Vict., Dec. V. 1877. Chapman, F. + Proc. R. Soc. Vict., vol. XX. (N.S.), pt. 2, 1908, pp. 210-212 + (_Ecionema_). Hinde, G. J. Quart. Journ. Geol. Soc., vol. LVI., + 1900, pp. 50-56 (calcisponges). Idem, Bull. Geol. Surv. W. + Austr., No. 36, 1910, pp. 7-21 (sponge-spicules). + + +ARCHAEOCYATHINAE. + + Etheridge, R. jnr., Trans. R. Soc. S. Austr., vol. XIII. 1890, pp. + 10-22. Taylor, T. G. Mem. Roy. Soc. S. Austr., vol. II., pt. 2, + 1910 (a monograph). + + +CORALS. + + Silurian.--Etheridge, R. jnr. Rec. Geol. Surv. New South Wales, vol. + II. pt. 1, 1890, pp. 15-21 (Silurian and Devonian). Idem, ibid., + vol. II. pt. 4, 1892, pp. 165-174 (Silurian and Devonian). Idem, + in Pal. and Geol. Queensland, 1892. Idem, Rec. Austr. Mus., vol. + I., No. 10, 1891, pp. 201-205 (_Rhizophyllum_). Id., ibid., vol. + III. No. 2, 1897, pp. 30-33 (_Columnaria_). Id., Prog. Rep. Geol. + Surv. Vict., No. 11, 1899, pp. 30-36. Idem, Mem. Geol. Surv. New + South Wales, No. 13, pt. I., 1904 (_Halysites_). Id., ibid., No. + 13, pt. 2, 1907 (_Tryplasma_). De Koninck, L. G. ibid., Pal. No. + 6, 1898. Shearsby, A. J. Geol. Mag., Dec. V., vol. III. 1906, + pp. 547-552. Chapman, F. Rec. Geol. Surv. Vict., vol. II. pt. 1, + 1907, pp. 67-80. + + Devonian.--Etheridge, R. jnr. and Foord, A. H. Ann. Mag. Nat. + Hist., ser. V., vol. XIV., 1884, pp. 175-179 (_Alveolites_ and + _Amplexopora_ = _Litophyllum_). Etheridge, R. jnr., in Geol. and + Pal. Queensland, 1892. Idem, Proc. Linn. Soc. New South Wales, + vol. IX. 1895, pp. 518-539. Id., Rec. Geol. Surv. New South + Wales, vol. VI. pt. 3, 1899, pp. 152-182 (Tamworth District). + Id., Rec. Austr. Mus., vol. IV. No. 7, 1902, pp. 253-260. De + Koninck, L. G. Mem. Geol. Surv. New South Wales, Pal. No. 6, + 1898. Chapman, F. Rec. Geol. Surv. Vict., vol. III, pt. 2, 1912, + pp. 215-222. + + Carbopermian.--Etheridge, R. jnr. Mem. Geol. Surv. New South Wales, + Pal. No. 5, 1891. Idem, in Geol. and Pal. Queensland, 1892. Id., + Bull. Geol. Surv., W. Austr., No. 10, 1903, pp. 8-10. + + Cainozoic.--Duncan, P. M. Quart. Journ. Geol. Soc., vol. XXVI. 1870, + pp. 284-318; vol. XXXI. 1875, pp. 673-678; vol. XXXII. 1876, pp. + 341-351. Woods, T. Proc. Linn. Soc. New South Wales, vol. XI., + 1878, pp. 183-195; ibid., vol. XXX. 1879, pp. 57-61. Idem, Trans. + Roy. Soc. S. Austr., vol. I., 1878, pp. 104-119. Dennant, J. + Trans. R. Soc. S. Austr., vols. XXIII. (1899) to XXVIII. (1904). + + +STROMATOPOROIDS. + + Hinde, G. J. Geol. Mag., Dec. III. vol. VII, 1890, p. 193. + + +GRAPTOLITES. + + McCoy, F. Prod. Pal. Vict., Decades I. (1874): II. (1875): V. (1877). + Hall, T. S. Proc. Roy. Soc. Vict., vol. IV. p. I. 1892, pp. 7, + 8 (_Dictyonema_). Idem, Geol. Mag. Dec. IV. vol. VI. 1899, pp. + 438-451; Id., Rep. Austr. Assoc. Adv. Sci., Brisbane, 1909, pp. + 318-320. Id., Rec. Geol. Surv. Vict., vol. I. pt. 4, 1906, pp. + 266-278. Id., ibid., vol. III. pt. 2, 1912, pp. 188-211. Idem, + Rec. Geol. Surv. New South Wales, vol. VII. part 1, 1910, pp. 16, + 17. Ibid., pp. 49-59. + + + + +CHAPTER VIII. + +FOSSIL SEA-LILIES, STARFISHES, BRITTLE-STARS AND SEA-URCHINS. + + +=Divisions of Echinodermata.--= + +The sub-kingdom of ECHINODERMATA includes the above groups comprised in +the Classes Crinoidea, Asteroidea, Ophiuroidea and Echinoidea. Besides +these are the less important classes of the Cystidea or sac-shaped +echinoderms (of which no definite remains are recorded from Australian +rocks); the Blastoidea or bud-shaped echinoderms (of which four genera are +known from Australia); the Edrioasteroidea or sessile starfishes (unknown +in Australia); and the Holothuroidea or sea-cucumbers (represented as +fossils by the skin spicules and plates, an example of which has been +recorded from Australia). + + +_CRINOIDEA, or Sea-lilies._ + + +=Crinoidea, their General Structure.--= + +These often beautiful and graceful animals resemble a starfish mounted +on a stalk. They are composed of calcareous joints and plates, and are +therefore important as rock-formers. The stalk or column may be either +short or long, and is generally rooted, in the adult stage, in the mud of +the sea-floor. Fossil Crinoids were sometimes furnished with a coiled +termination, which could be entwined around such objects as the stems of +sea-weeds. The crinoid column is composed of numerous plates, and is round +or pentagonal. Upon this is fixed the calyx or cup, with its attached +arms, which serve to bring food to the mouth, situated on the upper part +of the cup. The arms are grooved, and the water, being charged with food +particles (animalcula), pours down these channels into the mouth. The +stem elevates the animal above the mud or silt of the sea-floor, thus +making it more easy for it to obtain its food supply. The stalks of +fossil Crinoids sometimes reached the enormous length of 50 feet. Their +calcareous skeleton is built upon a plan having five planes of symmetry; +this pentamerism is found throughout the crinoids, the blastoids and +the free-moving echinoderma. Crinoids range from moderately shallow- +to deep-water, and at the present day are almost restricted to abyssal +conditions. The more ancient types usually found their habitats amongst +reefs or in comparatively clear water, where there was a marked freedom +from sediment, although that was not an essential, as seen by their +numerous remains in the Australian mudstones and sandstones. + + +=Cambrian Crinoids.--= + +The group of the Crinoidea first appears in the Upper Cambrian, and +persists to the present time. In North America the genus _Dendrocrinus_ +occurs in the Cambrian and Ordovician; and some stem-joints from the Upper +Cambrian limestone of the Mount Wellington district, Victoria, may be +provisionally referred to this genus. + +[Illustration: =Fig. 76.--FOSSIL CRINOIDS.= + + A--(?) Pisocrinus yassensis, Eth. fil. Side of calyx. Silurian. Yass, + New South Wales. + + B--(?) Pisocrinus yassensis, Eth. fil. Dorsal Surface. Silurian. N. S. + W. + + C--Botryocrinus longibrachiatus, Chapm. Silurian. Flemington, Vict. + + D--Helicocrinus plumosus, Chapm. Stem, distal end. Brunswick, Victoria. + + E--Phialocrinus konincki, Eth. fil. Carbopermian (Up. Mar. Ser.) + Nowra, New South Wales. + + F--Isocrinus australis, Moore sp. L. Cretaceous. Wollumbilla, Q'ld. + +] + + +=Ordovician Crinoids.--= + +No undoubted Crinoid remains have been found in the Australian Ordovician; +although many genera are found elsewhere in that system, chiefly +in N. America, as _Reteocrinus_, _Hybocrinus_, _Heterocrinus_ and +_Dendrocrinus_, and in Europe and North America, as _Rhodocrinus_ and +_Taxocrinus_. + + +=Silurian Crinoids.--= + +The Silurian Crinoidea of Australia are largely represented by the +remains of the columns or stalks, which are often found in such abundance +as to constitute large masses of sub-crystalline limestone, as that of +Toongabbie, Victoria. The columns of the Crinoids do not usually possess +sufficient characters to enable the forms to be identified. There are, +however, more perfect and identifiable remains of several very interesting +generic types in the Silurian faunas as follows:-- + +In New South Wales _Pisocrinus_ is represented with some reservation by +(?) _P. yassensis_, found at Limestone Creek, near Yass (Fig. 76 A, B). + +In Victoria, _Helicocrinus plumosus_ and _Botryocrinus longibrachiatus_ +occur at Brunswick and Flemington, respectively (Fig. 76). The former is +a delicate and handsome species, having a small cup with finely pinnate +arms, which are forked once, and with a pentagonal stem coiled at the +distal end (see Frontispiece). The genus _Botryocrinus_ is found in rocks +of a similar age in North America and England. _Hapalocrinus victoriae_, a +member of the Platycrinidae, has been described from the mudstone of South +Yarra, near Melbourne. The species above mentioned are of Melbournian age, +belonging to the lower stage of the Silurian system. + + +=Devonian Crinoids.--= + +In the Middle Devonian of Queensland, fragmentary crinoid stems are found +interbedded with the limestone of the Broken River. + +Thin slices of the limestone of the same age from Buchan, Victoria, show +numerous ossicles and stem-joints of Crinoids. + +Similar remains have also been recorded from the Devonian of the Kimberley +district and the Gascoyne River in Western Australia. + + +=Carboniferous Crinoids.--= + +The Carboniferous (Star Beds) of Queensland has yielded remains of +_Actinocrinus_. + +The Matai Series of New Zealand, which may be regarded as almost certainly +of Carboniferous age, contains remains of a _Cyathocrinus_, found in the +limestone of the Wairoa Gorge. + + +=Carbopermian Crinoids.--= + +The Carbopermian (Upper Marine Series) of New South Wales yields the +interesting Crinoid having a large, globular cup, known as _Phialocrinus_; +the best known species of this genus are _P. konincki_ (Fig. 76 E) and +_P. princeps_. Beds of the same age in New South Wales, also in the Upper +Marine Series, contain the aberrant Crinoid with strongly sculptured +plates of the calyx in the decorticated condition, _Tribrachiocrinus +clarkei_. + +_Poteriocrinus_ and _Platycrinus_ are, with some reservation, recorded +from the Gympie Series at Stanwell and the marine beds of the Bowen River +Coal-field respectively, both in Queensland. + +In Western Australia the Carbopermian rocks of the Gascoyne River +are known to contain crinoid stems, tentatively referred to either +the Rhodocrinidae or the Actinocrinidae. There is also a species of +_Platycrinus_ known from the Gascoyne and Irwin Rivers, and from the +Kimberley District. + + +=Triassic Crinoids.--= + +The Kaihiku Series of Nelson, New Zealand, has yielded some crinoid stems, +but the genus has not yet been determined. + + +=Cretaceous Crinoids.--= + +In the Lower Cretaceous Limestone of Queensland, at Mitchell Downs and +Wollumbilla, a typical Crinoid, closely allied to the living _Pentacrinus_ +is found, namely, _Isocrinus australis_ (Fig. 76 F). + +The Upper Cretaceous opal deposits of White Cliffs in Wilcannia, New South +Wales, contain many opalised fossil remains, amongst them being _Isocrinus +australis_, already noticed as occurring in the Lower Cretaceous of +Queensland. + + +=Cainozoic Crinoids.--= + +_Pentacrinus stellatus_ is a species founded on some deeply indented +pentagonal stem-joints found in the Oamaru Series (Miocene) at Curiosity +Shop, South Canterbury, New Zealand, and also occurring in the Chatham +Islands. This species has been identified in the Aire Coastal beds in +Victoria, of the same age. Another generic type, _Antedon_, the beautiful +"Feather Star," is frequently met with in Janjukian strata in Victoria +and South Australia, as at Batesford and Mount Gambier, represented by +the denuded crown and the ossicles of the arms of a comparatively large +species; whilst another and smaller form has been described from beds of +the same age from borings in the Victorian Mallee, under the name of _A. +protomacronema_. + + +_BLASTOIDEA--Bud-shaped Echinoderms._ + + +=Distribution and Characters of Blastoidea.--= + +This forms a small class which has a few representatives in the rocks +of Australia. Elsewhere they are chiefly of Devonian and Carboniferous +ages. In Australia they are confined, so far as known, to sediments of +the Carboniferous System. The animal was rooted to the sea-floor and a +jointed stem was usually present. The cup or theca, as before noted, is +bud-shaped, and consists of basal, radial and deltoid plates, the edges of +which are folded inwards into the thecal cavity, and thus the internal +organs came into contact with the incurrent water. The cup bears five +food grooves, bordered by numerous arms or brachioles, which directed the +incurrent particles into the thecal cavity. + + +=Carbopermian Blastoids.--= + +Three genera of blastoids have been recorded from the Gympie Beds, or +Carbopermian, of the Rockhampton District of Queensland. They are, +_Mesoblastus_, _Granatocrinus_ and _Tricoelocrinus_. A similar fossil in +beds of like age, and provisionally referred to the genus _Metablastus_, +has been lately recorded from Glenwilliam, Clarence Town, New South Wales. + + +_ASTEROIDEA, or Starfishes._ + + +=Characters of True Starfishes.--= + +These free-moving echinoderms are usually five-sided, though sometimes +star-shaped, with numerous arms surrounding a central disc. The mouth is +central on the under side of the disc, and the anus above and near the +centre (excentric), the latter being covered by a porous plate called +the madreporite. The hydraulic system of starfishes consists of tubes +extending along the grooved arms and giving off side branches which end +in processes called podia and terminating in suckers. The podia pass +through pores in the floor plates of the grooves, and communicate within +the body with distensions called ampulla. By this means the podia serve as +feet, and can be withdrawn by the expulsion of the water in them into the +ampulla. The stout flexible covering of the starfish is strengthened by +calcareous plates and bars, owing to the presence of which they are often +preserved as fossils. + +[Illustration: =Fig. 77.--FOSSIL STARFISH.= + + A--Palaeaster smythi. McCoy sp. Silurian. Flemington, Victoria. + + B--Urasterella selwyni, McCoy. Silurian. Kilmore, Victoria. + + C--Palaeaster giganteus, Eth. fil. Carbopermian. Near Farley, New + South Wales. + + D--Pentagonaster sp. Tertiary (Janjukian). Bore in Mallee, Victoria. + +] + + +=Silurian Starfishes.--= + +The oldest Australian fossil Starfishes are found in the Silurian. In +Victoria they occur in some abundance in the lower, Melbournian, series, +but appear to be absent or at all events very scarce in the upper, or +Yeringian series. The commonest genus is _Palaeaster_, of which there are +two species, _P. smythi_ (Fig. 77 A) and _P. meridionalis_, found alike in +the sandy and argillaceous strata near Melbourne. _Urasterella_ is another +genus found in the Silurian rocks near Melbourne, in which the marginal +series of plates seen in _Palaeaster_ are wanting, giving to the starfish +a slender, long-armed aspect (Fig. 77 B). + + +=Carbopermian Starfishes.--= + +In the Lower Marine Series of the Carbopermian of New South Wales a very +large species of _Palaeaster_ occurs (_P. giganteus_), measuring 7 inches +from point to point across the disc (Fig. 77 C). Two other species of the +same genus occur in this series (_P. stutchburii_ and _P. clarkei_) the +latter also ranging into the Upper Marine Series. + + +=Cainozoic Starfishes.--= + +No remains of true Starfishes have been recorded from Australia between +the Carbopermian and the Tertiary systems. In the Janjukian Series of +Victoria the marginal plates of a species of _Pentagonaster_ are typical +fossils. They have been recorded from Waurn Ponds, Spring Creek near +Torquay, and Batesford (Fig. 77 D). In the Mallee Bores, both marginal and +abactinal plates of this genus are found in polyzoal limestone (Miocene). +_Pentagonaster_ also occurs in the Lower Muddy Creek beds (Oligocene), +and the Upper beds of the same locality (Lower Pliocene). A species of +_Astropecten_ has been described from the Waikari River, New Zealand +(Oamaru Series). + + +_OPHIUROIDEA, or Brittle-stars._ + + +=Characters of Brittle-Stars.--= + +The Brittle-stars are frequently found at the present day cast up on +the fine sandy beaches of the coast. They are easily distinguished from +true starfishes by having a definite central disc, to which the arms are +attached. The arms are used for locomotion and prehension, and have their +grooves covered over with plates. The ossicles of the arms are moveable +and controlled by muscles which enable them to be used as feet. The lower +surface of the disc has a central arrangement of five rhomboidal sets of +jaws, formed of modified ossicles, called the mouth frame, whilst the +upper surface bears, between one set of arms, the madreporite or covering +plate to the water vascular system, as in starfishes. + +[Illustration: =Fig. 78.--Protaster brisingoides=, Gregory. + +Negative cast of the calcareous skeleton. Nat. size. Silurian Sandstone, +Flemington, Victoria. + + (_Nat. Mus. Coll._) +] + + +=Silurian Brittle-Stars.--= + +The Brittle-stars in Australia first appear in the Silurian, but +in England and Bohemia date back to the Ordovician. _Protaster_ is +the commonest genus, and is represented by _P. brisingoides_ of the +Melbournian stage of Silurian strata at Flemington (Fig. 78). It also +occurs rarely in the Yeringian beds at Yering, both Victorian localities. +A very ornamental form, _Gregoriura spryi_, occurs in the same division +of the Silurian at South Yarra. In this fossil the delicate spines +attached to the adambulacral ossicles are well preserved and form a +marginal fringe to the arm (Fig. 79). _Sturtzura_ is another Silurian +genus, found in the Wenlock of England and in the Melbournian of +Flemington, Victoria. + +[Illustration: =Fig. 79.--A Brittle-Star.= (Gregoriura spryi, Chapm.) + +Nat. size. From the Silurian Mudstone of South Yarra, Victoria. + + (_Nat. Mus. Coll._) +] + + +=Cainozoic Brittle-Stars.--= + +From the Victorian Cainozoic beds, in the Lower Pliocene of Grange Burn, +Hamilton, a vertebral ossicle of an ophiurian has been obtained, which has +been provisionally referred to the genus _Sigsbeia_. + + +_ECHINOIDEA, or Sea-urchins._ + +This group is an important one amongst Australian fossils, especially +those of Cainozoic age. + + +=Characters of Sea-urchins.--= + +Echinoids are animals enclosed in a spheroidal box or test composed of +numerous calcareous plates, disposed geometrically as in the Starfishes, +along five principal lines. The test in the living condition is more or +less densely covered with spines. The mouth is on the under surface. The +anus is either on the top of the test (dorso-central), or somewhere in +the median line between the two lower ambulacra. The ambulacra ("a garden +path") are the rows of perforated plates on the upper (abactinal) surface +sometimes extending to the lower surface, through which protrude the +podia, which in Starfishes are situated in grooves on the lower surface. + + +=Silurian Palaeechinoids.--= + +The Palaeechinoids are represented in the Silurian of Australia by +occasional plates, as at Bowning, New South Wales, and near Kilmore, +Victoria, whilst spines are not uncommon in certain Silurian limestones at +Tyer's River, Gippsland. + + +=Carbopermian Palaeechinoids.--= + +In the Carbopermian of New South Wales, tests of _Archaeocidaris_ have +been recorded, and also a plate of the same genus in the Gympie Beds of +Rockhampton, Queensland. + + +=Regular Echinoids.--= + +The regular Echinoids date from Permian times. They have two vertical rows +of plates for each ambulacrum and inter-ambulacrum. The mouth is on the +underside, and the anus abactinal (on the upper side) and near the centre. + +[Illustration: =Fig. 80.--CAINOZOIC SEA-URCHINS.= + + A--Cidaris (Leiocidaris) australiae, Duncan sp. Cainozoic (Janjukian). + Cape Otway, Victoria + + B--Psammechinus woodsi, Laube. Cainozoic (Janjukian). Murray River + Cliffs, S. Australia + + C--Fibularia gregata, Tate. Cainozoic (Janjukian). Aldinga, S.A. + + D--Echinocyamus (Scutellina) patella, Tate sp. Cainozoic (Janjukian). + Torquay, Victoria + + E--Clypeaster gippslandicus, McCoy. Cainozoic (Janjukian). Bairnsdale, + Victoria + + F--Studeria elegans, Laube. sp. Cainozoic (Janjukian). Murray River + Cliffs, S. Australia. + +] + + +=Cainozoic Regular Echinoids.--= + +In Australasia they make their first appearance in strata of Tertiary age, +and some species, as _Paradoxechinus novus_, range through Balcombian +strata to Kalimnan in Victoria, or Oligocene to Lower Pliocene, but are +more typically Janjukian. _Echinus_ (_Psammechinus_) _woodsi_ (Fig. 80 B) +is common in Janjukian strata in Victoria and South Australia and occurs +sparingly in the Kalimnan. Another common form of the regular Echinoids +in Southern Australia is _Cidaris australiae_ (Fig. 80 A), ranging from +Janjukian to Kalimnan, occurring more frequently in the older series. In +New Zealand a species of _Cidaris_ (_C. striata_), is known from the +Oamaru Series at Brighton. An _Echinus_ occurs in the Oamaru Series of +Broken River, and two species of that genus in the Wanganui formation of +Shakespeare Cliff. _Temnechinus macleayana_ has been recorded from the +Cainozoic (Miocene or Pliocene) of Yule Island, Papua. + + +=Irregular Echinoids.--= + +The irregular Echinoids are not known before the Upper Cretaceous in +Australia, and are very common in the Tertiaries. They are distinguished +by the anus (periproct) passing backward from the apex, as compared with +the regular forms, and by the elongation of the test and the loss of the +strong solid spines, which are replaced by thin, slender hair-like spines. +The animal is thus better fitted to burrow through the ooze on which it +feeds. + + +=Cretaceous Irregular Echinoids.--= + +An interesting form, _Micraster sweeti_, is found in the Upper Cretaceous +or Desert Sandstone of Maryborough in Queensland, which reminds one of +typical European species of this genus. + + +=Cainozoic Irregular Echinoids.--= + +Amongst the Australian Cainozoic Echinoids of the irregular type the +following may be mentioned. The little subglobular test of _Fibularia +gregata_, and _Echinocyamus_ (_Scutellina_) _patella_ (Fig. 80 C, D) are +Janjukian in age. The large _Clypeaster, C. gippslandicus_ (Fig. 80 E), +ranges from the Oligocene to Lower Pliocene in Victoria (Balcombian to +Kalimnan), and vies in size, especially in the Janjukian, with some large +species like those from Malta and Egypt. This genus includes some of the +largest known sea-urchins. The biscuit urchin, _Arachnoides (Monostychia) +australis_, is commonest in the Janjukian, but ranges from Balcombian +to Kalimnan. A common urchin from the polyzoal rock of Mt. Gambier is +_Echinolampas gambierensis_, which is also found in the Lower beds of +Muddy Creek. A typical Janjukian fossil is _Duncaniaster australiae_, +formerly thought to belong to the Cretaceous genus _Holaster_. Although +found living, the genus _Linthia_ attained its maximum development both in +size and abundance, in Janjukian or Miocene times, as seen in _L. gigas_ +(having a length of 7-1/2 inches) and _L. mooraboolensis_. _Echinoneus +dennanti_ is restricted to the Janjukian. Several species of _Eupatagus_ +occur in the Cainozoic or Tertiary beds of South Australia, Victoria and +New Zealand; _Lovenia forbesi_ (Fig. 81 C) is common in the Janjukian to +Kalimnan, both in Victoria and South Australia. In the latter State also +occur the following genera:--_Studeria_, _Cassidulus_, _Echinolampas_, +_Plesiolampas_, _Linthia_, _Schizaster_ and _Brissopsis_. In New +Zealand the following Cainozoic genera, amongst others of the irregular +sea-urchins, may be cited:--_Hemipatagus_, _Brissopsis_, _Hemiaster_, and +_Schizaster_ (Fig. 81). + +[Illustration: =Fig. 81--CAINOZOIC SEA-URCHINS.= + + A--Hemiaster planedeclivis, Gregory. Cainozoic (Janjukian). Morgan, S. + Australia + + B--Schizaster sphenoides, T. S. Hall. Cainozoic (Barwonian). + Sherbrooke River, Victoria + + C--Lovenia forbesi, T. Woods sp. Cainozoic (Janjukian). Murray River + Cliffs, S. Australia + +] + +A clypeastroid, _Peronella decagonalis_ has been described from the (?) +Lower Pliocene of Papua. + + +=Cainozoic Holothuroidea.--= + +The _HOLOTHUROIDEA_ (Sea-Cucumbers) are represented in Australian deposits +by a unique example of a dermal spicule of wheel-like form, referred to +_Chiridota_, obtained from the Cainozoic (Janjukian) beds of Torquay. This +genus is also known from the "calcaire grossier" or Middle Eocene of the +Paris Basin, and is found living in all parts of the world. + + * * * * * + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + +CRINOIDS. + + (?) _Pisocrinus yassensis_, Eth. fil. Silurian: New South Wales. + + _Helicocrinus plumosus_, Chapman. Silurian: Victoria. + + _Botryocrinus longibrachiatus_, Chapm. Silurian: Victoria. + + _Hapalocrinus victoriae_, Bather. Silurian: Victoria. + + _Actinocrinus_ sp. Carboniferous: Queensland. + + _Cyathocrinus_ sp. Carboniferous: New Zealand. + + _Phialocrinus konincki_, Clarke sp. Carbopermian: New South Wales. + + _Phialocrinus princeps_, Eth. fil. Carbopermian: New South Wales. + + _Tribrachiocrinus clarkei_, McCoy. Carbopermian: New South Wales. + + (?) _Platycrinus_ sp. Carbopermian: Queensland. + + _Platycrinus_ sp. Carbopermian: W. Australia. + + _Isocrinus australis_, Moore sp. Cretaceous: Queensland. + + _Pentacrinus stellatus_, Hutton. Miocene: New Zealand, Chatham Ids. + and Victoria. + + _Antedon protomacronema_, Chapman. Miocene: Victoria (deep borings). + + +BLASTOIDS. + + (?) _Mesoblastus australis_, Eth. fil. Carbopermian: Queensland. + + +STARFISHES. + + _Palaeaster smythi_, McCoy. Silurian: Victoria. + + _Palaeaster meridionalis_, Eth. fil. Silurian: Victoria. + + _Urasterella selwyni_, McCoy. Silurian: Victoria. + + _Palaeaster giganteus_, Eth. fil. Carbopermian (L. Mar. Ser.): New + South Wales. + + _Palaeaster clarkei_, de Koninck. Carbopermian (L. and Up. Mar. Ser.): + New South Wales. + + _Pentagonaster_ sp. Miocene: Victoria. + + _Astropecten_ sp. Miocene: New Zealand. + + +BRITTLE-STARS. + + _Protaster brisingoides_, Gregory. Silurian: Victoria. + + _Gregoriura spryi_, Chapman. Silurian: Victoria. + + _Sturtzura leptosomoides_, Chapman. Silurian: Victoria. + + (?) _Sigsbeia_ sp. Lower Pliocene: Victoria. + + +ECHINOIDS. + + _Palaeechinus_ sp. Silurian: Victoria. + + (?) _Archaeocidaris selwyni_, Eth. fil. Carbopermian: New South Wales. + + _Micraster sweeti_, Eth. fil. Cretaceous: Queensland. + + _Cidaris (Leiocidaris) australiae_, Duncan. Miocene and Lower + Pliocene: Victoria and S. Australia. + + _Cidaris striata_, Hutton. Miocene: New Zealand. + + _Echinus (Psammechinus) woodsi_, Laube sp. Miocene and L. Pliocene: + Victoria and S. Australia. + + _Temnechinus macleayana_, T. Woods. Cainozoic (? Lower Pliocene): + Papua. + + _Fibularia gregata_, Tate. Miocene: Victoria and S. Australia. + + _Echinocyamus (Scutellina) patella_, Tate sp. Oligocene to Miocene: + Victoria and S. Australia. + + _Clypeaster gippslandicus_, McCoy. Oligocene to L. Pliocene: Victoria. + + + _Arachnoides (Monostychia) australis_, Laube sp. Oligocene to L. + Pliocene: Victoria and S. Australia. + + _Echinoneus dennanti_, Hall. Miocene: Victoria. + + _Duncaniaster australiae_, Duncan sp. Miocene: Victoria. + + _Lovenia forbesi_, T. Woods sp. Miocene and L. Pliocene: Victoria and + S. Australia. + + _Hemiaster planedeclivis_, Gregory. Miocene: Victoria. + + +HOLOTHURIAN. + + _Chiridota_ sp. Miocene: Victoria. + + + * * * * * + + +LITERATURE. + + +CRINOIDS. + + Silurian.--Etheridge, R. jnr. Rec. Austr. Mus., vol. V. No. 5, 1904, + pp. 287-292 (_Pisocrinus_). Bather, F. A. Geol. Mag., Dec. + XV. vol. IV. 1897, pp. 337-345 (_Hapalocrinus_). Chapman, F. + Proc. R. Soc. Vict., vol. XV. (N.S.), pt. II. 1903, pp. 107-109 + (_Helicocrinus_ and _Botryocrinus_). Bather, F. A. Ottawa Nat., + vol. XX. No. 5, 1906, pp. 97, 98. + + Carboniferous and Carbopermian.--De Koninck, L. G. Mem. Geol. Surv. + New South Wales, Pal. No. 6, 1898, pp. 121-126. Etheridge, R. + jnr., in Geol. and Pal. Queensland, 1892, pp. 207-219. Idem, Mem. + Geol. Surv. New South Wales, Pal. No. 5, 1892, pp. 75-119. + + Cretaceous.--Moore, C. Quart. Journ. Geol. Soc., vol. XXVI. 1870, p. + 243. Etheridge, R. jnr., in Geol. and Pal. Queensland, 1892, p. + 439 (_Isocrinus_). + + Cainozoic.--Hutton, F. W. Cat. Tert. Moll. and Ech. of New Zealand, + 1873, p. 38. + + +BLASTOIDS. + + Carbopermian.--Etheridge, R. jnr., in Geol. and Pal. Queensland, 1892, + pp. 210-213. Taylor, T. G. Proc. Linn. Soc. New South Wales, + 1908, pp. 54-59 (_? Metablastus_). + + +STARFISHES. + + Silurian.--McCoy, F. Prod. Pal. Vict., Dec. I., 1874, pp. 41-43. + Etheridge, R. jnr. Rec. Austr. Mus., vol. I., No. 10, 1891, pp. + 199, 200. + + Carboniferous and Carbopermian.--Etheridge, R. jnr. Mem. Geol. Surv. + New South Wales, Pal. No. 5, pt. 2, 1892, pp. 70-75. De Koninck, + L. G. Ibid., Pal. No. 6, 1898, p. 127. + + Cainozoic.--Hall, T. S. Proc. R. Soc., Vict., vol. XV. (N.S.), pt. + I. 1902, pp. 81, 82 (_Pentagonaster_). Hutton, F. W. Cat. Tert. + Moll, and Ech. New Zealand, 1873, p. 38. + + +BRITTLE-STARS. + + Silurian.--Gregory, J. W. Geol. Mag., Dec. III. vol. VI. 1889, pp. + 24-27. Chapman, F. Proc. R. Soc. Vict., vol. XIX. (N.S.), pt. II. + 1907, pp. 21-27. + + Cainozoic.--Hall, T. S. Proc. R. Soc. Vict., vol. XV. (N.S.), pt. I. + 1902, p. 82 (cf. _Sigsbeia_). + + +ECHINOIDS. + + Silurian.--Chapman, F. Rec. Geol. Surv. Vict., vol. II. pt. 1, 1907, + pp. 77, 78. + + Carbopermian.--Etheridge, R. jnr. Mem. Geol. Surv. New South Wales, + Pal. No. 5, pt. 2, 1892, pp. 67-69. + + Cretaceous.--Etheridge, R. jnr., in Geol. and Pal. Queensland, 1892, + pp. 559, 560. + + Cainozoic.--T. Woods. Trans. Adelaide Phil. Soc., 1867. Laube, G. C. + Sitz, k. k. Ak. Wiss. Wien, vol. LIX. 1869, pp. 183-198. Hutton, + F. W. Cat. Tert. Moll, and Ech. New Zealand, 1873, pp. 38-43. + Duncan, P. M. Quart. Journ. Geol. Soc., vol. XXXIII. 1877, pp. + 42-73. Tate, R. Quart. Journ. Geol. Soc., vol. XXXIII. 1877, pp. + 256-258. Idem, Southern Science Record, 1885, p. 4. Idem, Trans. + R. Soc. S. Austr., vol. XIV. pt. 2, 1891, pp. 270-282. McCoy, F. + Prod. Pal. Vict., Dec. VI. VII. 1879, 1883. Gregory, J. W. Geol. + Mag., Dec. III. vol. VII. 1890, pp. 481-492. Ibid., Dec. III. + vol. IX. 1892, pp. 433-437. Cotteau, G. H. Mem. Zool. France, + vol. II. No. 4, 1889, p. 228; vol. III. No. 5, 1890, pp. 537-550; + vol. IV. No. 5, 1891, pp. 620-633. Bittner, A. Sitz. k.k. Ak. + Wiss. Wien, 1892, vol. 101, pp. 331-371. Hall, T. S. Proc. Roy. + Soc. Vic., vol. XIX. (N.S.), pt. II. 1906, pp. 48, 53. Chapman, + F. Proc. Roy. Soc. Vict., vol. XX. (N.S.), pt. II. 1908, pp. + 214-218. Pritchard, G. B. ibid., vol. XXI. (N.S.), pt. I. 1908, + pp. 392-400. + + +HOLOTHURIAN. + + Cainozoic.--Hall, T. S. Proc, R. Soc. Vict., vol. X. (N.S.), pt. I. + 1902, pp. 82, 83. + + + + +CHAPTER IX. + +FOSSIL WORMS, SEA-MATS and LAMP-SHELLS. + + +The first-named group, the ringed worms, belong to the phylum Annelida, +so-called because of the ring-like structure of their bodies. The +two remaining groups, the Polyzoa or Sea-mats and the Brachiopods or +Lamp-shells, are comprised in the phylum Molluscoidea, or mollusc-like +animals. + + +_WORMS (Annelida)._ + + +=Annelida and their Fossil Representatives.--= + +These animals, owing to the scarcity of hard parts within their bodies, +play a rather insignificant role as a fossil group. Worms are laterally +symmetrical animals, with a dorsal and a ventral surface. They are +segmented, the body being formed of numerous rings. Only those of the +Class Chaetopoda ("bristle-feet") are represented by identifiable fossil +remains. Fossil worms, moreover, chiefly belong to the Order Polychaeta +("many bristles"). The horny jaws of these worms are sometimes found in +the older rocks and are known as conodonts. + + +=Silurian Conodonts.--= + +Conodonts belonging to three genera are known from Australia. They are all +from the Silurian of the Bowning District, near Yass, New South Wales, and +are referred to the genera _Eunicites_, _Oenonites_ and _Arabellites_. + +[Illustration: =Fig. 82--FOSSIL WORMS.= + + A--Trachyderma crassituba, Chapm, Silurian. South Yarra, Vict. + + B--Cornulites tasmanicus, Eth. fil. Silurian. Heazlewood, Tas. + + C--Spirorbis ammonius, M. Edwards, var. truncata, Mid. Devonian. + Buchan, Victoria + + D--Torlessia mackayi, Bather. ? Trias. Mt. Torlesse, N. Zealand. + +] + + +=Palaeozoic Errant Worms.--= + +The wandering Worms (Polychaeta errantia) are also recognised by their +impressions, trails, borings and castings. Burrows formed by these worms +are seen in _Arenicolites_, found in the Silurian sandstone of New +South Wales, near Yass, and in the Carbopermian (Gympie Series) near +Rockhampton, Queensland. The membranous-lined burrows of _Trachyderma_ +(_T. crassituba_), occur in some abundance in the Silurian mudstones +in the neighbourhood of Melbourne, Victoria (Fig. 82 A). The genus +_Trachyderma_ is common also to Great Britain and Burmah, in beds of the +same age. + + +=Worm Tracks.--= + +Some of the curious markings on the Carboniferous sandstone of Mansfield, +Victoria, may be due to worm trails and castings, especially since they +are associated with sun-cracks and ripple-marks. + + +=Sedentary Worms.--= + +The sedentary or tube-making Worms (Polychaeta tubicola) are represented +by numerous forms. The long conical tube of _Cornulites tasmanicus_ is +recorded from the Silurian of Zeehan, Tasmania (Fig. 82 B). _Spirorbis_ +occurs in the Middle Devonian of Victoria (Fig. 82 C), and W. Australia, +and also in the Carbopermian of W. Australia. _Torlessia_ is found in +the Trias or Lower Jurassic of the province of Canterbury, New Zealand +(Fig. 82 D). The genus _Serpula_ is widely distributed, occurring in the +Carbopermian (Upper Jurassic Series), near East Maitland, New South Wales +(_S. testatrix_), in the Jurassic of W. Australia (_S. conformis_), in the +Lower Cretaceous of Wollumbilla, Queensland (_S. intestinalis_), and the +Darling River, north west of New South Wales, (_S. subtrachinus_), as well +as in Cainozoic deposits in Victoria (_S. ouyenensis_). _Ditrupa_ is very +abundant in some shelly deposits of Janjukian age in Victoria. + + +MOLLUSCOIDEA. + +The Sea-mats (Polyzoa) and the Lamp-shells (Brachiopoda) constitute a +natural group, the MOLLUSCOIDEA, which, although unlike in outward form, +have several physiological structures in common. The respiratory organs +lie in front of the mouth, and are in the form of fleshy tentacles or +spiral appendages. These animals are more nearly allied to the worms than +to the molluscs. + + +_POLYZOA._ + + +=Characters of Polyzoa.--= + +These are almost exclusively marine forms, and are important as fossils. +They form colonies (polypary or zoarium), and by their branching, +foliaceous or tufty growth resemble sea-weeds. The cells in which the +separate zoöids lived have peculiar characters of their own, which serve +to distinguish the different genera. + + +=Subdivisions of Polyzoa.--= + +Polyzoa are divided into the Sub-classes Phylactolaemata, in which the +mouth of the zoöid has a lip, and the series of tentacles is horse-shoe +shaped; and the Gymnolaemata, in which there is no lip to the mouth, and +the tentacles form a complete circle. The first group forms its polypary +of soft or horny material, which is not preserved fossil. The latter +has a calcareous polypary, and is of much importance as a fossil group. +This latter sub-class is further subdivided into the following Orders, +viz.:--Trepostomata ("turned mouths"), Cryptostomata ("hidden mouths"), +Cyclostomata ("round mouths"), and Cheilostomata ("lip mouths" furnished +with a moveable operculum). + + +=Trepostomata (Palaeozoic).--= + +The Order Trepostomata may include some genera as _Monticulipora_ and +_Fistulipora_, previously referred to under the corals. They become +extinct after Permian times. _Fistulipora_ occurs in certain Gippsland +limestones. + +[Illustration: =Fig. 83--PALAEOZOIC POLYZOA.= + + A--Fenestella margaritifera, Chapm. Silurian. Near Yering, Vict. + + B--Polypora australis, Hinde. Carbopermian. Gascoyne River, Western + Australia + + C--Rhombopora tenuis, Hinde. Carbopermian. Gascoyne River, Western + Australia + + D--Protoretepora ampla, Lonsdale sp. Carbopermian. N.S.W. + +] + + +=Cryptostomata (Palaeozoic).--= + +In the order Cryptostomata we have the genus _Rhombopora_ with its long, +slender branches, which occurs in the Silurian of Victoria and the +Carbopermian of Queensland and W. Australia (Fig. 83 C). Of this order a +very important Australian genus is _Fenestella_, the funnel-shaped zoaria +of which are found in the Silurian of Victoria and New South Wales, and +also in the Carboniferous of the latter State. _Fenestella_ also occurs in +the Carbopermian of W. Australia and Tasmania (Fig. 83 A). Accompanying +the remains of _Fenestella_ in the Carbopermian rocks, and closely related +to it, are found the genera _Protoretepora_ and _Polypora_ (Fig. 83 B, D). + +Polyzoa have been noticed in Jurassic rocks in W. Australia, but no +species have been described. + +[Illustration: =Fig. 84--CAINOZOIC POLYZOA.= + + A--Lichenopora australis, MacGillivray. Balcombian. Hamilton, Victoria + + B--Heteropora pisiformis, MacGillivray. Janjukian. Moorabool, Victoria + + C--Cellaria australis, MacGillivray. Balcombian. Hamilton, Vict. + + D--Selenaria cupola, T. Woods sp. Balcombian. Hamilton, Vict. + + E--Lepralia elongata, MacGill. Balcombian. Hamilton, Victoria + +] + + +=Cheilostomata (Cretaceous).--= + +Species of the genera (?) _Membranipora_ and (?) _Lepralia_, belonging to +the Cheilostomata, have been described from the Lower Cretaceous of the +Darling River, New South Wales, and Wollumbilla, Queensland, respectively. + + +=Cainozoic Polyzoa.--= + +A very large number of genera of the Polyzoa have been described from +the Tertiary strata of South Australia and Victoria. Some of the +principal of these are _Crisia_, _Idmonea_, _Stomatopora_, _Lichenopora_, +_Hornera_, _Entalophora_ and _Heteropora_ of the order Cyclostomata; and +_Catenicella_, _Cellaria_, _Membranipora_, _Lunulites_, _Selenaria_, +_Macropora_, _Tessarodoma_, _Adeona_, _Lepralia_, _Bipora_, _Smittia_, +_Porina_, _Cellepora_ and _Retepora_ of the order Cheilostomata. Many of +these genera, and not a few Australian species, are found also in the +Cainozoic or Tertiary beds of Orakei Bay, New Zealand (Fig. 84). + + +_BRACHIOPODA (Lamp-shells)._ + + +=Brachiopods: Their Structure.--= + +These are marine animals, and are enclosed in a bivalved shell. They +differ, however, from true bivalves (Pelecypoda) in having the shell +on the back and front of the body, instead of on each side as in the +bivalved mollusca. Each valve is equilateral, but the valves differ from +one another in that one is larger and generally serves to attach the +animal to rocks and other objects of support by a stalk or pedicle. Thus +the larger valve is called the pedicle valve and the smaller, on account +of its bearing the calcareous supports for the brachia or arms, the +brachial valve. Generally speaking, the shell of the valve is penetrated +by numerous canals, which give the shell a punctate appearance. Some +brachiopod shells, as _Atrypa_ and _Rhynchonella_, are, however, devoid of +these. + +[Illustration: =Fig. 85--LOWER PALAEOZOIC BRACHIOPODS.= + + A--Orthis (?) lenticularis, Wahlenberg. Up. Cambrian. Florentine + Valley, Tasmania + + B--Siphonotreta maccoyi, Chapm. Up. Ordovician. Bulla, Vict. + + C--Lingula yarraensis, Chapm. Silurian. South Yarra, Victoria + + D--Orbiculoidea selwyni, Chapm. Silurian. Merri Creek, Victoria + + E--Chonetes melbournensis, Chapm. Silurian. South Yarra, Vict. + + F--Stropheodonta alata, Chapm. Silurian. Near Lilydale, Vict. + +] + + +=Cambrian Brachiopods.--= + +Brachiopods are very important fossils in Australasian rocks. They first +appear in Cambrian strata, as for example, in the Florentine Valley, in +Tasmania, where we find _Orthis lenticularis_ (Fig. 85 A). In Victoria, +near Mount Wellington, in the mountainous region of N.E. Gippsland, +_Orthis platystrophioides_ is found in a grey limestone. In South +Australia the grey Cambrian limestone of Wirrialpa contains the genus +_Huenella_ (_H. etheridgei_). This genus is also found in the Middle and +Upper Cambrian of N. America. + + +=Ordovician Brachiopods.--= + +Coming to Ordovician rocks, the limestones of the Upper Finke Basin in +South Australia contain _Orthis leviensis_ and _O. dichotomalis_. The +Victorian mudstone at Heathcote may be of Ordovician age or even older; it +has afforded a limited fauna of brachiopods and trilobites, amongst the +former being various species of _Orthis_, _Chonetes_, and _Siphonotreta_. +The latter genus is represented in both the Lower and Upper Ordovician +rocks of slaty character in Victoria (Fig. 85 B). + + +=Silurian Brachiopods.--= + +The Silurian system in Australasia as in Europe, N. America and elsewhere, +is very rich in brachiopod life. It is impossible to enumerate even all +the genera in a limited work like the present, the most typical only being +mentioned. + +In New Zealand the palaeozoic fauna is at present imperfectly worked +out, but the following genera from the Wangapekian (Silurian) have been +identified, viz., _Chonetes_, _Stricklandinia_, _Orthis_, _Wilsonia_, +_Atrypa_, and _Spirifer_. The specific identification of these forms with +European types is still open to question, but the species are undoubtedly +closely allied to some of those from Great Britain and Scandinavia. + +The Victorian Silurian Brachiopods are represented by the horny-shelled +_Lingula_, the conical _Orbiculoidea_, a large species of _Siphonotreta_, +_Stropheodonta_ (with toothed hinge-line), _Strophonella_, _Chonetes_ +(with hollow spines projecting from the ventral valve, one of the species +_C. melbournensis_ being characteristic of the Melbournian division of +Silurian rocks), _Orthis_, _Pentamerus_, _Camarotoechia_, _Rhynchotrema_, +_Wilsonia_, _Atrypa_ (represented by the world-wide _A. reticularis_), +_Spirifer_ and _Nucleospira_ (Figs. 85, 86). + +New South Wales has a very similar assemblage of genera; whilst Tasmania +possesses _Camarotoechia_, _Stropheodonta_ and _Orthis_. + + +=Devonian Brachiopods.--= + +The Devonian limestones and associated strata are fairly rich in +Brachiopods. The Victorian rocks of this age at Bindi and Buchan contain +genera such as _Chonetes_ (_C. australis_), _Spirifer_ (_S. yassensis_ and +_S. howitti_) and _Athyris_. + +In New South Wales we again meet with _Spirifer yassensis_, veritable +shell-banks of this species occurring in the neighbourhood of Yass, +associated with a species of _Chonetes_ (_C. culleni_) (Fig. 86 D, E). + +[Illustration: =Fig. 86--SILURIAN and DEVONIAN BRACHIOPODS.= + + A--Camarotoechia decemplicata, Sow. Silurian. Victoria + + B--Nucleospira australis, McCoy. Silurian. Victoria + + C--Atrypa reticularis, L. sp. Silurian. Victoria + + D--Chonetes culleni, Dun. Mid. Devonian. New South Wales + + E--Spirifer yassensis, de Koninck. Devonian. New South Wales and + Victoria + +] + +In the Upper Devonian of New South Wales abundant remains occur of both +_Spirifer disjunctus_ and _Camarotoechia pleurodon_ (var.). + +The Upper Devonian Series at Nyrang Creek near Canowindra, New South +Wales, contains a _Lingula_ (_L. gregaria_) associated with the +_Lepidodendron_ plant beds of that locality. + +Queensland Devonian rocks contain _Pentamerus_, _Atrypa_ and _Spirifer_. +In Western Australia the Devonian species are _Atrypa reticularis_, +_Spirifer_ cf _verneuili_, _S. musakheylensis_ and _Uncinulus_ cf. +_timorensis_. + + +=Carboniferous Brachiopods.--= + +The Carboniferous Brachiopod fauna is represented in New South Wales at +Clarence Town and other localities by a species which has an extensive +time-range, _Leptaena rhomboidalis_ var. _analoga_, and the following, +a few of which extend upwards into the Carbopermian:--_Chonetes +papilionacea_, _Productus semireticulatus_, _P. punctatus_, _P. cora_, +_Orthothetes crenistria_, _Orthis (Rhipidomella) australis_, _O. +(Schizophoria) resupinata_, _Spirifer striatus_, _S. bisulcatus_, _Cyrtina +carbonaria_ and _Athyris planosulcatus_. + +In New Zealand the Matai series, referred to the Jurassic by Hutton, as +formerly regarded by Hector, and latterly by Park, as of Carboniferous +age, on the ground of a supposed discovery of _Spirifer subradiatus_ (_S. +glaber_) and _Productus brachythaerus_ in the Wairoa Gorge. Although these +species may not occur, the genera _Spirifer_ and _Productus_ are present, +which, according to Dr. Thomson, are distinctly of pre-Triassic types. + +[Illustration: =Fig. 87--CARBOPERMIAN BRACHIOPODS.= + + A--Productus brachythaerus, Sow. Carbopermian. New South Wales, &c. + + B--Strophalosia clarkei, Eth. sp. Carbopermian. N.S.W., &c. + + C--Spirifer convolutus, Phillips. Carbopermian. N.S.W., &c. + + D--Spirifer (Martiniopsis) subradiatus, Sow. Carbopermian. New South + Wales, &c. + +] + + +=Carbopermian Brachiopods.--= + +The Brachiopod fauna of Carbopermian age in New South Wales is rich +in species of _Productus_ and _Spirifer_. Amongst the former are _P. +cora_ (also found in Western Australia, Queensland and Tasmania), _P. +brachythaerus_ (also found in Western Australia and Queensland), (Fig. 87 +A), _P. semireticulatus_ (also found in Western Australia, Queensland and +the Island of Timor, and a common species in Europe), and _P. undatus_ +(also found in Western Australia and Queensland, as well as in Great +Britain and Russia). _Strophalosia_ is an allied genus to _Productus_. +It is a common form in beds of the same age in W. Australia, Tasmania, +and New South Wales. The best known species is _S. clarkei_ (Fig. 87 B). +This type of shell is distinguished from _Productus_ in being cemented by +the umbo of the ventral valve, which valve is also generally less spinose +than the dorsal. When weathered the shells present a peculiar silky or +fibrous appearance. The genus _Spirifer_ is represented in W. Australia +by such forms as _S. vespertilio_, _S. convolutus_, _S. hardmani_, _S. +musakheylensis_, and _S. striatus_; whilst _S. vespertilio_ and _S. +convolutus_ are common also to New South Wales (Fig. 87 C), and the latter +only to Tasmania. _S. vespertilio_ is found in the Gympie beds near +Rockhampton, Queensland; and _S. tasmaniensis_ in Queensland (Bowen River +Coal-field, Marine Series), New South Wales and Tasmania. Of the smoother, +stout forms, referred to the sub-genus _Martiniopsis_, we may mention _S. +(M.) subradiatus_, which occurs in W. Australia, New South Wales, and +Tasmania (Fig. 87 D). + +In the Queensland fauna, the Gympie series contains, amongst other +Brachiopods _Productus cora_, _Leptaena rhomboidalis_ var., _analoga_, +_Spirifer vespertilio_ and _S. strzeleckii_. + +Other Carbopermian Brachiopod genera found in Australian faunas are +_Cleiothyris_, _Dielasma_, _Hypothyris_, _Reticularia_, _Seminula_, +_Cyrtina_, and _Syringothyris_. + + +=Triassic Brachiopods.--= + +The Kaihiku Series of New Zealand (Hokonui Hills and Nelson) are probably +referable to the Trias. The supposed basal beds contain plants such as +_Taeniopteris_, _Cladophlebis_, _Palissya_ and _Baiera_. Above these +are marine beds containing Brachiopods belonging to _Spiriferina_, +_Rhynchonella_, _Dielasma_ and _Athyris_. The succession of these beds +presents some palaeontological anomalies still to be explained, for the +flora has a decided leaning towards a Jurassic facies. + +Next in order of succession the Wairoa Series, in the Hokonui Hills and +Nelson, New Zealand, contains _Dielasma_ and _Athyris wreyi_. + +The succeeding series in New Zealand, the Otapiri, or Upper Triassic +contains the Brachiopod genera _Athyris_[3] and _Spiriferina_, found at +Well's Creek, Nelson. + +[Footnote 3: Referred by Hector to a new sub-genus _Clavigera_, which +name, however, is preoccupied.] + + +=Jurassic Brachiopods.--= + +[Illustration: =Fig. 88--MESOZOIC BRACHIOPODS.= + + A--Rhynchonella variabilis Schloth. sp. Jurassic. W. Australia + + B--Terebratella davidsoni, Moore. L. Cretaceous. Queensland + + C--Lingula subovalis, Davidson. L. Cretaceous. S. Australia + + D--Rhynchonella croydonensis, Eth. fil. Up. Cretaceous. Queensland + +] + +The marine Jurassic beds of W. Australia, as at Shark Bay and Greenough +River, contain certain _Rhynchonellae_ allied to European species, as _R. +variabilis_ (Fig. 88 A), and _R._ cf. _solitaria_. + +=Lower Cretaceous Brachiopods.--= + +The Lower Cretaceous or Rolling Downs Formation of Queensland has +yielded a fair number of Brachiopods, principally from Wollumbilla,--as +_Terebratella davidsoni_ (Fig. 88 B), (?) _Argiope wollumbillensis_, (?) +_A. punctata_, _Rhynchonella rustica_, _R. solitaria_, _Discina apicalis_ +and _Lingula subovalis_. From beds of similar age in Central South +Australia and the Lake Eyre Basin _Lingula subovalis_ (Fig. 88 C), and +_Rhynchonella eyrei_ have been recorded; the latter has been compared with +a species (_R. walkeri_) from the Middle Neocomian of Tealby in Yorkshire. + + +=Upper Cretaceous Brachiopod.--= + +A solitary species of the Brachiopoda occurs in the Upper Cretaceous of +Australia, namely, _Rhynchonella croydonensis_ (Fig. 88 D) of the Desert +Sandstone of the Croydon Gold-fields and Mount Angas, Queensland. + + +=Cainozoic Brachiopods.--= + +The Brachiopoda of the Cainozoic or Tertiary strata of Australia +and New Zealand are well represented by the genera _Terebratula_, +_Magellania_, _Terebratulina_, _Terebratella_, _Magasella_ and +_Acanthothyris_. In the Balcombian or Oligocene of southern Australia +occur the following:--_Terebratula tateana_, _Magellania corioensis_, +_M. garibaldiana_ and _Magasella compta_ (Figs. 89 A, D); and most of +these range into the next stage, the Janjukian, whilst some extend even +to the Kalimnan. _Terebratulina suessi_, Hutton sp. (= _T. scoulari_, +Tate) ranges through the Balcombian and Janjukian, but is most typical +of the Janjukian beds in Victoria: it also occurs in the Oamaru Series +of New Zealand (= Janjukian). _Acanthothyris squamosa_ (Fig. 89 F) is +typical of the Janjukian of southern Australia, and it occurs also in +the Pareora beds of the Broken River, New Zealand. The latter are green, +sandy, fossiliferous strata immediately succeeding the Oamaru stone of the +Hutchinson Quarry beds. _A. squamosa_ is said to be still living south of +Kerguelen Island. _Magellania insolita_ is a Victorian species which is +also found in the Oamaru Series of New Zealand. + +[Illustration: =Fig. 89--CAINOZOIC BRACHIOPODS.= + + A--Terebratula tateana, T. Woods. Cainozoic. Victoria + + B--Magellania corioensis, McCoy, sp. Cainozoic. Victoria + + C--Magellania garibaldiana, Dav. sp. Cainozoic. Victoria + + D--Magasella compta, Sow. sp. Cainozoic. Victoria + + E--Terebratulina catinuliformis, Tate. Cainozoic. S. Australia + + F--Acanthothyris squamosa, Hutton sp. Cainozoic. Tasmania + +] + +Whilst many of the older Tertiary brachiopods range into the next +succeeding stage of the Kalimnan in Victoria, such as _Magellania +insolita_, _Terebratulina_ catinuliformis_ (Fig. 89 E) and _Magasella +compta_, one species, _Terebratella pumila_, is restricted to the +Kalimnan, occurring at the Gippsland Lakes. + +The next stage, the Werrikooian, typical in upraised marine beds on the +banks of the Glenelg River in western Victoria, contains _Magellania +flavescens_, a species still living (see _antea_, Fig. 23), and _M. +insolita_, having the extraordinarily wide range of the whole of the +Cainozoic stages in southern Australia. + + * * * * * + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + +WORMS. + + _Eunicites mitchelli_, Eth. fil. Silurian: New South Wales. + + _Oenonites hebes_, Eth. fil. Silurian: New South Wales. + + _Arabellites bowningensis_, Eth. fil. Silurian: New South Wales. + + _Arenicolites_ sp. Silurian: New South Wales. + + _Trachyderma crassituba_, Chapm. Silurian: Victoria. + + _Cornulites tasmanicus_, Eth. fil. Silurian: Tasmania. + + _Spirorbis ammonius_, M. Edw. var. _truncata_, Chapm. Mid. Devonian: + Victoria. + + _Spirorbis omphalodes_, Goldfuss. Devonian: W. Australia. + + _Serpula testatrix_, Eth. fil. Carbopermian: New South Wales. + + _Torlessia mackayi_, Bather. Lower Mesozoic: New Zealand. + + _Serpula conformis_, Goldfuss. Jurassic: W. Australia. + + _Serpula intestinalis_, Phillips. Lower Cretaceous: Queensland. + + _Serpula subtrachinus_, Eth. fil. Lower Cretaceous: New South Wales. + + _Serpula ouyenensis_, Chapm. Cainozoic: Victoria. + + _Ditrupa cornea_, L. sp. var. _wormbetiensis_, McCoy. Cainozoic: + Victoria. + + +POLYZOA. + + _Rhombopora gippslandica_, Chapm. Silurian: Victoria. + + _Fenestella australis_, Chapm. Silurian: Victoria. + + _Protoretepora ampla_, Lonsdale. Carbopermian: W. Australia, New South + Wales, Queensland, and Tasmania. + + _Polypora australis_, Hinde. Carbopermian: W. Australia. + + _Rhombopora tenuis_, Hinde. Carbopermian: W. Australia. + + _Rhombopora laxa_, Etheridge sp. Carbopermian: Queensland. + + _Membranipora wilsonensis_, Eth. fil. Lower Cretaceous: New South + Wales. + + (?) _Lepralia oolitica_, Moore. Lower Cretaceous: Queensland. + + _Lichenopora australis_, MacGillivray. Cainozoic: Victoria. + + _Heteropora pisiformis_, MacGillivray. Cainozoic: Victoria. + + _Cellaria australis_, MacGillivray. Cainozoic: Victoria. + + _Membranipora macrostoma_, Reuss. Cainozoic: Victoria (also living). + + _Selenaria marginata_, T. Woods. Cainozoic: Victoria (also living). + + _Macropora clarkei_, T. Woods sp. Cainozoic: Victoria. + + _Adeona obliqua_, MacGill. Cainozoic: Victoria. + + _Lepralia burlingtoniensis_, Waters. Cainozoic: Victoria. + + _Bipora philippinensis_, Busk sp. Cainozoic: Victoria (also living). + + _Porina gracilis_, M. Edwards sp. Cainozoic: Victoria (also living). + + _Cellepora fossa_, Haswell, sp. Cainozoic: Victoria (also living). + + _Retepora fissa_, MacGill. sp. Cainozoic: Victoria (also living). + + +BRACHIOPODA. + + _Orthis lenticularis_, Wahlenberg sp. Cambrian: Tasmania. + + _Orthis platystrophioides_, Chapm. Cambrian: Victoria. + + _Huenella etheridgei_, Walcott. Cambrian: S. Australia. + + _Orthis leviensis_, Eth. fil. Ordovician: S. Australia, (?) Victoria. + + _Siphonotreta discoidalis_, Chapm. Ordovician: Victoria. + + _Siphonotreta maccoyi_, Chapm. Ordovician: Victoria. + + _Lingula yarraensis_, Chapm. Silurian: Victoria. + + _Orbiculoidea selwyni_, Chapm. Silurian: Victoria. + + _Chonetes melbournensis_, Chapm. Silurian: Victoria. + + _Stropheodonta alata_, Chapm. Silurian: Victoria. + + _Orthis elegantula_, Dalman. Silurian: Victoria. + + _Pentamerus australis_, McCoy. Silurian: Victoria and New South Wales. + + _Conchidium knightii_, Sow. sp. Silurian: Victoria and New South Wales. + + _Camarotoechia decemplicata_, Sow. sp. Silurian: Victoria. + + _Rhynchotrema liopleura_, McCoy sp. Silurian: Victoria. + + _Atrypa reticularis_, L. sp. Silurian: New South Wales and Victoria. + Devonian: New South Wales, W. Australia and Queensland. + + _Spirifer sulcatus_, Hisinger sp. Silurian: Victoria. + + _Nucleospira australis_, McCoy. Silurian: Victoria. + + _Chonetes australis_, McCoy. Mid. Devonian: Victoria. + + _Chonetes culleni_, Dun. Mid. Devonian: New South Wales. + + _Spirifer yassensis_, de Koninck. Mid. Devonian: New South Wales and + Victoria. + + _Spirifer_ cf. _verneuili_, de Kon. Mid. Devonian: New South Wales and + W. Australia. + + _Lingula gregaria_, Eth. fil. Upper Devonian: New South Wales. + + _Spirifer disjunctus_, Sow. Up. Devonian: New South Wales. + + _Productus cora_, d'Orb. Carboniferous: New South Wales and Queensland. + + _Orthothetes crenistria_, Sow. sp. Carboniferous: New South Wales. + + _Spirifer striatus_, Sow. Carboniferous: New South Wales. + + _Productus brachythaerus_, Sow. Carbopermian: New South Wales, + Queensland, W. Australia. + + _Strophalosia clarkei_, Eth. sp. Carbopermian: New South Wales, + Tasmania and W. Australia. + + _Spirifer (Martiniopsis) subradiatus_, Sow. Carbopermian: New South + Wales, Tasmania and W. Australia. + + _Spirifer convolutus_, Phillips. Carbopermian: New South Wales, + Tasmania and W. Australia. + + _Cleiothyris macleayana_, Eth. fil. sp. Carbopermian: W. Australia. + + _Dielasma elongata_, Schlotheim sp. Trias (Kaihiku Series): New + Zealand. + + _Athyris wreyi_, Suess sp. Trias (Wairoa Series): New Zealand. + + _Athyris_ sp. Trias (Otapiri Series): New Zealand. + + _Rhynchonella variabilis_, Schlotheim sp. Jurassic: W. Australia. + + _Terebratella davidsoni_, Moore. Lower Cretaceous: Queensland. + + _Rhynchonella solitaria_, Moore. Lower Cretaceous: Queensland. + + _Lingula subovalis_, Davidson. Lower Cretaceous: Queensland and S. + Australia. + + _Rhynchonella croydonensis_, Eth. fil. Upper Cretaceous: Queensland. + + _Terebratula tateana_, T. Woods. Cainozoic (Balcombian and Janjukian): + Victoria and S. Australia. + + _Magellania corioensis_, McCoy, sp. Cainozoic (Balcombian and + Janjukian): Victoria and S. Australia. + + _Magellania garibaldiana_, Davidson sp. Cainozoic (Balcombian and + Janjukian): Victoria and S. Australia. + + _Magasella compta_, Sow. sp. Cainozoic (Balcombian to Kalimnan): + Victoria and S. Australia. + + _Terebratula suessi_, Hutton sp. Cainozoic (Balcombian and Janjukian): + Victoria, S. Australia, and New Zealand (Oamaru Series.) + + _Acanthothyris squamosa_, Hutton sp. Cainozoic (Janjukian): Victoria + and S. Australia, New Zealand (Oamaru Series) (also living). + + _Terebratella pumila_, Tate. Cainozoic (Kalimnan): Victoria. + + _Magellania flavescens_, Lam. sp. Pleistocene: Victoria (also living). + + + * * * * * + + +LITERATURE. + + +WORMS. + + Silurian.--Etheridge, R. jnr. Geol. Mag., Dec. III. vol. VII. 1890, + pp. 339, 340. Idem, Proc. Roy. Soc. Tas. (for 1896), 1897, p. 37. + Chapman, F. Proc. R. Soc. Vict., vol. XXII. (N.S.), pt. II. 1910, + pp. 102-105. + + Devonian.--Hinde, G. J. Geol. Mag., Dec. II. vol. VII. 1890, p. 199. + Chapman, F. Rec. Geol. Surv. Vict., vol. III. pt. 2, 1912, p. 220. + + Carboniferous.--Etheridge, R. jnr. Bull. Geol. Surv. W. Australia, No. + 10, 1903, p. 10. + + Carbopermian.--Etheridge, R. jnr. Mem. Geol. Surv. New South Wales. + Pal. No. 5, 1892, pp. 119-121. + + Lower Mesozoic.--Bather, F. A. Geol. Mag., Dec. V. vol. II. 1905, pp. + 532-541. + + Lower Cretaceous.--Etheridge, R. jnr. Mem. Soc. Geol. Surv. New South + Wales, Pal. No. 11. 1902, pp. 12, 13. + + Cainozoic.--Chapman, F. Proc. R. Soc. Vict., vol. XXVI. (N.S.) pt. I. + 1913, pp. 182-184. + + +POLYZOA. + + Silurian.--Chapman, F. Proc. R. Soc. Vict., vol. XVI. (N.S.), pt. I. + 1903, pp. 61-63. Idem, Rec. Geol. Surv. Vic., vol. II., pt. 1, + 1907, p. 78. + + Carboniferous.--Hinde, G. J. Geol. Mag. Dec. III. vol. VII. 1890, pp. + 199-203. + + Carbopermian.--De Koninck Mem. Geol. Surv. New South Wales, Pal. No. + 6, 1898, pp. 128-140. + + Cainozoic.--Stolicka, F. Novara Exped., Geol. Theil., vol. I. pt. 2, + pp. 87-158. Waters, A. W. Quart. Journ. Geol. Soc., vol. XXXVII. + 1881, pp. 309-347; ibid., vol. XXXVIII. 1882, pp. 257-276 and pp. + 502-513; ibid., vol. XXXIX. 1883, pp. 423-443; ibid., vol. XL. + 1884, pp. 674-697; ibid., vol. XLI. 1885, pp. 279-310; ibid., + vol. XLIII. 1887, pp. 40-72 and 337-350. MacGillivray, P. H. + Mon. Tert. Polyzoa Vict., Trans. Roy. Soc. Vict., Vol. IV. 1895. + Maplestone, C. M. "Further Descr. Polyzoa Vict.," Proc. Roy. Soc. + Vict., vol. XI. (N.S.), pt. I. 1898, pp. 14-21, et seqq. + + +BRACHIOPODA. + + Cambrian.--Tate, R. Trans. R. Soc. S. Austr., vol. XV. 1892, pp. 185, + 186. Etheridge, R. jnr. Rec. Austr. Mus., vol. V. pt. 2, 1904, p. + 101. Walcott, C. D. Smiths. Misc. Coll., vol. LIII. 1908, p. 109. + Chapman, F. Proc. R. Soc. Vic., vol. XXIII. (N.S.), pt. I. 1911, + pp. 310-313. + + Ordovician.--Etheridge, R. jnr. Parl. Papers, S. Aust., No. 158, 1891, + pp. 13, 14. Tate, R. Rep. Horn Exped., pt. 3, 1896, pp. 110, 111. + Chapman, F. Rec. Geol. Surv. Vict., vol. I. pt. 3, 1904, pp. + 222-224. + + Silurian.--McCoy, F. Prod. Pal. Vic. Dec. V. 1877, pp. 19-29. Eth., R. + jnr. Rec. Geol. Surv. New South Wales, vol. 3, pt. 2, 1892, pp. + 49-60 (Silurian and Devonian _Pentameridae_). Idem, Proc. Roy. + Soc., Tas., (for 1896), 1897, pp. 38-41. De Koninck, L. G. Mem. + Geol. Surv. New South Wales, Pal. No. 6, 1898, pp. 20-29. Dun, + W. S. Rec. Geol. Surv. New South Wales, vol. VII. pt. 4, 1904, + pp. 318-325 (Silurian to Carboniferous). Ibid., vol. VIII. pt. + 3, 1907, pp. 265-269. Chapman, F. Proc. R. Soc. Vict., vol. XVI. + (N.S.), pt. 1, 1903, pp. 64-79. Ibid., vol. XXI. (N.S.), pt. 1, + 1908, pp. 222, 223. Ibid., vol. XXVI. (N.S.) pt. 1. 1913, pp. + 99-113. + + Devonian.--McCoy, F. Prod. Pal. Vict., Dec. IV., 1876, pp. 16-18. + Foord, A. H. Geol. Mag., Dec. III. vol. VII. 1890, pp. 100-102. + Etheridge, R. jnr. Geol. and Pal. Queensland, 1892, pp. 64-68. + De Koninck, L. G. Mem. Geol. Surv. New South Wales, Pal., No. 6, + 1898, pp. 64-85. Chapman, F. Proc. R. Soc. Vict., vol. XVIII. + (N.S.), pt. 1, 1905, pp. 16-19. + + Carboniferous.--Etheridge, R. jnr. Rec. Austr. Mus., vol. IV. No. 3, + 1901, pp. 119, 120. Idem, Geol. Surv. W. Austr., Bull. No. 10, + 1903, pp. 12-23. Dun, W. S. Rec. Geol. Surv. New South Wales, + vol. VII., pt. 2, 1902, pp. 72-88 and 91-93. + + Carbopermian.--Sowerby, G. B., in Strzelecki's Phys. Descr. of New + South Wales, etc., 1845, pp. 275-285. McCoy, F. Ann. Mag. Nat. + Hist., vol. XX. 1847, pp. 231-236. Foord, A. H. Geol. Mag. Dec. + III. vol. VII. 1890, pp. 105 and 145-154. Etheridge, R. jnr. + Geol. and Pal. Queensland, 1892, pp. 225-264. De Koninck, L. G. + Mem. Geol. Surv. New South Wales, Pal., No. 6, 1898, pp. 140-203. + Dun, W. S. Rec. Geol. Surv. New South Wales, vol. VIII. pt. 4, + 1909, pp. 293-304. + + Lower Cretaceous.--Moore, C. Quart. Journ. Geol. Soc., vol. XXVI. + 1870, pp. 243-245. Etheridge, R. jnr. Mem. R. Soc. S. Austr., + vol. II. pt. 1, 1902, pp. 8, 9. + + Upper Cretaceous.--Etheridge, R. jnr. Geol. and Pal. Queensland, 1892, + p. 560. + + Cainozoic.--McCoy, F. Prod. Pal. Vict., Dec. V. 1877, pp. 11-13. Tate, + R. Trans. R. Soc. S. Austr., vol. III. 1880, pp. 140-170. Idem, + ibid., vol. XXIII. 1899, pp. 250-259. Hutton, F. W. Trans. N.Z. + Inst., vol. XXXVII. 1905, pp. 474-481 (Revn. Tert. Brach.). + + + + +CHAPTER X. + +FOSSIL SHELL-FISH (MOLLUSCA). + + +=Molluscan Characters.--= + +The phylum or sub-kingdom Mollusca is a group of soft-bodied animals +(mollis, soft), which, although having no external skeleton, usually +possess the protective covering of a shell. This shell is secreted +from the outer skin or mantle, and is composed of carbonate of lime +(calcareous) with a varying proportion of organic material. + + +=Hard Parts.--= + +Fossil molluscan remains consist practically of the shells, but the +calcareous apertural lid (operculum) of some kinds is often preserved, as +in _Turbo_ and _Hyolithes_; or the horny lids of others, as _Bithynia_ +of the European Pleistocene "brick earths." The cuttle-fishes have hard, +horny beaks and internal bones, and the latter are frequently found fossil +in Australia. + + +=Characters of Pelecypoda.--= + +The class for first consideration is the important one of the Bivalved +Mollusca, the _LAMELLIBRANCHIATA_ ("plate-gills") or _PELECYPODA_ +("hatchet foot"). The shells are double, hinged dorsally and placed on +either side of the animal, that is, they are left and right. The height is +measured on a vertical line drawn from the beaks or umbones to the ventral +margin. The length is the greatest distance between the margins parallel +with a line drawn through the mouth and posterior adductor impression. The +thickness is measured by a line at right angles to the line of height. The +shell being placed mouth forward, the valves are thus left and right. The +anterior is usually shorter, excepting in some cases, as in _Donax_ and +_Nucula_. + + +=Hinge Structure.--= + +In the absence of the animal, the character of the hinge-structure is very +important. Some are without teeth (edentulous). The oldest forms have been +grouped as the "Palaeoconcha," and it has been shown that here, although +well-developed teeth were absent, the radial ribs of the surface and +ventral areas were carried over to the dorsal margin and became a fixed +character in the form of crenulations or primitive teeth. + +The taxodont type of hinge teeth shows alternating teeth and sockets, as +in _Nucula_. + +The schizodont type is seen in the heavy, variable teeth of _Trigonia_ and +_Schizodus_. + +The isodont type of hingement is a modification of the taxodont, +represented by two ridges originally divergent below the beak, and +forming an interlocking series of two pairs of teeth and sockets as in +_Spondylus_; or where the primitive hinge disappears as in _Pecten_, the +divergent ridge-teeth (crura) may only partially develop. + +The dysodonts have a feeble hinge-structure derived from the external +sculpture impinging on the hinge-line, as in _Crenella_. + +The pantodonta are an ancient palaeozoic group which seems allied to the +modern teleodont or long toothed shells, but the laterals may exceed a +pair in a single group, as in _Allodesma_. + +The diogenodonta have lateral and cardinal teeth upon a hinge-plate, but +never more than two laterals and three cardinals in any one group, as in +_Crassatellites_. + +The cyclodonta have extremely arched teeth, which curve out from under the +beaks, as in _Cardium_. + +[Illustration: =Fig. 90--LOWER PALAEOZOIC BIVALVES.= + + A--Ambonychia macroptera, Tate. Cambrian. S. Australia + + B--Grammysia cuneiformis, Eth. fil. Silurian. Victoria + + C--Panenka gippslandica, McCoy sp. Silurian. Victoria + + D--Nucula melbournensis, Chapm. Silurian. Victoria + + E--Nuculites maccoyianus. Chapm. Silurian. Victoria + + F--Palaeoneilo victoriae, Chapm. Silurian. Victoria + +] + +The teleodonts include the more highly developed types of hinge, with +attenuated teeth and sockets. Common shells of our coast, and from +Cainozoic beds, belonging to this group are _Venus_, _Mactra_ and +_Meretrix_. + +The asthenodonta are boring and burrowing molluscs that have lost the +hinge dentition from disuse as _Corbula_ and _Pholas_. + + +=Cambrian Bivalve.--= + +The earliest example of a bivalved shell in Australian rocks is +_Ambonychia macroptera_ (Fig. 90 A), which occurs in the Cambrian +Limestone of Curramulka, S. Australia. It is quite a small form, being +less than a quarter of an inch in length. + + +=Ordovician Bivalve.--= + +In the basal Ordovician mudstone of Heathcote, Victoria, there is +a bivalve which in some respects resembles a _Modiolopsis_ (?_M. +knowsleyensis_), but the exact relationship is still doubtful. + + +=Silurian Bivalves.--= + +The Silurian sandstones, mudstones, slates and limestones of Australia +and New Zealand, unlike the older rocks just mentioned, contain a +rich assemblage of bivalve fossils. In Victoria the lower division or +Melbournian stage contains the following principal genera:--_Orthonota_, +_Grammysia_, _Leptodomus_, _Edmondia_, _Cardiola_, _Ctenodonta_, +_Nuculites_, _Nucula_, _Palaeoneilo_, _Conocardium_, _Modiolopsis_ and +_Paracyclas_. The upper division or Yeringian stage contains other +species of similar genera to those in the Melbournian, as _Grammysia_, +_Palaeoneilo_ and _Conocardium_; whilst _Panenka_, _Mytilarca_, +_Sphenotus_, _Actinodesma_, _Lunulicardium_, _Actinopteria_ and +Cypricardinia are, so far as known, peculiar to this and a still higher +stage. _Cardiola_ is a widely distributed genus, occurring as well in +Tasmania; whilst in Europe it is found both in Bohemia and Great Britain. +Its time-range in the northern hemisphere is very extensive, being found +in beds ranging from Upper Ordovician to Devonian. _Actinopteria_ is found +also in New South Wales and New Zealand, and _Pterinea_ and _Actinodesma_ +in New South Wales. + +The molluscs with a taxodont hinge-line (beset with numerous little +teeth and sockets) are quite plentiful in the Australian Silurian; such +as _Nucula_, a form common around Melbourne (_N. melbournensis_ (Fig. +90 D)); _Nuculites_, which has an internal radial buttress or clavicle +separating the anterior muscle-scar from the shell-cavity, and which is +found likewise in the Melbourne shales (_N. maccoyianus_ (Fig. 90 E)); +_Ctenodonta_, represented in both the Melbournian and Yeringian stages +(_C. portlocki_); and _Palaeoneilo_, a handsome, subrostrate generic +type with concentric lamellae or striae, commonest in the Melbournian, +but occasionally found in the younger stage (_P. victoriae_ Fig. 90 F, +Melbournian;--_P. raricostae_, Yeringian). _Conocardium_ is represented +by two species in Victoria (_C. bellulum_ and _C. costatum_); whilst in +New South Wales _C. davidis_ is found at Oakey Creek. In New Zealand +_Actinopteria_ and _Pterinea_ occur in the Wangapeka series (Silurian). + + +=Devonian Bivalves.--= + +The compact limestone and some shales of Middle Devonian age in the N.E. +Gippsland area in Victoria, contain several as yet undescribed species +belonging to the genera _Sphenotus_, _Actinodesma_ and _Paracyclas_. + +[Illustration: =Fig. 91--PALAEOZOIC BIVALVES.= + + A--Mytilarca acutirostris, Chapm. Silurian. Victoria + + B--Modiolopsis melbournensis, Chapm. Silurian. Victoria + + C--Goniophora australis, Chapm. Silurian. Victoria + + D--Paracyclas siluricus, Chapm. Silurian. Victoria + + E--Actinopteria australis, Dun. Devonian. New South Wales + + F--Lyriopecten gracilis, Dun. Devonian. New South Wales + +] + +The genera _Paracyclas_, _Aviculopecten_ and _Pterinea_ have been recorded +from New South Wales, chiefly from the Yass district. The derived +boulders found in the Upper Cretaceous beds forming the opal-fields at +White Cliffs, New South Wales, have been determined as of Devonian age. +They contain, amongst other genera, examples of _Actinopteria_ (_A. +australis_), _Lyriopecten_ (_L. gracilis_) (Fig. 91 F), and _Leptodesma_ +(_L. inflatum_ and _L. obesum_). + +[Illustration: =Fig. 92--CARBOPERMIAN BIVALVES.= + + A--Stutchburia farleyensis, Eth. fil. Carbopermian. N.S. Wales + + B--Deltopecten limaeformis, Morris sp. Carbopermian. N.S. Wales + + C--Aviculopecten sprenti, Johnston. Carbopermian. N.S. Wales + + D--Chaenomya etheridgei, de Kon. Carbopermian. N.S. Wales + + E--Pachydomus globosus J. de C. Sow. Carbopermian. N.S. Wales + +] + + +=Carbopermian Bivalves.--= + +One of the most prolific palaeozoic series for bivalved mollusca is the +Carbopermian. To select from the numerous genera and species we may +mention _Stutchburia farleyensis_ (Fig. 92 A) and _Edmondia nobilissima_ +from Farley, New South Wales; and _Deltopecten limaeformis_ (Fig. 92 +B), found in the Lower Marine Series at Ravensfield, New South Wales, +and in the Upper Marine Series at Burragorang and Pokolbin in the same +State, in Queensland at the Mount Britton Gold-field, and in Maria Id., +Tasmania. _Deltopecten fittoni_ occurs in both series in New South Wales, +and in the Upper Marine Series associated with "Tasmanite shale" in +Tasmania. _Aviculopecten squamuliferus_ is a handsome species found alike +in Tasmania and New South Wales; whilst _A. tenuicollis_ is common to +W. Australia and New South Wales. Other characteristic bivalves of the +Carbopermian of New South Wales are _Chaenomya etheridgei_ (Fig. 92 D) +and _Pachydomus globosus_ (Fig. 92 E). The gigantic _Eurydesma cordatum_ +is especially characteristic of the New South Wales Lower Marine Series, +and is also found in Tasmania. All three species are found in Queensland. + + +=Triassic Bivalves.--= + +The Triassic rocks of New South Wales were accumulated under either +terrestrial, lacustrine, or brackish (estuarine) conditions. Hence the +only bivalved mollusca found are referred to the freshwater genera _Unio_ +(_U. dunstani_) and _Unionella_ (_U. bowralensis_ and _U. carnei_ (Fig. 93 +A)). The latter genus differs from Unio in the structure of the adductor +muscle-impressions. + +[Illustration: =Fig. 93--LOWER MESOZOIC BIVALVES.= + + A--Unionella carnei, Eth. fil. Triassic. New South Wales + + B--Mytilus problematicus, Zittel. Triassic. New Zealand + + C--Monotis salinaria, Zittel. Triassic. New Zealand + + D--Trigonia moorei, Lycett. Jurassic. W. Australia + + E--Astarte cliftoni, Moore. Jurassic. W. Australia + +] + +The Queensland Trias (Burrum Formation) contains a solitary species of +bivalved mollusca, _Corbicula burrumensis_. This genus is generally found +associated with freshwater or brackish conditions. + +In New Zealand marine Triassic beds occur, containing, amongst other +genera, a species of _Leda_. In the succeeding Wairoa Series the +interesting fossil, _Daonella lommeli_ occurs. This shell is typical of +the Norian (Upper Trias) of the Southern Tyrol. Above the _Daonella_ bed +occurs the _Trigonia_ bed, with that genus and _Edmondia_. In the next +younger stage, the Otapiri Series, near Nelson, there are fine-grained +sandstones packed full of the remains of _Mytilus problematicus_ (Fig. 93 +B) and _Monotis salinaria_ (Fig. 93 C), the latter also a Norian fossil. + + +=Jurassic Bivalves.--= + +Jurassic bivalved molluscs are plentiful in the W. Australian limestones, +as at Greenough River. Amongst others may be mentioned _Cucullaea +semistriata_, _Ostrea_, _Gryphaea_, _Trigonia moorei_ (Fig. 93 D), _Pecten +cinctus_, _Ctenostreon pectiniforme_ and _Astarte cliftoni_ (Fig. 93 E). +Several of the species found are identical with European Jurassic fossils. + +Jurassic strata in Victoria, being of a freshwater and lacustrine nature, +yield only species of _Unio_, as _U. dacombei_, and _U. stirlingi_. + +The Jurassic beds of S. Australia contain a species of _Unio_ named _U. +eyrensis_. In the same strata which contains this shell, plant remains +are found, as _Cladophlebis_ and _Thinnfeldia_, two well-known types of +Jurassic ferns. + + +=Lower Cretaceous Bivalves.--= + +In Queensland the Lower Cretaceous limestones and marls contain a large +assemblage of bivalves, the more important of which are _Nucula truncata_ +(Fig. 94 A), _Maccoyella reflecta_ (Fig. 94 B), _M. barkleyi_, _Pecten +socialis_ and _Fissilunula clarkei_ (Fig. 94 C), from Wollumbilla; and +_Inoceramus pernoides_, _I. carsoni_ and _Aucella hughendenensis_ from the +Flinders River (the latter also from New South Wales). + +In the Lake Eyre District of S. Australia we find _Maccoyella barkleyi_, +which also occurs in Queensland and New South Wales (at White Cliffs), +_Trigonia cinctuta_, _Mytilus rugocostatus_ and _Modiola eyrensis_. The +handsome bivalve, _Pleuromya plana_ occurs near Broome in W. Australia. + +[Illustration: =Fig. 94--CRETACEOUS BIVALVES.= + + A--Nucula truncata, Moore. L. Cretaceous. South Australia + + B--Maccoyella reflecta, Moore sp. Up. and L. Cretaceous. Q'land. + + C--Fissilunula clarkei, Moore sp. Up. and L. Cretaceous. Q'land. + + D--Inoceramus carsoni, McCoy. L. Cretaceous. Queensland + + E--Cyrenopsis opallites, Eth. fil. Up. Cretaceous. New South Wales + + F--Conchothyra parasitica, Hutton. Cretaceous. New Zealand + +] + + +=Upper Cretaceous Bivalves.--= + +The Upper Cretaceous or Desert Sandstone at Maryborough, Queensland, +has yielded amongst others, the following shells:--(_Nucula gigantea_, +_Maccoyella reflecta_ also found in the Lower Cretaceous of Queensland, +New South Wales and S. Australia), and _Fissilunula clarkei_ (also found +in the L. Cretaceous of New South Wales, Queensland and S. Australia). +Some of these beds, however, which were hitherto believed to belong +to the Upper and Lower Series respectively may yet prove to be on one +horizon--the Lower Cretaceous. _Cyrenopsis opallites_ (Fig. 94 E) of White +Cliffs, New South Wales, appears to be a truly restricted Upper Cretaceous +species. + +The Cretaceous of New Zealand (Amuri System) contains _Trigonia sulcata_, +_Inoceramus_ sp. and the curious, contorted shell, _Conchothyra +parasitica_ (Fig. 94 F) which is related to _Pugnellus_, a form usually +considered as a sub-genus of _Strombus_. + +From Papua an _Inoceramus_ has been recorded from probable Cretaceous beds. + + +=Cainozoic Bivalves.--= + +In Victoria, South Australia, and the N.W. of Tasmania, as well as in +New Zealand, Cainozoic marine beds are well developed, and contain an +extensive bivalved molluscan fauna. Of these fossils only a few common and +striking examples can here be noticed, on account of the limits of the +present work. + +The commonest genera are:--_Ostrea_, _Placunanomia_, _Dimya_, _Spondylus_, +_Lima_, _Pecten_, _Arca_, _Barbatia_, _Plagiarca_, _Cucullaea_, +_Glycimeris_, _Limopsis_, _Nucula_, _Leda_, _Trigonia_, _Cardita_, _Cuna_, +_Crassatellites_, _Cardium_, _Protocardium_, _Chama_, _Meretrix_, _Venus_ +(_Chione_), _Dosinea_, _Gari_, _Mactra_, _Corbula_, _Lucina_, _Tellina_, +_Semele_ and _Myodora_. + +[Illustration: =Fig. 95--CAINOZOIC BIVALVES.= + + A--Dimya dissimilis, Tate. Balcombian. Victoria + + B--Spondylus pseudoradula, McCoy. Balcombian. Victoria + + C--Pecten polymorphoides, Zittel. Janjukian. South Australia + + D--Leda vagans, Tate. Janjukian. South Australia + + E--Modiola praerupta, Pritchard. Balcombian. Victoria + +] + + +=Persistent Species.--= + +To mention a few species of persistent range, from Balcombian to +Kalimnan, we may cite the following from the Cainozoic of southern +Australia:--_Dimya dissimilis_ (Fig. 95 A), _Spondylus pseudoradula_ (Fig. +95 B), _Lima (Limatula) jeffreysiana_, _Pecten polymorphoides_ (found also +in the Oamaru Series, New Zealand) (Fig. 95 C), _Amusium zitteli_ (found +also in both the Waimangaroa and Oamaru Series of New Zealand), _Barbatia +celleporacea_, _Cucullaea corioensis_, _Limopsis maccoyi_, _Nucula +tenisoni_, _Leda vagans_ (Fig. 95 D), _Corbula ephamilla_ and _Myodora +tenuilirata_. + + +=Balcombian Bivalves.--= + +On the other hand, many species have a restricted range, and these are +invaluable for purposes of stratigraphical correlation. For example, +in the Balcombian we have _Modiola praerupta_ (Fig. 95 E), _Modiolaria +balcombei_, _Cuna regularis_, _Cardium cuculloides_, _Cryptodon +mactraeformis_, _Verticordia pectinata_ and _V. excavata_. + +[Illustration: =Fig. 96--CAINOZOIC BIVALVES.= + + A--Modiola pueblensis, Pritchard. Janjukian. Victoria + + B--Cardita tasmanica, Tate. Janjukian. Tasmania + + C--Lucina planatella, Tate. Janjukian. Tasmania + + D--Ostrea manubriata, Tate. Kalimnan. Victoria + + E--Limopsis beaumariensis, Chap. Kalimnan. Victoria + + F--Venus (Chione) subroborata, Tate sp. Kalimnan. Victoria + +] + + +=Janjukian Bivalves.--= + +In the Janjukian Series restricted forms of bivalves are exceptionally +numerous, amongst them being:--_Dimya sigillata_, _Plicatula ramulosa_, +_Lima polynema_, _Pecten praecursor_, _P. eyrei_, _P. gambierensis_, +_Pinna cordata_, _Modiola pueblensis_ (Fig. 96 A), _Arca dissimilis_, +_Limopsis multiradiata_, _L. insolita_, _Leda leptorhyncha_, _L. +crebrecostata_, _Cardita maudensis_, _C. tasmanica_ (Fig. 96 B), _Cuna +radiata_, _Lepton crassum_, _Cardium pseudomagnum_, _Venus (Chione) +multitaeniata_, _Solenocurtus legrandi_, _Lucina planatella_ (Fig. 96 C), +_Tellina porrecta_ and _Myodora lamellata_. + +In Papua a _Pecten_ (_P. novaeguineae_) has been recorded from the ? Lower +Pliocene of Yule Island. + + +=Kalimnan Bivalves.--= + +The Kalimnan beds contain the following restricted or upward ranging +species:--_Ostrea arenicola_, _O. manubriata_ (Fig. 96 D), _Pecten +antiaustralis_ (also in the Werrikooian Series), _Perna percrassa_, +_Mytilus hamiltonensis_, _Glycimeris halli_, _Limopsis beaumariensis_ +(also Werrikooian) (Fig. 96 E), _Leda crassa_ (also living), _Trigonia +howitti_, _Cardita solida_, _C. calva_ (also living), _Erycina micans_, +_Meretrix paucirugata_, _Sunetta gibberula_, _Venus (Chione) subroborata_ +(Fig. 96 F), _Donax depressa_, _Corbula scaphoides_ (also living), _Barnea +tiara_, _Lucina affinis_, _Tellina albinelloides_ and _Myodora corrugata_. + + +=Werrikooian Bivalves.--= + +The next stage, the Werrikooian (Upper Pliocene), contains a large +percentage of living species, as _Ostrea angasi_, _Placunanomia ione_ +(ranging down into Janjukian), _Glycimeris radians_, _Leda crassa_ (also +a common Kalimnan fossil), various species of _Venus (Chione)_, as _V. +strigosa_ and _V. placida_, and _Barnea australasiae_. + + +=Pleistocene Bivalves.--= + +The bivalved shells of the Pleistocene are similar to those now found +living round the Australian coast, as _Pecten asperrimus_, _Mytilus +latus_, _Leda crassa_, _Soletellina biradiata_ and _Spisula parva_. + +Pleistocene shells of bivalved genera occur in the coastal hills of Papua, +including the following:--_Cultellus_, _Corbula_, _Mactra_, _Tellina_, +_Venus (Chione)_, _Dione_, _Dosinea_, _Leda_ and _Arca_. + +The _SCAPHOPODS_ ("digger foot") or the "Elephant-tusk shells" are +adapted, by their well-developed foot, to burrow into the mud and sand. + +[Illustration: =Fig. 97--FOSSIL SCAPHOPODS and CHITONS.= + + A--Dentalium huttoni, Bather. Jurassic. New Zealand + + B--Dentalium mantelli, Zittel. Cainozoic. Victoria + + C--Chelodes calceoloides, Eth. fil. Silurian. New South Wales + + D--Ischnochiton granulosus, Ashby and Torr sp. Cainozoic (Balc). + Victoria + + E--Cryptoplax pritchardi, Hall. Cainozoic (Kalimnan). Victoria + +] + + +=Devonian Scaphopods.--= + +This group of mollusca makes its first appearance in Australasian +sediments in the Middle Devonian (Murrumbidgee beds) of New South Wales, +represented by _Dentalium tenuissimum_. + + +=Jurassic Scaphopods.--= + +In the Jurassic strata of the Mataura Series of New Zealand, _Dentalium +huttoni_ (Fig. 97 A) occurs at the Kowhai River and Wilberforce. + + +=Cretaceous Scaphopods.--= + +_Dentalium wollumbillensis_ occurs in the drab and dark-coloured +limestones of the Lower Cretaceous of the Lake Eyre Basin in S. Australia, +and the same species is also found in the Lower Cretaceous (Rolling Downs +Formation) of Wollumbilla, Queensland. + + +=Cainozoic Scaphopods.--= + +The Cainozoic beds both of New Zealand and southern Australia yield +many species of _Dentalium_, the commonest and most widely distributed +being the longitudinally ribbed _D. mantelli_ (Fig. 97 B), which ranges +from the Balcombian to the Werrikooian stages in Australia, and is also +typical of the Oamaru Series in New Zealand, where it is accompanied by +the ponderous species, _D. giganteum_, which attained a length of over six +inches. Another form common in our Cainozoics is the smooth-shelled _D. +subfissura_; this also has a wide range, namely Balcombian to Kalimnan. + + +=Palaeozoic Chitons.--= + +The _POLYPLACOPHORA_ or Chitons ("Mail-shells"), first appeared in the +Ordovician. In Australia _Chelodes calceoloides_ (Fig. 97 C) is found in +the Silurian of Derrengullen Creek, Yass, New South Wales; and another +species of the genus is found in beds of the same age at Lilydale, +Victoria. Between that period and the Cainozoic or Tertiary there is a gap +in their history in Australia. + + +=Cainozoic Chitons.--= + +_Ischnochiton granulosus_ (Fig. 97 D) is a Balcombian species of the +modern type of "mail-shell," occurring not infrequently in the clays of +Balcombe's Bay, Port Phillip, Victoria. _Cryptoplax pritchardi_ (Fig. 97 +E) is a curious form belonging to the attenuated, worm-like group of the +Cryptoplacidae, until lately unknown in the fossil state; it is found in +the Kalimnan Series near Hamilton, Victoria. Several other genera of the +chitons are found fossil in the Australian Cainozoics which still live +on our coasts, as _Lorica_, _Plaxiphora_ and _Chiton_. The first-named +genus is represented fossil by _Lorica duniana_ from the _Turritella_ bed +(Janjukian) of Table Cape, Tasmania. + + +=Characters of Gasteropoda.--= + +The _GASTEROPODA_ ("belly-foot") or univalve shells possess a muscular +foot placed beneath the stomach and viscera. In the Heteropoda this foot +is modified as a vertical fin, and in the Pteropoda as two wing-like +swimming membranes close to the head. The mantle lobe is elevated along +the back like a hood, and its surfaces and edges secrete the shell +which contains the animal. The shell is typically a cone (example, +_Patella_ or Limpet) which is often spirally coiled either in a plane +(ex. _Planorbis_), conically turbinoid (ex. _Trochus_), or turreted (ex. +_Turritella_). The body and shell are attached by muscles, the spiral +forms being attached to the columella or axial pillar, and the bowl-shaped +forms to the inner surface of the shell. + +Gasteropod shells are normally right-handed (dextral), but a few genera +as _Clausilia_, _Bulinus_ and _Physa_, are left-handed (sinistral). The +height or length of the shell is measured from the apex to the lower +margin of the mouth. In coiled shells we may regard them as a more or less +elongated cone wound round a central pillar, the columella, or around a +central tube. A turn or coil of the shell is a whorl, and together, with +the exception of the last, form the spire. The line between two adjacent +whorls is the suture. When the columella is solid the shell is said to +be imperforate, and when a central tube is left by the imperfect fusion +of the whorls, it is perforate. The opening of the tubular columella is +termed the umbilicus, and this is sometimes contracted by the encroachment +of shell matter termed the callus. The aperture is entire when the rim +is uninterrupted; and channelled when there is a basal notch, where the +siphon which conducts water to the gills is lodged. + +As a rule the large heavy gasteropods inhabit shallow water. The following +living genera are characteristic of rocky shore-lines; _Risella_, +_Buccinum_, _Purpura_ and _Patella_. Genera typical of sandy shores are +_Nassa_, _Natica_, _Cypraea_, _Turritella_ and _Scala_. + + +=Cambrian Gasteropods.--= + +From the Cambrian of South Australia Prof. Tate described some minute +Gasteropods which he referred to the genera _Stenotheca_ (_S. rugosa_, +var. _paupera_), _Ophileta (O. subangulata)_ (Fig. 98 A), and _Platyceras +(P. etheridgei)_. In these beds at Curramulka the following Pteropods were +found by the same authority, viz., _Salterella planoconvexa_, _Hyolithes +communis_ (Fig. 98 C) and _H. conularioides_. + +The Cambrian Limestone of the Kimberley District, W. Australia, contains +the characteristic Pteropod _Salterella hardmani_ (Fig. 98 B). The shell +is a conical tube, straight or slightly curved, and measuring scarcely an +inch in length. + +[Illustration: =Fig. 98--LOWER PALAEOZOIC GASTEROPODA.= + + A--Ophileta subangulata, Tate. Cambrian. South Australia + + B--Salterella hardmani, Foord. Cambrian. West Australia + + C--Hyolithes communis, Billings. Cambrian. South Australia + + D--Scenella tenuistriata, Chapm. Cambrian. Victoria + + E--Raphistoma browni, Eth. fil. Ordovician. South Australia + + F--Helicotoma johnstoni, Eth. fil. Silurian. Tasmania + +] + +The Upper Cambrian of the Mersey River District in Tasmania has afforded +some doubtful examples of the genus _Ophileta_. + +In the Upper Cambrian Limestones of the Dolodrook Valley, near Mt. +Wellington, Victoria, a minute limpet shaped Gasteropod occurs, named +_Scenella tenuistriata_ (Fig. 98 D). + + +=Ordovician Gasteropods.--= + +Ordovician limestones with fossil shells occur in the Leigh's Creek +District in South Australia, and also at Tempe Downs and Petermann and +Laurie's Creeks, W. of Alice Springs. The euomphaloid shell _Ophileta +gilesi_ was described from Laurie's Creek, and _Eunema larapinta_ from +the Tempe Downs. A pleurotomarid, _Raphistoma browni_ (Fig. 98) occurs +near Leigh's Creek, and at Laurie's and Petermann Creeks. A Pteropod, +_Hyolithes leptus_, has been described from the Lower Ordovician of Coole +Barghurk Creek, near Meredith, Victoria. + + +=Silurian Gasteropods.--= + +The Silurian Gasteropods are fairly well represented, especially in +the upper stage, and are widely distributed throughout the Australian +fossiliferous localities. Moreover, some of the species are identical +with those found as far off as North America and Europe. In Victoria +the shales and sandstones of the lower stage (Melbournian) contain the +genera _Bellerophon_, _Cyrtolites_ and _Loxonema_. The Pteropoda include +_Tentaculites_, _Coleolus_, _Hyolithes_ and _Conularia_ (_C. sowerbii_ +(Fig. 99 F), a species also found in Great Britain). The Victorian +limestones and mudstones of the upper stage (Yeringian) are somewhat rich +in Gasteropods, such genera occurring as _Pleurotomaria_, _Phanerotrema_ +(with cancellated shell and large slit-band), _Murchisonia_, _Gyrodoma_, +_Bellerophon_, _Trematonotus_ (a spiral shell with a large trumpet-shaped +mouth and a dorsal row of perforations in place of a slit-band), +_Euomphalus_, _Cyclonema_, _Trochus (Scalaetrochus)_, _Niso (Vetotuba)_, +_Loxonema_, _Platyceras_ and _Capulus_. The section Pteropoda contains +_Tentaculites_, _Hyolithes_ and _Conularia_. + +[Illustration: =Fig. 99--SILURIAN GASTEROPODA.= + + A--Hyolithes spryi, Chapm. Silurian (Melb.) Victoria + + B--Gyrodoma etheridgei, Cressw. sp. Silurian (Yeringian). Vict. + + C--Bellerophon cresswelli. Eth. fil. Silurian (Yeringian). Victoria + + D--Euomphalus northi, Eth. fil. sp. Silurian (Yeringian). Victoria + + E--Trochonema montgomerii. Eth. fil. sp. Silurian. Tasmania + + F--Conularia sowerbii, Defr. Silurian (Yeringian). Victoria + +] + +In the Silurian of New South Wales the chief Gasteropod genera are +_Bellerophon (B. jukesi)_, _Euomphalus_, _Omphalotrochus_, and _Conularia +(C. sowerbii.)_. + +In Tasmania are found _Raphistoma_, _Murchisonia_, _Bellerophon_, +_Helicotoma_, _Trochonema_ and _Tentaculites_. + + +=Devonian Gasteropods.--= + +The derived boulders of the White Cliffs opal field have been referred to +the Devonian system, but of this there is some doubt, as the Gasteropods +noted from these boulders closely resemble those of the Silurian fauna: +they are _Murchisonia Euomphalus_ (_E. culleni_), and _Loxonema_. The +genus _Murchisonia_ has also been recorded from the Baton River, New +Zealand (Wangepeka Series) by MacKay. + +The Middle Devonian Gasteropod fauna in Victoria, as found in the Buchan +and Bindi Limestones, comprises _Murchisonia_, _Trochus_, and _Platyceras_. + +[Illustration: =Fig. 100--UPPER PALAEOZOIC GASTEROPODA.= + + A--Gosseletina australis, Eth. fil. sp. Carboniferous. N.S. Wales + + B--Yvania konincki, Eth. fil. Carboniferous. N.S. Wales + + C--Loxonema babbindoonensis, Eth. fil. Carboniferous. N.S. Wales + + D--Pleurotomaria (Ptychomphalina) morrisiana, McCoy. Carbopermian. + N.S. Wales + + E--Platyschisma oculum, Sow. sp. Carbopermian. N.S. Wales + + F--Murchisonia carinata, Eth. Carbopermian. Queensland + +] + +In New South Wales the best known genera are _Pleurotomaria_, +_Murchisonia_, _Bellerophon_, _Euomphalus_ and _Loxonema_. The two latter +genera have also been obtained at Barker Gorge, Western Australia. + + +=Carboniferous Gasteropods.--= + +Carboniferous Gasteropoda have been found in New South Wales, belonging +to the genera _Gosseletina_ (_G. australis_) (Fig. 100 A) and _Yvania_ +(_Y. konincki_) (Fig. 100 B), both of which have their countertypes in the +Carboniferous of Belgium. _Y. konincki_ is also found in the Carbopermian +(Gympie beds) of Rockhampton, Queensland, while _Y. levellii_ is found in +the Carbopermian of Western Australia. + + +=Carbopermian Gasteropods.--= + +The Carbopermian gasteropods of New South Wales are _Pleurotomaria_ +(_Mourlonia_), _Keeneia platyschismoides_, _Murchisonia_, _Euomphalus_, +_Platyschisma_ (_P. oculum_) (Fig. 100 E), _Loxonema_ and _Macrocheilus_. +Examples of the genus _Conularia_ are sometimes found, probably attaining +a length, when complete, of 40 centimetres. + +In Tasmania we find _Conularia tasmanica_, a handsome Pteropod, also +of large dimensions. _Platyschisma_, _Pleurotomaria_ (_Mourlonia_), +_Bellerophon_ and _Porcellia_ are amongst the Carbopermian Gasteropods of +Queensland. + +In Western Australia _Pleurotomaria_ (_Mourlonia_), _Bellerophon_, +_Euomphalus_, _Euphemus_, _Platyceras_, and _Loxonema_ occur in the +Carbopermian. + + +=Jurassic Gasteropods.--= + +Jurassic gasteropods are found sparingly in the limestone of the +Geraldton District and other localities in Western Australia. The more +important of these are _Pleurotomaria_ (_P. greenoughiensis_), _Turbo_ +(_T. australis_) (Fig. 101 A) and _Rissoina_ (_R. australis_) (Fig. 101 B). + +[Illustration: =Fig. 101--MESOZOIC GASTEROPODA.= + + A--Turbo australis, Moore. Jurassic. West Australia + + B--Rissoina australis, Moore. Jurassic. West Australia + + C--Natica ornatissima, Moore. Cretaceous. Queensland + + D--Pseudamaura variabilis, Moore sp. Cretaceous. Queensland + + E--Rostellaria waiparensis, Hector.--Cretaceous. New Zealand + +] + + +=Cretaceous Gasteropods.--= + +The Queensland gasteropod fauna comprises _Cinulia_ a typical Cretaceous +genus, _Actaeon_ and _Natica_. These occur in the Lower Cretaceous or +Rolling Downs Formation. _Cinulia_ is also found in South Australia at +Lake Eyre with _Natica_ (_N. ornatissima_) (Fig. 101 C). _Pseudamaura +variabilis_ (Fig. 101 D) is found in New South Wales, Queensland and South +Australia; whilst _Anchura wilkinsoni_ occurs in Queensland and South +Australia. + +In New Zealand the Waipara Greensands (Cretaceous) contain a species of +_Rostellaria_ (_R. waiparensis_) (Fig. 101 E). + + +=Cainozoic Gasteropods.--= + +Cainozoic Gasteropods are exceedingly abundant in beds of that system +in Australasia. The Cainozoic marine fauna in Australia is practically +restricted to the States of Victoria, South Australia, and Tasmania; +whilst New Zealand has many species in common with Australia. + + +=Genera.--= + +The commonest genera of the marine Cainozoic or Tertiary deposits +are:--_Haliotis_, _Fissurellidea_, _Emarginula_, _Subemarginula_, +_Astralium_, _Liotia_, _Gibbula_, _Eulima_, _Niso_, _Odostomia_, +_Scala_, _Solarium_, _Crepidula_, _Calyptraea_, _Natica_, _Rissoa_, +_Turritella_, _Siliquaria_, _Cerithium_, _Newtoniella_, _Tylospira_, +_Cypraea_, _Trivia_, _Morio_, _Semicassis_, _Lotorium_, _Murex_, _Typhis_, +_Columbella_, _Phos_, _Nassa_, _Siphonalia_, _Euthria_ (_Dennantia_), +_Fusus_, _Columbarium_, _Fasciolaria_, _Latirus_, _Marginella_, _Mitra_, +_Volutilithes_, _Voluta_, _Harpa_, _Ancilla_, _Cancellaria_, _Terebra_, +_Pleurotoma_, _Drillia_, _Conus_, _Bullinella_ and _Vaginella_. + + +=Persistent Species.--= + +Amongst the Cainozoic Gasteropoda of southern Australia which have a +persistent range through Balcombian to Kalimnan times, we find:--_Niso +psila_, _Crepidula unguiformis_ (also Werrikooian and Recent), _Natica +perspectiva_, _N. hamiltonensis_, _Turritella murrayana_, _Cerithium +apheles_, _Cypraea leptorhyncha_, _Lotorium gibbum_, _Volutilithes +antiscalaris_ (also in Werrikooian), _Marginella propinqua_, _Ancilla +pseudaustralis_, _Conus ligatus_ and _Bullinella exigua_. + + +=Balcombian Gasteropods.--= + +Species restricted to the Balcombian stage include _Scala dolicho_, +_Seguenzia radialis_, _Dissocheilus eburneus_, _Trivia erugata_, _Cypraea +ampullacea_ (Fig. 102 A), _C. gastroplax_, _Colubraria leptoskeles_, +_Murex didymus_ (Fig. 102 B), _Eburnopsis aulacoessa_ (Fig. 102 C), +_Fasciolaria concinna_, _Mitra uniplica_, _Harpa abbreviata_, _Ancilla +lanceolata_, _Cancellaria calvulata_ (Fig. 102 D), _Buchozia oblongula_, +_Pleurotoma optata_, _Terebra leptospira_ and _Vaginella eligmostoma_ +(Fig. 102 E), (also found at Gellibrand River). + +[Illustration: =Fig. 102--CAINOZOIC GASTEROPODA.= + + A--Cypraea ampullacea, Tate. Cainozoic (Balc.) Victoria + + B--Murex didymus, Tate. Cainozoic (Balc.) Victoria + + C--Eburnopsis aulacoessa, Tate. Cainozoic (Balc.) Victoria + + D--Cancellaria calvulata, Tate. Cainozoic (Balc.) Victoria + + E--Vaginella eligmostoma, Tate. Cainozoic (Balc.) Victoria + +] + +[Illustration: =Fig. 103--CAINOZOIC GASTEROPODA.= + + A--Eutrochus fontinalis, Pritchard. Cainozoic (Janjukian). Vict. + + B--Morio wilsoni, Tate. Cainozoic (Janjukian). Victoria + + C--Scala lampra, Tate sp. Cainozoic (Janjukian). South Australia + + D--Natica gibbosa, Hutton. Cainozoic (Janjukian). South Australia + + E--Volutilithes anticingulatus, McCoy sp. Cainozoic (Janjukian). + Victoria + + F--Struthiolaria sulcata, Hutton. Cainozoic (Awatere series). New + Zealand + +] + + +=Janjukian Gasteropods.--= + +Species of Gasteropods restricted to the Janjukian stage +include:--_Pleurotomaria tertiaria_, _Haliotis mooraboolensis_, _Liotia +lamellosa_, _Thalotia alternata_, _Eutrochus fontinalis_ (Fig. 103 A), +_Astralium hudsonianum_, _Turbo atkinsoni_, _Odostomia polita_, _Scala +lampra_ (Fig. 103 C), _Natica gibbosa_ (Fig. 103 D) (also found in the +Pareora Series of the Oamaru system and in the Wanganui beds of New +Zealand), _Calyptraea subtabulata_, _Turritella aldingae_, _Cerithiopsis +mulderi_, _Cerithium flemingtonense_, _Cypraea platyrhyncha_, _C. +consobrina_, _Morio wilsoni_ (Fig. 103 B), _Lotorium abbotti_, _Murex +otwayensis_, _Eburnopsis tesselatus_, _Tudicla costata_, _Latirus +semiundulatus_, _Fusus meredithae_, _Columbarium spiniferum_, _Voluta +pueblensis_, _V. heptagonalis_, _V. macroptera_ (also recorded from +Hall's Sound, Papua) (Fig. 103 E), _Volutilithes anticingulatus_ (also +from Papua), _Harpa clathrata_, _Bela woodsi_, _Bathytoma paracantha_ and +_Volvulella inflatior_. + +_Dolium costatum_, allied to the "Fig-Shell" has been noted from the +Cainozoic clays (? Lower Pliocene), Yule Island, Papua. + +[Illustration: =Fig. 104--LATE CAINOZOIC and PLEISTOCENE GASTEROPODA= + + A--Bankivia howitti, Pritchard. Cainozoic (Kal.) Victoria + + B--Eglisia triplicata, Tate sp. Cainozoic (Kal.) Victoria + + C--Voluta masoni, Tate. Cainozoic (Kal.) Victoria + + D--Ancilla papillata. Tate sp. Cainozoic (Kal.) Victoria + + E--Terebra geniculata, Tate. Cainozoic (Kal.) Victoria + + F--Helix simsoniana, Johnston. Pleistocene. Tasmania + +] + + +=Kalimnan Gasteropods.--= + +Species of Gasteropods restricted to the Kalimnan Stage, or only +passing upwards include:--_Bankivia howitti_ (Fig. 104 A), _Liopyrga +quadricingulata_, _Calyptraea corrugata_, _Natica subvarians_, +_Turritella pagodula_, _Eglisia triplicata_ (Fig. 104 B), _Tylospira +clathrata_, _Cypraea jonesiana_, _Lotorium ovoideum_, _Sistrum +subreticulatum_, _Voluta masoni_ (Fig. 104 C), _Ancilla papillata_ (Fig. +104 D), _Cancellaria wannonensis_, _Drillia wanganuiensis_ (also in the +Petane Series of New Zealand), _Terebra catenifera_, _T. geniculata_ (Fig. +104 E) and _Ringicula tatei_. + + +=New Zealand Cainozoic Gasteropods.--= + +Characteristic Gasteropoda of the Oamaru Series in New Zealand are +_Pleurotomaria tertiaria_ (also in the Australian Janjukian), _Scala +lyrata_, _Natica darwinii_, _Turritella cavershamensis_, _Ancilla hebera_ +(also in the Australian Balcombian and Janjukian) and _Pleurotoma +hamiltoni_. Gasteropods of the Awatere Series in New Zealand are _Natica +ovata_, _Struthiolaria sulcata_ (Fig. 103 F), and _Scaphella corrugata_ +(found also in the Oamaru Series). The Putiki beds of the Petane Series in +New Zealand contain _Trophon expansus_, _Pisania drewi_ and _Pleurotoma +wanganuiensis_. + + +=Werrikooian Gasteropods.--= + +The marine gasteropods of the Werrikooian of southern Australia, as found +at Limestone Creek, Glenelg River, Western Victoria, and the Moorabool +Viaduct near Geelong, are nearly all living at the present time, with +the exception of a few older Cainozoic species. Amongst these latter are +_Conus ralphi_, _Pleurotoma murndaliana_, _Volutilithes antiscalaris_ and +_Columbarium craspedotum_. + + +=Pleistocene Gasteropoda.--= + +The Pleistocene land mollusca, and especially the gasteropods of +Australia, present some striking points of interest, for whilst most of +the species are still living, some appear to be extinct. The travertine +deposits of Geilston, near Hobart, Tasmania contain _Helix geilstonensis_ +and _H. stanleyana_, the latter still living. The calcareous _Helix_ +sandstone of the islands in Bass Strait are largely composed of shells +of that genus and generally represent consolidated sand-dunes which have +undergone a certain amount of elevation. One of the prevalent species is +_Helix simsoniana_ (Fig. 104 F), a handsome keeled form, somewhat related +to the living _H. launcestonensis_. It is found in some abundance in the +Kent's Group and in the adjacent islands. + +The large ovoid land-shells, _Panda atomata_, although still existing, +are found associated with extinct marsupials, as _Thylacoleo_, in the +stalagmitic floor of the Buchan Caves, Gippsland. + +The _Diprotodon_-breccias of Queensland have afforded several species +of _Helix_ and other land-shells, as well as the brackish-water genus +_Melania_. The Raised Beaches of Queensland, New South Wales, Victoria, +and Tasmania all contain species of land and freshwater shells identical +with those now found living in the same localities. + +The Raised Beaches of New Zealand contain numerous marine shells all +having living representatives. Some of these elevated beaches occur as +high as 150 feet above sea-level at Taranaki, and at 200 feet near Cape +Palliser in Cook Strait. + +Many species of Pleistocene Mollusca identical with those now living +in Torres Strait, the China Sea and the Philippine Islands are +found in Papua. They occur in the greenish sandy clay of the hills +near the present coast line and comprise the following genera of +Gasteropods:--_Ranella_, _Nassa_, _Mitra_, _Oliva_, _Terebra_, _Conus_, +_Strombus_, _Bulla_ and _Atys_. + + +=Characters of Cephalopoda.--= + +The highest class of the mollusca is the _CEPHALOPODA_ ("head-feet"). +In these shell-fish the extremity of the body or foot is modified, and +furnished with eyes, a funnel and tentacles. It has also strong horny +beaks or jaws which make it a formidable enemy to the surrounding life in +the sea. In the chambered forms of this group the animal partitions off +its shell at regular intervals, like the Pearly Nautilus and the Ammonite, +inhabiting only the last chamber cavity, but still communicating with the +earlier series by a continuous spiral tube (siphuncle). In some forms like +the living squid and the extinct Belemnite, the shell is internal and +either spoon-shaped, or dart-shaped, that is, subcylindrical and pointed. + + +=Characters of Cephalopod Shells.--Nautiloidea.--= + +In geological times the nautiloid forms were the first to appear (in the +Ordovician), and they were either straight shells, as _Orthoceras_, or +only slightly curved, as _Cyrtoceras_. Later on they became more closely +coiled, and as they were thus less likely to be damaged, they gradually +replaced the straight forms. + +The Ammonites have the siphuncle close to the outside of the shell, whilst +in the Nautilus it is more or less median. The sutures or edges of the +septa in _Nautilus_ and its allies are curved or wavy, but not so sharply +flexed or foliaceous as in _Ammonites_. The Nautiloidea range from the +Ordovician and are still found living. + + +=Ammonoidea.--= + +The Ammonoidea appear in Devonian times and die out in the Cretaceous. +They were very abundant in Jurassic times, especially in Europe. + + +=Belemnoidea.--= + +The Belemnoidea, ranging from the Trias to Eocene, comprise the extinct +_Belemnites_, the interesting genus _Spirulirostra_ of Miocene times, and +the living _Spirula_. + + +=Sepioidea.--= + +The Sepioidea or true Cuttle-fishes ("pen-and-ink fish") range from the +Trias to the present day. + + +=Octopoda.--= + +The Octopoda, with _Octopus_ and _Argonauta_ (the paper "Nautilus") are +present-day modifications. The male of the latter is without a shell, the +female only being provided with a delicate boat-shaped shell secreted by +the mantle and the two fin-like expansions of the dorsal arms. + + +=Ordovician Cephalopods.--= + +The Ordovician cephalopods of Australasia are not numerous, and are, +so far as known, practically restricted to the limestones of the +Larapintine series at Laurie's Creek and Tempe Downs, in Central South +Australia. Amongst them may be mentioned _Endoceras warburtoni_ (Fig. +105 A), (a straight form in which the siphuncle is partially filled with +organic deposits); _Orthoceras gossei_; _O. ibiciforme_; _Trochoceras +reticostatum_ (a coiled form); and _Actinoceras tatei_ (a genus +characterised by swollen siphuncular beads between the septa). + +[Illustration: =Fig. 105--PALAEOZOIC CEPHALOPODA.= + + A--Endoceras warburtoni, Eth. fil. Ordovician. South Australia + + B--Orthoceras lineare, Münster sp. Silurian (Yer.) Victoria + + C--Cycloceras ibex, Sow. sp. Silurian (Melb.) Victoria + + D--Phragmoceras subtrigorium, McCoy. Mid Devonian. Victoria + + E--Gastrioceras jacksoni, Eth. fil. Carbopermian. W. Australia + + F--Agathiceras micromphalum, Morris sp. Carbopermian. N.S.W. + +] + + +=Silurian Cephalopods.--= + +Silurian cephalopods are more generally distributed, and in Victoria +constitute an important factor in the molluscan fauna of that system. +_Orthoceras_ and _Cycloceras_ are the best known genera, represented by +_Orthoceras capillosum_, found near Kilmore, Victoria; _O. lineare_ (Fig. +105 B), from the Upper Yarra; _Cycloceras bullatum_, from the Melbournian +of Collingwood and Whittlesea; and _C. ibex_ (Fig. 105 C) from South +Yarra and Flemington, in both Melbournian shale and sandstone. The latter +species occurs also at Rock Flat Greek, New South Wales. Other Victorian +species are _Kionoceras striatopunctatum_, a well-known European fossil +with a reticulated and beaded ornament, found near Warburton and at +McMahon's Creek, Upper Yarra. + +_Orthoceras_ is also recorded from Tasmania and from the Wangapeka beds of +Baton River, New Zealand. _Cyclolituites_, a partially coiled nautilian +is recorded from Bowning, near Yass, New South Wales; whilst the closely +related _Lituites_ is noted from the Silurian of Tasmania. + + +=Devonian Cephalopods.--= + +The only genus of cephalopoda at present recorded from the Devonian of +Victoria is _Phragmoceras_ (_P. subtrigonum_) (Fig. 105 D), which occurs +in the Middle Devonian Limestone of Buchan, E. Gippsland. From beds of +similar age in New South Wales _Orthoceras_, _Cyrtoceras_ and _Goniatites_ +have been noted; whilst the latter genus also occurs near Kimberley, +Western Australia. In Queensland _Gyroceras philpi_ is a characteristic +shell, found in the Fanning and Reid Gap Limestones of the Burdekin +Formation (Middle Devonian). + + +=Carbopermian Cephalopods.--= + +The Carbopermian rocks of New South Wales have yielded _Orthoceras +striatum_, _Cameroceras_, _Nautilus_ and _Agathiceras micromphalum_ +(Fig. 105 F). In Queensland the Gympie Formation contains _Orthoceras_, +_Gyroceras_, _Nautilus_, _Agathiceras micromphalum_ and _A. +planorbiforme_. In Western Australia the Kimberley rocks contain +_Orthoceras_, _Glyphioceras sphaericum_ and _Agathiceras micromphalum_; +whilst the largest known Australian goniatite, _Gastrioceras jacksoni_ +(Fig. 105 E) is found in the Irwin River District. _Actinoceras hardmani_ +is an interesting fossil from the Carbopermian of Lennard River, N.W. +Australia. In Tasmania the genera _Orthoceras_ and _Goniatites_ have been +recorded from beds of similar age. + + +=Triassic Cephalopods.--= + +For Triassic cephalopoda we look to New Zealand, where, in the Mount Potts +_Spiriferina_ Beds of the Kaihiku Series a species of _Orthoceras_ has +been recorded. The Wairoa Series next in succession contains _Orthoceras_ +and an Ammonite. + + +=Jurassic Cephalopods.--= + +[Illustration: =Fig. 106--MESOZOIC and CAINOZOIC CEPHALOPODA.= + + A--Perisphinctes championensis, Crick. Jurassic. West Australia + + B--Nautilus hendersoni, Eth. fil. L. Cretaceous. Queensland + + C--Haploceras daintreei, Eth. sp. L. Cretaceous. Queensland + + D--Crioceras australe, Moore. L. Cretaceous. Queensland + + E--Aturia australis, McCoy. Cainozoic. Victoria + + F--Spirulirostra curta, Tate. Cainozoic (Janjukian). Victoria + +] + +The Jurassic of Western Australia yields a rich cephalopod fauna, from +which may be selected as typical examples the _Nautilus_, _N. perornatus_ +and the following Ammonites: _Dorsetensia clarkei_; _Normanites +australis_; and _Perisphinctes championensis_ (Fig. 106 A). These all +occur in the Greenough River District, and at several other Jurassic +localities in Western Australia. + +The Jurassic system of New Zealand (Putataka Series) contains _Ammonites +aucklandicus_ and _Belemnites aucklandicus_, both from the upper marine +horizon of that series. + +Upper Jurassic Ammonites belonging to the genera _Macrocephalites_ (_M._ +cf. _calloviensis_) and _Erymnoceras_ (_E._ cf. _coronatum_) have been +recorded from Papua. + + +=Lower Cretaceous Cephalopods.--= + +Remains of Cephalopoda are fairly abundant in the Lower Cretaceous of +Australasia. From amongst them may be selected the following--_Nautilus +hendersoni_ (Fig. 106 B) (Q.); _Haploceras daintreei_ (Fig. 106 C) (Q. and +N.S.W.); _Desmoceras flindersi_ (Q. and N.S.W.); _Schloenbachia inflatus_ +(Q.); _Scaphites cruciformis_ (N. Terr.); _Ancyloceras flindersi_ (Q. +and N.S.W.); _Crioceras australe_ (Fig. 106 D) (Q. and S.A.); _Belemites +australis_ (Q.); _B. oxys_ (Q., N.S.W., and S.A.); _B. sellheimi_ (Q. and +S.A.); _B. diptycha_, = _canhami_, Tate, (Q., N.S.W., and S.A.); and _B. +eremos_ (Centr. S.A.). + + +=Upper Cretaceous Cephalopods.--= + +In the Upper Cretaceous (Desert Sandstone) of Queensland there occurs +a Belemnite somewhat resembling _Belemnites diptycha_, but with a very +pointed apex. + + +=Cretaceous Cephalopods, New Zealand.--= + +In New Zealand the Amuri System (Cretaceous) contains fossils which +have been referred to the genera _Ammonites_, _Baculites_, _Hamites_, +_Ancyloceras_ and _Belemnites_, but probably these determinations require +some further revision. A species of Belemnite has also been noted from +probable Cretaceous beds in Papua. + +The Cainozoic System in Victoria contains a true _Nautilus, N. +geelongensis_; and _Aturia australis_ (Fig. 106 E), a nautiloid shell +having zig-zag suture lines and septal necks enclosing the siphuncle. +_A. australis_ is also found in the Oamaru Series of New Zealand; in +Victoria it has an extensive vertical range, from Balcombian to Kalimnan +(Oligocene to Lower Pliocene). Species of _Nautilus_ are also found in the +Janjukian of the Murray River Cliffs; where, in some cases the shell has +been infilled with clear gypsum or selenite, through which can be seen the +tubular siphuncle in its original position. _Spirulirostra curta_ (Fig. +106 F) is an interesting cuttle-bone of rare occurrence. The genus is +represented by two other species only, occurring in the Miocene of Italy +and Germany. In Victoria it is occasionally found in the Janjukian marly +limestone at Bird Rock near Torquay. + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + +PELECYPODA. + + _Ambonychia, macroptera_, Tate. Cambrian: S. Australia. (?) + _Modiolopsis knowsleyensis_, Chapm. L. Ordovician: Victoria. + + _Orthonota australis_, Chapm. Silurian (Melbournian): Victoria. + + _Grammysia cuneiformis_, Eth. fil. Silurian (Melbournian): Victoria. + + _Leptodomus maccoyianus_, Chapm. Silurian (Melbournian): Victoria. + + _Edmondia perobliqua_, Chapm. Silurian (Melbournian): Victoria. + + _Cardiola cornucopiae_, Goldfuss sp. Silurian (Melbournian): Victoria. + + _Panenka gippslandica_, McCoy sp. Silurian (Tanjilian): Victoria. + + _Ctenodonta portlocki_, Chapm. Silurian: Victoria. + + _Nuculites maccoyianus_, Chapm. Silurian: Victoria. + + _Nucula melbournensis_, Chapm. Silurian (Melb.): Victoria. + + _Palaeoneilo victoriae_, Chapm. Silurian (Melb.): Victoria. + + _Pterinea lineata_, Goldfuss. Silurian (Yeringian): Victoria. + + _Lunulicardium antistriatum_, Chapm. Silurian (Tanj.): Victoria. + + _Conocardium costatum_, Cressw. sp. Silurian: Victoria. + + _Conocardium davidis_, Dun. Silurian: New South Wales. + + _Actinopteria boydi_, Conrad sp. Silurian (Yer.): Victoria. + + _Aviculopecten spryi_, Chapm. Silurian (Melb.): Victoria. + + _Modiolopsis complanata_, Sowerby sp. Silurian (Melb.): Victoria. + + _Goniophora australis_, Chapm. Silurian (Yer.): Victoria. + + _Cypricardinia contexta_, Barrande. Silurian (Yer.): Victoria. + + _Paracyclas siluricus_, Chapm. Silurian (Melb.): Victoria. + + _Actinopteria australis_, Dun. Devonian: New South Wales. + + _Lyriopecten gracilis_, Dun. Devonian: New South Wales. + + _Leptodesma inflatum_, Dun. Devonian: New South Wales. + + _Stutchburia farleyensis_, Eth. fil. Carbopermian: New South Wales. + + _Edmondia nobilissima_, de Koninck. Carbopermian: New South Wales. + + _Deltopecten limaeformis_, Morris sp. Carbopermian: New South Wales, + Queensland and Tasmania. + + _Aviculopecten squamuliferus_, Morris sp. Carbopermian: New South + Wales and Tasmania. + + _Aviculopecten tenuicollis_, Dana sp. Carbopermian: New South Wales + and W. Australia. + + _Chaenomya etheridgei_, de Koninck sp. Carbopermian: New South Wales + and Queensland. + + _Maeonia elongata_, Dana. Carbopermian: New South Wales. + + _Pachydomus globosus_, J. de C. Sow. sp. Carbopermian: New South + Wales, Tasmania and Queensland. + + _Eurydesma cordatum_, Morris. Carbopermian: New South Wales and + Queensland. + + _Unio dunstani_, Eth. fil. Trias: New South Wales. + + _Unionella carnei_, Eth. fil. Trias: New South Wales. + + _Corbicula burrumensis_, Eth. fil. Trias: Queensland. + + _Daonella lommeli_, Wissm. sp. Trias: New Zealand. + + _Mytilus problematicus_, Zittel. Trias: New Zealand. + + _Monotis salinaria_, Zittel. Trias: New Zealand. + + _Cucullaea semistriata_, Moore. Jurassic: W. Australia. + + _Trigonia moorei_, Lycett. Jurassic: W. Australia. + + _Ctenostreon pectiniforme_, Schlotheim sp. Jurassic: W. Australia. + + _Astarte cliftoni_, Moore. Jurassic: W. Australia. + + _Unio dacombei_, McCoy. Jurassic: Victoria. + + _Unio eyrensis_, Tate. Jurassic: S. Australia. + + _Nucula truncata_, Moore. Lower Cretaceous: Queensland and S. + Australia. + + _Maccoyella reflecta_, Moore sp. L. Cretaceous: New South Wales, + Queensland (also U. Cretaceous), and S. Australia. + + _Maccoyella barkleyi_, Moore sp. L. Cretaceous: New South Wales, + Queensland and S. Australia. + + _Fissilunula clarkei_, Moore sp. L. Cretaceous: New South Wales, + Queensland, and S. Australia; also Up. Cret. in Queensland and + South Australia. + + _Inoceramus carsoni_, McCoy. Lower Cretaceous: Queensland. + + _Trigonia cinctuta_, Eth. fil. Lower Cretaceous: S. Australia. + + _Mytilus rugocostatus_, Moore. Lower Cretaceous: Queensland and S. + Australia. + + _Cyrenopsis opallites_, Eth. fil. Upper Cretaceous: New South Wales. + + _Conchothyra parasitica_, Hutton. Cretaceous: New Zealand. + + _Dimya dissimilis_, Tate. Cainozoic (Balc.-Kal.): Victoria and South + Australia. + + _Spondylus pseudoradula_, McCoy. Cainozoic (Balc.-Kal.): Victoria and + South Australia. + + _Pecten polymorphoides_, Zittel. Cainozoic (Balc.-Kal.): Victoria and + South Australia; also New Zealand. + + _Cucullaea corioensis_, McCoy. Cainozoic (Balc.-Kal.): Victoria and + South Australia. + + _Leda vagans_, Tate. Cainozoic (Balc.-Kal.): Victoria and South + Australia. + + _Corbula ephamilla_, Tate. Cainozoic (Balc.-Kal.): Victoria and South + Australia. + + _Modiola praerupta_, Pritchard. Cainozoic (Balc.): Victoria. + + _Pecten praecursor_, Chapm. Cainozoic (Janjukian): Victoria. + + _Modiola pueblensis_, Pritchard. Cainozoic (Janjukian): Victoria. + + _Limopsis insolita_, Sow. sp. Cainozoic (Janjukian): Victoria and S. + Australia. Also Oamaru Ser., N.Z. + + _Cardita tasmanica_, Tate. Cainozoic (Janj.): Tasmania. + + _Lucina planatella_, Tate. Cainozoic (Janj.): Victoria and Tasmania. + + _Pecten novaeguineae_, T. Woods. Cainozoic (?Lower Pliocene), Yule + Island, Papua. + + _Ostrea manubriata_, Tate. Cainozoic (Kal.): Victoria. + + _Glycimeris halli_, Pritch. Cainozoic (Kal.): Victoria. + + _Limopsis beaumariensis_, Chapm. Cainozoic (Kalimnan and Werrikooian): + Victoria. + + _Trigonia howitti_, McCoy. Cainozoic (Kal.): Victoria. + + _Meretrix paucirugata_, Tate sp. Cainozoic (Kal.): Victoria. + + _Venus (Chione) subroborata_, Tate, sp. Cainozoic (Kal.): Victoria and + South Australia. + + +SCAPHOPODA. + + _Dentalium tenuissimum_, de Koninck. Mid. Devonian: New South Wales. + + _Dentalium huttoni_, Bather. Jurassic: New Zealand. + + _Dentalium wollumbillensis_, Eth. fil. L. Cretaceous: Queensland. + + _Dentalium, mantelli_, Zittel. Cainozoic: Victoria, S. Australia and + New Zealand. + + +POLYPLACOPHORA. + + _Chelodes calceoloides_, Eth. fil. Silurian: New South Wales. + + _Ischnochiton granulosus_, Ashby and Torr sp. Cainozoic (Balc.): + Victoria. + + _Lorica duniana_, Hull. Cainozoic (Janjukian): Tasmania. + + _Cryptoplax pritchardi_, Hall. Cainozoic (Kal.): Victoria. + + +GASTEROPODA. + + _Ophileta subangulata_, Tate. Cambrian: S. Australia. + + _Platyceras etheridgei_, Tate. Cambrian: S. Australia. + + _Salterella planoconvexa_, Tate. Cambrian: S. Australia. + + _Salterella hardmani_, Foord. Cambrian: W. Australia. + + _Hyolithes communis_, Billings. Cambrian: S. Australia. + + _Scenella tenuistriata_, Chapm. Cambrian (Upper): Victoria. + + _Ophileta gilesi_, Tate. Ordovician: S. Australia. + + _Raphistoma browni_, Tate. Ordovician: S. Australia. + + _Hyolithes leptus_, Chapm. Lower Ordovician: Victoria. + + _Helicotoma johnstoni_, Eth. fil. Ordovician: Tasmania. + + _Coleolus (?) aciculum_, J. Hall. Silurian (Melb.): Victoria. + + _Hyolithes spryi_, Chapm. Silurian (Melb.): Victoria. + + _Conularia ornatissima_, Chapm. Silurian (Melb.): Victoria. + + _Phanerotrema australis_, Eth. fil. Silurian (Yer.): Victoria. + + _Gyrodoma etheridgei_, Cressw. sp. Silurian (Yer.): Victoria. + + _Trematonotus pritchardi_, Cressw. Silurian (Yer.): Victoria. + + _Bellerophon cresswelli_, Eth. fil. sp. Silurian (Yer.) Victoria. + + _Euomphalus northi_, Eth. fil. sp. Silurian (Yer.): Victoria. + + _Cyclonema australis_, Eth. fil. Silurian (Yer.): Victoria. + + _Trochonema montgomerii_, Eth. fil. sp. Silurian: Tasmania. + + _Bellerophon jukesii_, de Koninck. Silurian: New South Wales. + + _Conularia sowerbii_, Defrance. Silurian: Victoria and New South Wales. + + _Euomphalus culleni_, Dun. Devonian: New South Wales. + + _Gosseletina australis_, Eth. fil. Carboniferous: New South Wales. + + _Yvania konincki_, Eth. fil. Carboniferous: New South Wales; and + Carbopermian: Queensland. + + _Bellerophon costatus_, Sow. Carbopermian: W. Australia. + + _Mourlonia humilis_, de Koninck. Carbopermian: West Australia and New + South Wales. + + _Pleurotomaria (Ptychomphalina) morrisiana_, McCoy. Carbopermian: New + South Wales. + + _Keeneia platyschismoides_, Eth. fil. Carbopermian (Lower Marine): New + South Wales. + + _Platyschisma oculum_, Sow. sp. Carbopermian: New South Wales and + Queensland. + + _Macrocheilus filosus_, Sow. Carbopermian: New South Wales. + + _Loxonema babbindonensis_, Eth. fil. Carbopermian: New South Wales. + + _Conularia tenuistriata_, McCoy. Carbopermian: New South Wales and + Queensland. + + _Conularia tasmanica_, Carbopermian: Tasmania. + + _Murchisonia carinata_, Etheridge. Carbopermian: Queensland. + + _Pleurotomaria greenoughiensis_, Eth. fil. Jurassic: W. Australia. + + _Turbo australis_, Moore. Jurassic: W. Australia. + + _Rissoina australis_, Moore. Jurassic: W. Australia. + + _Cinulia hochstetteri_, Moore. Cretaceous: Queensland and S. Australia. + + _Natica ornatissima_, Moore. Cretaceous: S. Australia. + + _Pseudamaura variabilis_, Moore sp. Cretaceous: New South Wales, + Queensland and S. Australia. + + _Anchura wilkinsoni_, Eth. fil. Cretaceous: Queensland and S. + Australia. + + _Rostellaria waiparensis_, Hector. Cretaceous: New Zealand. + + _Niso psila_, T. Woods. Cainozoic (Balc.-Kal.): Victoria and S. + Australia. + + _Crepidula unguiformis_, Lam. Cainozoic (Balc.-Recent): Victoria and + Tasmania. + + _Natica hamiltonensis_, Tate. Cainozoic (Balc.-Recent): Victoria and + South Australia. + + _Turritella murrayana_, Tate. Cainozoic (Balc.-Kal.): Victoria, S. + Australia and Tasmania. + + _Cerithium apheles_, T. Woods. Cainozoic (Balc.-Kal.): Victoria. + + _Volutilithes antiscalaris_, McCoy sp. Cainozoic (Balc.-Werrikooian): + Victoria. + + _Ancilla pseudaustralis_, Tate sp. Cainozoic (Balc.-Kal.): Victoria, + S. Australia and Tasmania. + + _Cypraea ampullacea_, Tate. Cainozoic (Balc.): Victoria. + + _Murex didyma_, Tate. Cainozoic (Balc.): Victoria. + + _Eburnopsis aulacoessa_, Tate. Cainozoic (Balc.): Victoria. + + _Cancellaria calvulata_, Tate. Cainozoic (Balc.): Victoria. + + _Vaginella eligmostoma_, Tate. Cainozoic (Balc.): Victoria. + + _Eutrochus fontinalis_, Pritchard. Cainozoic (Janjukian): Victoria. + + _Turbo atkinsoni_, Pritchard. Cainozoic (Janjukian): Tasmania and + Victoria. + + _Scala lampra_, Tate sp. Cainozoic (Janjukian): S. Australia. + + _Natica gibbosa_, Hutton. Cainozoic (Janjukian): Victoria. Also Oamaru + and Wanganui Series: New Zealand. + + _Morio wilsoni_, Tate. Cainozoic (Janjukian): Victoria. + + _Voluta heptagonalis_, Tate. Cainozoic (Janjukian): S. Australia. + + _Volutilithes anticingulatus_, McCoy sp. Cainozoic (Janjukian): + Victoria and Tasmania. Also Papua. + + _Bathytoma paracantha_, T. Woods sp. Cainozoic (Janj.): Victoria and + Tasmania. Also Papua. + + _Dolium costatum_, Deshayes. Cainozoic. (? Lower Pliocene): Yule + Island, Papua. + + _Bankivia howitti_, Pritch. Cainozoic (Kal.): Victoria. + + _Eglisia triplicata_, Tate sp. Cainozoic (Kal.): Victoria. + + _Voluta masoni_, Tate. Cainozoic (Kal.): Victoria. + + _Ancilla papillata_, Tate sp. Cainozoic (Kal.): Victoria. + + _Drillia wanganuiensis_, Hutton. Cainozoic (Kal.): Victoria. Also + Petane Series: New Zealand. + + _Terebra geniculata_, Tate. Cainozoic (Kal.): Victoria. + + _Pleurotomaria tertiaria_, McCoy. Cainozoic (Kal.): Victoria. Also + Oamaru Series: New Zealand. + + _Scala lyrata_, Zittel sp. Cainozoic (Oamaru): New Zealand. + + _Natica darwinii_, Hutton. Cainozoic (Oamaru): New Zealand. + + _Turritella cavershamensis_, Harris. Cainozoic (Oamaru): New Zealand. + + _Ancilla hebera_, Hutton sp. Cainozoic (Oamaru): New Zealand. Also + (Balc. and Janj.): Victoria, South Australia and Tasmania. + + _Pleurotoma hamiltoni_, Hutton. Cainozoic (Oamaru): New Zealand. + + _Natica ovata_, Hutton. Cainozoic (Awatere Series): New Zealand. + + _Struthiolaria sulcata_, Hutton. Cainozoic (Awatere Series): New + Zealand. + + _Trophon expansus_, Hutton. Cainozoic (Petane Series): New Zealand. + + _Pisania drewi_, Hutton. Cainozoic (Petane Series): New Zealand. + + _Bankivia fasciata_, Menke. Cainozoic (Werrikooian-Recent): Victoria. + + _Astralium aureum_, Jonas sp. Cainozoic (Werrikooian-Recent): Victoria. + + _Natica subinfundibulum_, Tate. Cainozoic (Balc.-Werr.): Victoria and + S. Australia. + + _Nassa pauperata_, Lam. Cainozoic (Werr.-Rec.): Victoria. + + _Helix tasmaniensis_, Sow. Cainozoic (Pleistocene): Tasmania. + + _Helix geilstonensis_, Johnston. Cainozoic (Pleistocene): Tasmania. + + _Panda atomata_, Gray sp. Cainozoic (Pleist.-Rec.): Victoria and New + South Wales. + + +CEPHALOPODA. + + _Endoceras warburtoni_, Eth. fil. Ordovician: S. Australia. + + _Orthoceras gossei_, Eth. fil. Ordovician: S. Australia. + + _Orthoceras ibiciforme_, Tate. Ordovician: S. Australia. + + _Trochoceras reticostatum_, Tate. Ordovician: S. Australia. + + _Actinoceras tatei_, Eth. fil. sp. Ordovician: S. Australia. + + _Orthoceras capillosum_, Barrande. Silurian: Victoria. + + _Orthoceras lineare_, Münster sp. Silurian (Yer.): Victoria. + + _Cycloceras bullatum_, Sow. sp. Silurian (Melbournian): Victoria. + + _Cycloceras ibex_, Sow. sp. Silurian (Melbournian): Victoria. + + _Kionoceras striatopunctatum_, Münster sp. Silurian (Tanjilian): + Victoria. + + _Phragmoceras subtrigonum_, McCoy. Mid. Devonian: Victoria. + + _Gyroceras philpi_, Eth. fil. Mid. Devonian: Queensland. + + _Orthoceras striatum_, Sow. Carbopermian: New South Wales. + + _Agathiceras micromphalum_, Morris sp. Carbopermian: New South Wales + and W. Australia. + + _Gastrioceras jacksoni_, Eth. fil. Carbopermian: W. Australia. + + _Actinoceras hardmani_, Eth. fil. Carbopermian: N.W. Australia. + + _Nautilus perornatus_, Crick. Jurassic: W. Australia. + + _Dorsetensia clarkei_, Crick. Jurassic: W. Australia. + + _Normanites australis_, Crick sp. Jurassic: W. Australia. + + _Perisphinctes championensis_, Crick. Jurassic: W. Australia. + + _Ammonites aucklandicus_, Hector. Jurassic: New Zealand. + + _Belemnites aucklandicus_, Hector. Jurassic: New Zealand. + + _Nautilus hendersoni_, Eth. fil. Lower Cretaceous: Queensland. + + _Haploceras daintreei_, Etheridge sp. Lower Cretaceous: Queensland and + New South Wales. + + _Ancyloceras flindersi_, McCoy. Lower Cretaceous: Queensland and New + South Wales. + + _Crioceras australe_, Moore. Lower Cretaceous: Queensland and S. + Australia. + + _Scaphites eruciformis_, Eth. fil. Lower Cretaceous: Northern + Territory. + + _Belemnites diptycha_, McCoy. Lower Cretaceous: Queensland, New South + Wales, and S. Australia. + + _Belemnites eremos_, Tate. Lower Cretaceous: S. Australia. + + _Nautilus geelongensis_, Foord. Cainozoic (Janjukian): Victoria. + + _Aturia australis_, McCoy. Cainozoic (Balc.-Kal.): Victoria. Oamaru + Series: New Zealand. + + _Spirulirostra curta_, Tate. Cainozoic (Janjukian): Victoria. + + + * * * * * + + +LITERATURE. + + +MOLLUSCA. + + Cambrian.--Foord, A. H. Geol. Mag., Dec. III. vol. VII. 1890, pp. + 98, 99 (Pteropoda). Tate, R. Trans. R. Soc. S. Austr., vol. XV. + 1892, pp. 183-185 (Pelec. and Gastr.), pp. 186, 187 (Pteropoda). + Etheridge, R. jnr. Trans. R. Soc. S. Austr., vol. XXIX. 1905, p. + 251 (Pteropoda). Chapman, F. Proc. R. Soc. Vict., vol. XXIII. pt. + II. 1910, pp. 313, 314 (Gastr.). + + Ordovician.--Etheridge, R. jnr. Parl. Papers, Leg. Assemb., S. Austr., + No. 158, 1891, pp. 9, 10 (Gastr. and Ceph.). Tate, R. Rep. Horn. + Sci. Exped., pt. 3, 1896, pp. 98-110. Chapman, F. Proc. R. Soc. + Vic., vol. XV. pt. II. 1903, pp. 119, 120 (_Hyolithes_). + + Silurian.--McCoy, F. Prod. Pal. Vic., Dec. VI. 1879, pp. 23-29. + Etheridge, R. jnr. Rec. Austr. Mus., vol. I. No. 3, 1890, pp. + 62-67 (Gastr.). Idem, ibid., vol. I. No. 7, 1891, pp. 126-130 + (Pelec. and Gastr.). Cresswell, A. W. Proc. R. Soc. Vict., vol. + V. 1893, pp. 41-44. Etheridge, R. jun. Rec. Austr. Mus., vol. + III. No. 4, 1898, pp. 71-77 (Gastr.). Idem, Rec. Geol. Surv. New + South Wales, vol. V. pt. 2, 1898, pp. 67-70 (_Chelodes_). De + Koninck, L. G. Mem. Geo. Surv. New South Wales, Pal. No. 6, 1898, + pp. 29-35. Etheridge, R. jnr. Prog. Rep. Geol. Surv. Vict., No. + XI. 1899, pp. 34, 35 (Pelec.). Idem, Rec. Austr. Mus., vol. V. + No. 2, 1904, pp. 75-77 (Ceph.). Chapman, F. Proc. R. Soc., Vict., + vol. XVI. pt. 11. 1904, pp. 336-341 (Pteropoda). Idem, Mem. Nat. + Mus. Melbourne, No. 2, 1908 (Pelecypoda). + + Devonian.--McCoy, F. Prod. Pal., Vict., Dec. IV. 1876, pp. 18, 19 + (Ceph.). Etheridge, R. jnr. Geol. and Pal. Queensland, 1892, p. + 69 (_Gyroceras_). De Koninck, L. G. Mem. Geol. Surv. New South + Wales, Pal. No. 6, 1898, pp. 85-105. + + Carboniferous.--Etheridge, R. jnr. Rec. Austr. Mus., vol. III. No. 1, + 1897, pp. 7-9 (_Actinoceras_). Idem, Geol. Surv. W.A., Bull. No. + 27, 1907, pp. 32-37. + + Carbopermian.--Morris, J., in Strzelecki's Phys. Descr. of New South + Wales, etc., 1845, pp. 270-278 and 285-291. Foord, A. H. Geol. + Mag., Dec. III. vol. VII. 1890, pp. 103, 104. Etheridge, R. + jnr. Geol. and Pal. Queensland, 1892, pp. 264-296. Idem., Proc. + Linn. Soc. New South Wales, vol. IX. 1895, pp. 530-537 (Pelec. + and Gastr.). De Koninck, L. G. Mem. Geol. Surv. New South Wales, + Pal. No. 6, 1898, pp. 203-274. Etheridge, R. jnr. and Dun, W. + S. Mem. Geol. Surv. New South Wales, Pal. No. 5, vol. II. pt. + I. 1906 (_Palaeopecten_). Idem, ibid., vol. II., pt. 2, 1910 + (_Eurydesma_). + + Trias.--Zittel, K. Novara Exped., vol. I. Abth. II. Geol. Theil., + 1864, pp. 26-29. Etheridge, R. jnr. Mem. Geol. Surv. New South + Wales, Pal. No. 1, 1888, pp. 8-14. + + Jurassic.--Zittel, K. Novara Exped., vol. I., Abth. II. Geol. Theil., + 1864, pp. 20-34. Moore, C. Quart. Journ. Geol. Soc., vol. XXVI. + pp. 245-260 (Jurassic and Cretaceous Moll.). Etheridge, R. jnr. + ibid., vol. XXVIII. 1872, pp. 317-359 (Palaeozoic, Jur. and Cret. + Moll.). Crick, G. C. Geol. Mag., Dec. IV. vol. I. 1894, pp. + 385 393 and 433-441 (Ceph.). Chapman, F. Proc. R. Soc. Vict., + vol. XVI. pt. II. 1904, pp. 327-332. Marshall, P. Trans. New + Zealand Inst., vol. XLI. 1909, pp. 143-145 (New Zealand Ceph.). + Etheridge, R. jnr. Geol. Surv. W.A. Bull. No. 36, 1910, pp. 30-40. + + Cretaceous.--Etheridge, R. jnr. Geol. and Pal. Queensland, 1892, pp. + 445-503 and 561-574. Idem, Geol. Surv. Queensland, Bull. No. 13, + 1901, pp. 13-35. Idem, Mem. Roy. Soc. S. Aust., vol. II. pt. 1, + 1902 (S.A. Moll.). Idem, Mem. Geol. Surv. New South Wales, Pal. + No. 11, 1902, pp. 16-49 (New South Wales Moll.). + + Cainozoic.--Zittel, K. Novara Exped. Geol. Theil., vol. I. Abth. II. + 1864, pp. 34-55 (Pelec. and Gastr. New Zealand). McCoy, F. Prod., + Pal. Vict., Dec. I. 1874; Dec. II. 1875; Dec. III. 1876; Dec. V. + 1877; Dec. VI. 1879. Woods, J. E. T. Proc. R. Soc. Tas. (1875), + 1876, pp. 13-26 (Table Cape Moll.). Idem, Proc. Linn. Soc. New + South Wales, vol. III. 1879, pp. 222-240 (Muddy Creek Moll.). + Idem, ibid., vol. IV. 1880, pp. 1-24. Hutton, F. W. Trans. New + Zealand Inst. vol. IX. 1877, pp. 593-598. Ibid., vol. XVII. + 1885, pp. 313-332 (New Zealand Pelec. and Gastr.). Idem, Proc. + Linn. Soc. New South Wales, vol. I. 2nd ser. (1886), 1887, pp. + 205-237, (distr. lists, Pareora and Oamaru). Idem, Macleay, Mem. + Vol. Linn. Soc. New South Wales, 1893, pp. 35-92 (Pliocene Moll. + New Zealand). Tate, R. Trans. R. Soc. S. Austr., vol. VII. 1886, + pp. 96-158, and vol. IX., 1887, pp. 142-189 (Pelec.); ibid., pp. + 190-194 (Scaphopoda); ibid., 194-196 (Pteropoda). Idem, ibid., + vol. X. 1888, pp. 91-176; vol. XI. 1889, pp. 116-174; vol. XIII. + 1890, pp. 185-235; and vol. XVII. 1893, pp. 316-345 (Gastr.). + Idem, Journ. R. Soc., New South Wales, vol. XXVII. 1893, pp. + 169-191. Idem, ibid., vol. XXXI. 1897, pp. 392-410 (Gastr. and + Pelec.). Idem, Trans. Roy. Soc. S. Austr., vol. XXIII. 1899, pp. + 260-277 (Revision of Moll.). Pritchard, G. B. Proc. Roy. Soc. + Vic., vol. VII. 1895, pp. 225-231 (Pelec.). Idem, ibid., vol. + VIII. 1896, pp. 79-141 (Moll. of T. Cape). Idem, ibid., vol. XI. + pt. I. 1898, pp. 96-111 (Gastr.). Idem, ibid., vol. XIV. pt. I. + 1901, pp. 22-31 (Pelec.). Idem, ibid., vol. XVI. pt. II. 1903, + pp. 87-103 (Pelec.). Idem, ibid., vol. XVI. pt. I. 1903, pp. + 83-91 (_Pleurotomaria_). Idem, ibid., vol. XVII. pt. I. 1904, pp. + 320-337 (Gastr.) Idem, ibid., vol. XXVI. (N.S.) pt. I. 1913, pp. + 192-201 (Volutes). Hall, T. S. Proc. R. Soc. Vict., vol. XVII. + pt. II. 1905, pp. 391-393 (Chitons). Ashby, E. and Torr. W. G. + Trans. R. Soc. S. Austr., vol. XXV. 1901, pp. 136-144 (Chitons). + Thomson, J. A. Trans. New Zealand Inst., Vol. XL. 1908, pp. 102, + 103 (N.Z. Moll.). Chapman, F. Proc. R. Soc. Vict. vol. XX. pt. + II. 1908, pp. 218-220 (Chiton). Idem, ibid., vol. XXV. pt. I. + 1912, pp. 186-192 (Gastr.). + + + + +CHAPTER XI. + +FOSSIL TRILOBITES, CRUSTACEA AND INSECTS. + + +=Arthropods and their Structure.--= + +The above-named fossil groups are included by zoologists in the +sub-kingdom Arthropoda ("joint-footed animals"). The Arthropods possess a +body and limbs composed of a number of jointed segments covered externally +with a hard, shelly material and separated by a softer, flexible skin. +They have no internal skeleton, and therefore the only portion which can +be preserved in the fossil state is the harder part of the outer covering. +Under exceptional conditions of fossilisation, however, even frail insects +such as ants, wasps and dragon-flies are sometimes found more or less +wholly preserved and showing their original minute structure. + + +=Subdivisions of Arthropoda.--= + +The principal representatives of the group of the Arthropods which are +found as fossils include the Trilobites; various Crustacea proper, as +Crabs, Lobsters, Shrimps, Pod-shrimps and Water-fleas; the Insects; +and occasionally Spiders and Scorpions (Arachnida). The King-crabs and +Eurypterids (as the extinct _Pterygotus_) form a separate sub-class, the +Merostomata, which are placed by some authors in the group of Spiders and +Scorpions: their remains date back to the time when the older Palaeozoic +strata were deposited. + + +=Crustacea, an Archaic Group.--= + +A typical division of the Arthropod group, and one which was well +represented from the earliest period up to the present day, is the +_CRUSTACEA_. As the name denotes, these animals are generally invested +with a strong shelly covering or "crust," usually of horny or chitinous +material, which in some forms is strengthened by deposits of phosphate +of lime. Of the horny condition of the shell the groups of the bivalved +Crustacea (Ostracoda) and the "water-fleas" (Entomostraca) supply notable +instances; whilst the limy-structured shell is seen in the common crab. +Some authorities separate the great extinct group of the Trilobites +from the rest of the Crustacea; but it will here be convenient, in a +preliminary study, to consider them together. + + +=Development of Crustacea.--= + +The development of the lower forms of the Crustacea is interesting, +from the fact that the young usually escapes from the egg in a larval +state known as a "nauplius." In this stage there are no segments to +the body, and but a solitary median eye, such as may be seen in the +common water-flea known to microscopists as _Cyclops_. The three pairs +of appendages seen in this larval crustacean represent the two pairs of +antennae and the jaws or mandibles of the full-grown form. + +Among the higher Crustacea, however, there is no larval form; the young +escaping from the egg in a more or less highly developed condition +resembling the adult. The group of the Crabs, Lobsters and Shrimps (or +Decapoda, _i.e._, having ten ambulatory feet) exhibit a larval stage in +which the young form ("zoea") has a segmented abdomen and seven pairs of +appendages. + + +=Trilobites.--= + +The first group of arthropods here described is that of the _TRILOBITES_. +These were so named on account of the three-lobed form of the body. +This particular feature distinguishes them from the Crustacea proper; +which includes the Phyllopods (with leaf-like limbs), as the freshwater +_Estheria_, the Ostracoda or Bivalved Water-fleas, the Barnacles or +Cirripedia and the Higher Crustacea (Malacostraca), including Shrimps, +Crabs, and Lobsters, of which the oldest representatives are the +Pod-shrimps (Phyllocarida). + + +=Habits of Trilobites.--= + +The remains of these primitive but often strikingly ornamented +crustacean-like animals, the trilobites, are found in comparative +abundance in the limestones, mudstones, and even the sandstones of the +older sedimentary rocks of Australasia. They were amongst the most +prolific types of animal life existing in the seas of Palaeozoic times, +and are especially characteristic of Cambrian, Ordovician and Silurian +rocks. Trilobites, as a group, seem to have adapted themselves to almost +all conditions of marine life: some are found in the hardened black mud +of shallow waters, whilst others are to be looked for in the limestones +and excessively fine sediments of deeper waters. In all probability +certain of these forms crawled over the soft, oozy sea-bed in order to +obtain their food, and consequently their remains in the stratified rocks +would be restricted to the fine black shales; whilst the freely swimming +forms could change their habitat at will, and would be found alike in +sandy or clayey deposits. As some indication of their varied habits, the +eyes of trilobites differ greatly in size. They are always compound like +the eye of the house-fly, though of a semi-lunar shape. In some forms the +eyes are very small or even absent, whilst in others they are exceedingly +large and prominent. This latter feature probably indicates their +frequenting moderately deep water. + +[Illustration: =Fig. 107--Diagram-restoration of an Australian Trilobite.= + +(Dalmanites meridianus, Eth. fil. and Mitch, sp.) + +To show the sutures or joints, and the structure of the back of the +carapace. + + About 2/3 natural size. +] + + +=Structure of Trilobites.--= + +The complete structure and zoological relationship of the trilobites has +always been open to some doubt. As regards the former, within recent +years exceptionally well-preserved specimens from the Utica Slates and +the Cincinnati Limestone of Ohio, rocks of Ordovician age, have been +discovered and dissected, whereby our knowledge of the organisation of +this group is greatly advanced. These remarkable fossil remains show that +the Trilobites bore on their under surface a number of appendages, one +pair to each segment, except that of the anal. The front pair is whip-like +and served as antennae; the others are branched, the forward portion being +a crawling limb, and the hinder, which was fringed with bristles or thin +plates, may have served either for swimming or breathing. At the base of +the four pairs of appendages attached to the head there was an arrangement +for biting the food, from whence it was passed to the mouth. Taking one +of the commonest Australasian trilobites, _Dalmanites meridianus_, for +an example of general structure, and looking at the back of the shell or +upper surface, we see the trilobate (three-lobed) form well defined (Fig. +107). The central ridge is termed the axis, and on either side of this are +arranged the pleural lobes, each well marked transverse division of which, +in the central or thoracic region, being a pleuron or rib. The whole body +is divided into three more or less distinct portions,--the head-shield or +cephalon, the thorax, and the tail-shield or pygidium. The central area +of the head-shield is called the glabella or cranidium, against which, on +either side, are placed the free cheeks carrying the compound sessile eyes +when present. The appendages of the head are pediform or leglike, arranged +in five pairs, and biramous or forked, excepting the antennae, which are +simple and used as sensory organs. In front of the mouth is the hypostoma +or forelip, and behind it is the metastoma or hind-lip. The segments of +the head-shield are most closely united, and in all the trilobites are of +the same number. Those of the thorax have flexible joints and are variable +in number. The segments of the abdomen are fused together and form a +caudal shield or pygidium. + +The larval stage of the trilobite was a protonauplian form (that is more +primitive than the nauplius), the protoaspis; the adult stage, being +attained by the addition of segments at the successive moults. + +The earliest known trilobites in Australia are some Cambrian species from +South Australia, Western Australia, Victoria, and Tasmania. + + +=Lower Cambrian Trilobites.--= + +[Illustration: =Fig. 108--CAMBRIAN TRILOBITES.= + + A--Ptychoparia howchini, Eth. fil. L. Cambrian. South Australia + + B--Dolichometopus tatei, H. Woodw. L. Cambrian. South Australia + + C--Agnostus australiensis, Chapm. Up. Cambrian. Victoria + + D--Ptychoparia thielei, Chapm. Up. Cambrian. Victoria + + E--Dikellocephalus florentinensis, Eth. fil. L. Cambrian. Tasmania + +] + +In the Lower Cambrian Limestone of Yorke Peninsula, South Australia, the +following trilobites occur:--a species doubtfully referred to _Olenellus_ +(? _O. pritchardi_); _Ptychoparia howchini_ (Fig. 108 A); _P. australis_; +_Dolichometopus tatei_ (Fig. 108 B); and _Microdiscus subsagittatus_. +The Cambrian of the Northern Territory contains _Olenellus brownii_. In +Western Australia _Olenellus forresti_ is found in similar beds. + + +=Upper Cambrian Trilobites.--= + +The Dolodrook Limestone (Upper Cambrian) of Gippsland, Victoria, +contains the remains of the primitive little trilobite _Agnostus_ (_A. +australiensis_, Fig. 108 C); _Crepicephalus_ (_C. etheridgei_); and +_Ptychoparia_ (_P. thielei_ (Fig. 108 D) and _P. minima_). The Upper +Cambrian sandstones of Caroline Creek, Tasmania, contain _Dikellocephalus_ +(_D. tasmanicus_); a species of _Asaphus_ and _Ptychoparia_ (_P. +stephensi_). Beds of the same age in the Florentine Valley, Tasmania, have +yielded _Dikellocephalus_ (_D. florentinensis_, Fig. 108 E). + + +=Ordovician Trilobites.--= + +Trilobites of Lower Ordovician age or even older, are found in the +Knowsley beds near Heathcote in Victoria. They are referred to two genera, +_Dinesus_ and _Notasaphus_. Both forms belong to the ancient family +of the Asaphidae. Associated with these trilobites are some doubtful +species of sea-weed, spicules of siliceous sponges, traces of threadlike +hydrozoa, some fragments of graptolites allied to _Bryograptus_, and +several brachiopods. At the Lyndhurst Gold-fields, near Mandurama, New +South Wales, trilobites related to the genus _Shumardia_ have been found +associated with brachiopods (lamp-shells), pteropods (sea-butterflies), +and graptolites (hydrozoa) of an Upper Ordovician facies. + +The limestone beds at Laurie's Creek and other localities in Central +Australia contain remains of _Asaphus illarensis_, _A. howchini_ and _A. +lissopelta_; whilst in the limestone and quartzite of Middle Valley, Tempe +Downs, _A. thorntoni_ also occurs. + + +=Silurian Trilobites.--= + +[Illustration: =Fig. 109--OLDER SILURIAN TRILOBITES.= + + A--Ampyx parvulus, Forbes, var. jikaensis, Chapm. Silurian (Melb.) + Victoria + + B--Cypaspis spryi, Gregory. Silurian (Melb.) Victoria + + C--Homalonotus harrisoni, McCoy. Silurian (Melb.) Victoria + + D--Phacops latigenalis, Eth. fil. and Mitch. Silurian. N.S. Wales + +] + +Trilobites are well-known fossils in the Australasian Silurian strata. As +they occur rather abundantly along with other fossils in rocks of this age +they are extremely useful aids in separating the system into the different +beds or zones. In Victoria the Silurian is divisible into two sets of +beds: an older, or Melbournian stage (the bed-rock of Melbourne) and a +younger, Yeringian (Lilydale series). Trilobites of Melbournian age are +found to belong to the genera _Ampyx_, _Illaenus_, _Proetus_, _Cyphaspis_, +_Encrinurus (Cromus)_ and _Homalonotus_. The commonest species are +_Cyphaspis spryi_ (Fig. 109 B), and _Encrinurus (Cromus) spryi_ from the +South Yarra mudstones; and _Ampyx parvulus_, var. _jikaensis_ (Fig. 109 +A), and _Homalonotus harrisoni_ (Fig. 109 C), from the sandstone of Moonee +Ponds Creek. + +The handsome _Dalmanites meridianus_ and _Homalonotus vomer_ occur at +Wandong in what appear to be passage beds between the Melbournian and +Yeringian. + +The Yeringian of Victoria is far richer in trilobites than the preceding +series, and includes the genera _Proetus_, _Cyphaspis_, _Bronteus_, +_Lichas_, _Odontopleura_, _Encrinurus_, _Calymene_, _Homalonotus_, +_Cheirurus_, and _Phacops_. The rocks in this division occur as mudstones, +limestones, and occasionally sandstones and conglomerates. The mudstones, +however, prevail, and these pass insensibly into impure limestones of +a blue-black colour, weathering to brown, as at Seville; the change of +structure indicating less turbid water. At Lilydale, and on the Thomson +River, as well as at Loyola and Waratah Bay, almost pure limestone +occurs, which represents clear water conditions, not necessarily deep; +there, however, trilobites are scarce, and the prevailing fauna is that +of an ancient coral reef. Some described Yeringian species are _Lichas +australis_ (Fig. 110 A), _Odontopleura jenkinsi_ (Fig. 110 B) (found +also in New South Wales), _Encrinurus punctatus_ (Fig. 110 C), _Calymene +tuberculosa_, _Bronteus enormis_, _Phacops sweeti_, and _P. serratus_ +(Fig. 110 E). In _Calymene_ ("covered up") the joints of the thorax +are facetted at the angles, so that each pleuron could work over that +immediately behind; in consequence of this it could roll itself up like +a woodlouse or slater, hence the name of the genus. This trilobite also +occurs in England, and is there known amongst the quarry men and fossil +collectors as the "Dudley Locust." Perhaps the most characteristic and +common trilobite of the Yeringian series in Victoria is _Phacops sweeti_ +(Fig. 110 D), formerly identified with Barrande's _P. fecundus_, from +which it differs in the longer and larger eye with more numerous lenses. +It is found in Victoria in the Upper Yarra district near the junction +of the Woori Yallock and the Yarra Rivers; north-west of Lilydale; near +Seville; at Loyola near Mansfield; and at Fraser's Creek near Springfield, +Kilmore. + +[Illustration: =Fig. 110--NEWER SILURIAN TRILOBITES.= + + A--Lichas australis, McCoy. Silurian (Yeringian). Victoria + + B--Odontopleura jenkinsi. Eth. fil. and Mitch. Silurian. N.S. Wales + + C--Encrinurus punctatus, Brunnich sp. Silurian. N.S. Wales. + + D--Phacops sweeti, Eth. fil. and Mitch. Silurian. N.S. Wales + + E--Phacops serratus, Foerste. Silurian. N.S. Wales + +] + +In New South Wales trilobites are abundant in the Yass district, amongst +other localities, where the upper beds, corresponding to the Yeringian +of Victoria, are well developed. _Dalmanites meridianus_ is common to +the Silurian of New South Wales, Victoria, and Tasmania. In Victoria +this handsome species is found in the hard, brown, sandy mudstone of +Broadhurst's and Kilmore Creeks, and, as previously noted, in the hard, +blue mudstone of Wandong. At the latter locality specimens may be found in +the railway ballast quarry, where they are known to the workmen as "fossil +butterflies." The species also occurs at the famous fossil locality of +Hatton's Corner, Yass; at Bowning; and at Limestone Creek, all in New +South Wales. Other trilobites occurring in the Silurian of New South Wales +are _Odontopleura jenkinsi_, _O. bowningensis_, _Cheirurus insignis_ and +_Phacops latigenalis_ (Fig. 109 D). + +In the Wangapeka series of New Zealand the calcareous shales and +limestones of the upper division contain _Calymene blumenbachii_, +_Homalonotus knightii_ and _H. expansus_. + + +=Devonian Trilobites.--= + +Trilobites suddenly became rare in the Australian Devonian. The only +known examples of trilobite remains belong to a species of _Cheirurus_ +occasionally found in the Middle Devonian limestone of Buchan, Victoria; +and a species of _Proetus_ in the Devonian of Barker Gorge, Napier Range, +West Australia. + + +=Carbopermian Trilobites.--= + +Trilobites of Carbopermian age are found in New South Wales, Queensland, +and Western Australia. All the genera belong to the family Proetidae. +The genera _Phillipsia_ (_P. seminifera_, Fig. 111 A), _Griffithides_ +(_G. eichwaldi_, Fig. 111 B), and _Brachymetopus_ (_B. strzelecki_, Fig. +111 C) occur in New South Wales. _Griffithides eichwaldi_ is also found +in Queensland. Other Queensland species are _Phillipsia woodwardi_, _P. +seminifera_ var. _australasica_ and _P. dubia_. _Phillipsia grandis_ is +found in the Carbopermian of the Gascoyne River, Western Australia. + +[Illustration: =Fig. 111--CARBONIFEROUS TRILOBITES and a PHYLLOPOD.= + + A--Phillipsia seminifera, Phillips. Carboniferous. N.S. Wales + + B--Griffithides eichwaldi, Waldheim. Carboniferous. N.S. Wales + + C--Brachymetopus strzelecki, McCoy. Carboniferous. N.S. Wales + + D--Estheria coghlani, Cox. Triassic. N.S. Wales + +] + + +=Phyllopoda in Carboniferous, Triassic and Jurassic.= + +The _PHYLLOPODA_, which belong to the Crustacea in the strict sense of +the term, comprise the Estheriidae and Cladocera (water-fleas). The +former group is represented by _Leaia mitchelli_, which is found in the +Upper Carboniferous or Carbopermian of the Newcastle District, New South +Wales. In the still later Hawkesbury series (Triassic) of New South Wales, +_Estheria coghlani_ (Fig. 111 D) occurs. This species is a minute form, +the carapace measuring from 1.25mm. to 2mm. in the longer diameter of +the shell. In the upper part of the Wairoa Series (Triassic) of Nelson, +New Zealand, there is found another species of _Estheria_, identified +with a European form _E. minuta_. _Estheria mangaliensis_ is another form +occurring in the Jurassic (Ipswich series) of Queensland. At the present +day these little _Estheriae_ sometimes swarm in countless numbers in +freshwater lakes or salt marshes. + + +=Ostracoda: Their Structure.--= + +Passing on to the next group, the bivalved _OSTRACODA_, we note that these +have existed from the earliest geological periods to the present day. They +are usually of minute size, commonly about the sixteenth of an inch in +length, although some attained a length of nearly one inch (_Leperditia_). +Their bodies are indistinctly segmented, and are enclosed within a +horny or calcareous shell. This shell consists of two valves which are +joined along the back by a ligament or hinge, the ends and ventral edge +remaining quite free. The pairs of appendages present are the antennae +(2), mandibles (1), maxillae (2), and thoracic feet (2). The only portion +found in the fossil state is the bivalved carapace, the two valves being +frequently met with still united, especially when these tiny animals have +settled down quietly on the sea-bed and have been quickly covered with +sediment. + + +=Features of the Ostracod Carapace.--= + +Since the body parts of the ostracod are wanting in the fossil examples, +the generic determination is attended with some difficulty, especially +in regard to the smooth or bean-shaped forms. The chief distinctive +characters to note are, the contour of the carapace seen in three +directions (top, side and end views), the structure of the hinge, and +the position and figure of the muscle-spots or points of adhesion of the +muscular bands which hold or relax the two valves. The valves in certain +genera fit closely upon one another. In others, one overlaps the other, +the larger being sometimes the right (as in _Leperditia_), sometimes +the left (as in _Leperditella_). The hinge-line is often simple or +flange-like, or it may consist of a groove and corresponding bar, or there +may be a series of teeth and sockets. Lateral eye-tubercles are sometimes +seen on the surface of the valve, whilst in the animal there was also a +small eye. + + +=Habits of Ostracoda.--= + +Ostracoda swarmed in many of the streams, lakes and seas of past +geological times, and they still exist in vast numbers under similar +conditions. Like some other minute forms of life, they played a most +important part in building up the rock formations of the sedimentary +series of the earth's crust; and by the decomposition of the organism +itself they are of real economic value, seeing that in some cases their +decay resulted in the subsequent production of oil or kerosene shales +and bituminous limestones. The Carboniferous oil shales in the Lothians +of Scotland, for example, are crowded with the carapaces of Ostracoda +associated with the remains of fishes. + + +=Cambrian Ostracoda.--= + +Some undescribed forms of the genus _Leperditia_ occur in the hard, +sub-crystalline Cambrian Limestone of Curramulka, South Australia. + + +=Silurian Ostracoda.--= + +In Victoria and New South Wales the oldest rocks from which we have +obtained the remains of Ostracoda up to the present, are the uppermost +Silurians, in which series they occur both in the limestone and the +mudstone. In Victoria their bivalved carapaces are more often found in the +limestone; but one genus, _Beyrichia_, is also met with in abundance in +the mudstone. These mudstones, by the way, must have originally contained +a large percentage of carbonate of lime, since the casts of the shells of +mollusca are often excessively abundant in the rock, and the mudstone is +cavernous, resembling an impure, decalcified limestone. These Yeringian +mudstones of Victoria seem, therefore, to be the equivalent of the +calcareous shales met with in the Wenlock and Gotland Series in Europe; +a view entirely in accordance with the character of the remainder of +the fauna. One of the commonest of the Silurian ostracods is _Beyrichia +kloedeni_, a form having an extensive distribution in Europe. It occurs +in the Silurian mudstone of the Upper Yarra District. Other species of the +same genus are _B. wooriyallockensis_ (Fig. 112 A), distinguished from the +former by differences in the shape of the lobes and its longer valves; +also a form with narrow lobes, _B. kilmoriensis_; and the ornate _B. +maccoyiana_, var. _australis_. Of the smooth-valved forms, mention may be +made of _Bythocypris hollii_, _B. caudalis_ (Fig. 112 D), and the striking +form, _Macrocypris flexuosa_. Regarding the group of the _Primitiae_, of +which as many as thirteen species and varieties have been described from +the Lilydale Limestone, we may mention as common forms _P. reticristata_ +(Fig. 112 E) and _P. punctata_. This genus is distinguished by the +bean-shaped or purse-shaped carapace, with its well developed marginal +flange and mid-dorsal pit. Other genera which occur in our Silurians and +are of great interest on account of their distribution elsewhere. are +_Isochilina_, _Aparchites_, _Xestoleberis_, _Aechmina_, and _Argilloecia_. + +[Illustration: =Fig. 112--SILURIAN OSTRACODA.= + + A--Beyrichia wooriyallockensis, Chapm. Silurian (Yer.) Victoria + + B--Xestoleberis lilydalensis, Chapm. Silurian (Yer.) Victoria + + C--Argilloecia acuta, Jones and Kirkby. Silurian (Yer.) Victoria + + D--Bythocypris caudalis, Jones. Silurian (Yer.) Victoria + + E--Primitia reticristata, Jones. Silurian (Yer.) Victoria + +] + +The largest ostracod yet described from Australia, measuring more than a +quarter of an inch in length, occurs in the Upper Silurian of Cliftonwood, +near Yass, New South Wales. It belongs to the genus _Leperditia_ (_L. +shearsbii_), and is closely related to _L. marginata_, Keyserling sp.; +which occurs in strata of similar age in the Swedish and Russian Baltic +area. A limestone at Fifield, New South Wales, probably of Silurian age, +contains _Primitia_, _Kloedenia_, and _Beyrichia_. + + +=Devonian Ostracoda.--= + +The little _Primitia cuneus_ (Fig. 113 A) with a bean-shaped carapace and +median pit or depression occurs somewhat frequently in the Middle Devonian +Limestone of Buchan, Victoria. Another species, _Primitia yassensis_, is +found in the shaly rock of Narrengullen Greek, New South Wales. It is +probable that many other species of the group of the ostracoda remain to +be described from Australian Devonian rocks. + + +=Carboniferous Ostracoda.--= + +In Queensland a conspicuous little ostracod is _Beyrichia varicosa_ from +the Star Beds of Corner Creek. + + +=Carbopermian Ostracoda.--= + +In the Carbopermian of Cessnock, New South Wales, _Primitia dunii_ occurs; +and in that of Farley is found _Jonesina etheridgei_. From both these +localities _Leperditia prominens_ was also obtained. Another species from +New South Wales is _Entomis jonesi_ (Fig. 113 B), described from the Muree +Sandstone by de Koninck. + +[Illustration: =Fig. 113--UPPER PALAEOZOIC and MESOZOIC OSTRACODA.= + + A--Primitia cuneus, Chapm. Mid. Devonian. Victoria + + B--Entomis jonesi, de Kon. Carboniferous. New South Wales + + C--Synaphe mesozoica, Chapm. sp. Triassic. New South Wales + + D--Cythere lobulata, Chapm. Jurassic. West Australia + + E--Paradoxorhyncha foveolata, Chapm. Jurassic. West Australia + + F--Loxoconcha jurassica, Chapm. Jurassic. West Australia + + G--Cytheropteron australiense, Chapm. Jurassic. West Australia + +] + + +=Triassic Ostracoda.--= + +The Triassic (Wiannamatta Shales) of Grose Vale, New South Wales has +afforded a few specimens of ostracoda belonging to _Synaphe_ (_S. +mesozoica_, Fig. 113 C), _? Darwinula_, and _? Cytheridea_. + + +=Jurassic Ostracoda.--= + +The marine Jurassic strata of Western Australia at Geraldton, have yielded +a small but interesting series of ostracoda, largely of modern generic +types. The genera, which were found in a rubbly _Trigonia_-Limestone, are +_Cythere_, _Paradoxorhyncha_, _Loxoconcha_, and _Cytheropteron_. + +[Illustration: =Fig. 114--CAINOZOIC OSTRACODA.= + + A--Bairdia amygdaloides, G. S. Brady. Balcombian. Victoria + + B--Cythere clavigera, G. S. Brady. Balcombian. Victoria + + C--Cythere scabrocuneata, G. S. Brady. Balcombian. Victoria + + D--Cytherella punctata, G. S. Brady. Balcombian. Victoria + +] + + +=Cainozoic Ostracoda.--= + +The fossiliferous clays and calcareous sands of the southern Australian +Cainozoic beds often contain abundant remains of ostracoda. The moderately +shallow seas in which the fossiliferous clays, such as those of Balcombe's +Bay, were laid down, teemed with these minute bivalved Crustacea. All the +forms found in these beds are microscopic. They either belong to living +species, or to species closely allied to existing forms. Some of the more +prominent of the Balcombian species are _Cythere senticosa_, a form which +is now found living at Tenedos, and _C. clavigera_ (Fig. 114 B), with +the young form sometimes referred to as _C. militaris_, a species which +may still be dredged alive in Hobson's Bay. Other genera common in these +clays are _Bairdia_, with its broad, pear-shaped carapace, represented by +the still living _B. amygdaloides_ (Fig. 114 A). _Cytherella_, with its +compressed, subquadrate carapace, as seen in _C. punctata_ (Fig. 114 D), +a species having an elaborate series of muscle-spots, and which, like the +previous species, is found living in Australian seas; and _Macrocypris_, +with its slender, pointed, pear-shaped outline. + + +=Cirripedia: Their Habits and Structure.--= + +_CIRRIPEDIA OR BARNACLES._--These curious modifications of the higher +group of Crustacea (Eucrustacea) date back to Ordovician times. They +appear to have tried every possible condition of existence; and although +they are mostly of shallow water habits, some are found at the great +depth of 2,000 fathoms (over two miles). Those which secrete lime or +have calcareous shells, attach themselves to stones, pieces of wood, +shell-fish, crabs, corals and sea-weeds. Others are found embedded in the +thick skin of whales and dolphins, or in cavities which they have bored +in corals or shells of molluscs. Some are found parasitic in the stomachs +of crabs and lobsters, or within other cirripedes. They begin life, after +escaping from the egg, as a free-swimming, unsegmented larva ("nauplius" +stage), and before settling down, pass through the free-swimming, +segmented "cypris" stage, which represents the pupa condition, and in +which state they explore their surroundings in search of a suitable +resting place for their final change and fixed condition. Just before +this occurs, glands are developed in the pupa barnacle, which open into +the suckers of the first pair of appendages or antennae. When a suitable +place for fixation has been found, these glands pour out a secretion which +is not dissolved by water, and thus the barnacle is fixed head downwards +to its permanent position. The compound eyes of the "cypris" stage +disappear, and henceforth the barnacle is blind. The characteristic plates +covering the barnacle are now developed, and the six pairs of swimming +feet become the cirri or plumes, with which the barnacle, by incessant +waving, procures its food. In short, as remarked by one authority, it is +a crustacean "fixed by its head, and kicking the food into its mouth with +its legs." + +Cirripedes may be roughly divided into two groups, the Acorn Barnacles +and the Goose Barnacles. Although dissimilar in general appearance, they +pass through identical stages, and are closely related in most of their +essential characters. The latter forms are affixed by a chitinous stalk or +peduncle, whilst the acorn barnacles are more or less conical and affixed +by the base. + + +=Silurian Cirripedes.--= + +The stalked barnacles are probably the oldest group, being found as far +back as the Ordovician period. In Australia the genus _Turrilepas_ occurs +in Silurian rocks, _T. mitchelli_ (Fig. 115 A) being found at Bowning in +the Yass District of New South Wales. The isolated plume-like plates of +_T. yeringiae_ (Fig. 115 B) are not uncommon in the olive mudstone of the +Lilydale District in Victoria. + +[Illustration: =Fig. 115--FOSSIL CIRRIPEDIA.= + + A--Turrilepas mitchelli, Eth. fil. Silurian. New South Wales + + B--Turrilepas yeringiae, Chapm. Silurian. Victoria + + C--(?) Pollicipes aucklandicus, Hector sp. Cainozoic (Oamaru series). + New Zealand + +] + +[Illustration: =Fig. 116--LIVING AND FOSSIL CIRRIPEDES.= + + A--Lepas anatifera, L. Common Goose Barnacle. Living + + B--Lepas pritchardi, Hall. Cainozoic. Victoria + +] + + +=Cainozoic Lepadidae.--= + +The genus _Lepas_ (the modern goose barnacles) is represented by isolated +plates in the Cainozoic (Janjukian) limestones and marls of Waurn +Ponds, and Torquay near Geelong: it also occurs in a stratum of about +the same age, the nodule bed, at Muddy Creek, near Hamilton, Victoria +(_L. pritchardi_, Fig. 116). In New Zealand the gigantic cirripede, +_?Pollicipes aucklandicus_ (Fig. 115 C), occurs in the Motutapu beds. + + +=Cainozoic Balanidae.--= + +The Acorn Barnacles are represented in our Cainozoic shell marls and +clays by a species of _Balanus_ from the Janjukian of Torquay; whilst +two species of the genus occur in the Kalimnan beds at Beaumaris, Port +Phillip, in similar beds in the Hamilton District, and at the Gippsland +Lakes. + + +=Phyllocarida: Their Structure.--= + +A large and important group of the higher Crustacea, but confined to the +older rocks of Victoria, is the order _PHYLLOCARIDA_. This seems to form +a link between the Entomostraca, including the bivalved Ostracoda and the +well-known group of the lobsters, shrimps and crabs. The body of these +phyllocarids consists of five segments to the head, eight to the thorax, +and from two to eight to the abdomen. The portion usually preserved in +this group is the carapace, which covers the head and thorax, and although +often in one piece, is sometimes hinged, or otherwise articulated along +the back. In front of the carapace there is a moveable plate, the rostrum +or beak (Fig. 117). There are two pairs of antennae to the head, and the +animal is provided with a pair of stalked compound eyes. The thoracic +segments are furnished with soft leaf-like legs as in the Phyllopods. +The abdomen is formed of ring-like segments, and generally terminates +in a sharp tail-piece or telson, often furnished with lateral spines. +In many respects the ancient phyllocarids correspond with the living +genus _Nebalia_, which is found inhabiting the shallow waters of the +Mediterranean and elsewhere. + +[Illustration: =Fig. 117--Ceratiocaris papilio, Salter.= + +Silurian. Lanarkshire. + + (_After H. Woodward_) +] + +[Illustration: =Fig. 118--ORDOVICIAN PHYLLOCARIDS.= + + A--Rhinopterocaris maccoyi, Eth. fil. sp. L. Ordovician. Victoria + + B--Caryocaris angusta, Chapm. L. Ordovician. Victoria + + C--Saccocaris tetragona, Chapm. L. Ordovician. Victoria + +] + +[Illustration: =Fig. 119--SILURIAN PHYLLOCARIDS.= + + A--Ceratiocaris pritchardi, Chapm. Silurian. Victoria + + B--Ceratiocaris cf. murchisoni, Agassiz sp. Silurian. Victoria + + C--Ceratiocaris pinguis, Chapm. Silurian. Victoria + +] + + +=Ordovician Phyllocarids.--= + +Phyllocarids of the Lower Ordovician slates are referred to the genera +_Rhinopterocaris_, _Caryocaris_, _Saccocaris_ and _Hymenocaris_. The +first-named is the commonest type; and is found in slates of the +Lancefield, Bendigo and Castlemaine Series at the localities named, +as well as at Dromana. _Rhinopterocaris_ (Fig. 118 A) is readily +distinguished by its long--ovate outline, and this, together with its +wrinkled chitinous appearance makes it resemble the wing of a dipterous +insect. _Caryocaris_ (Fig. 118 B) is a smaller and narrower form which +occurs in the Victorian Lower Ordovician slates, as well as in ice-borne +blocks derived from the Ordovician, at Wynyard, in N.W. Tasmania. + + +=Silurian Phyllocarids.--= + +The chief type of Phyllocarid in the Silurian is _Ceratiocaris_ (Fig. +119). The carapace is typically ovate, straight on one edge, the dorsal, +and convexly curved on the other, the ventral. They resemble bean-pods in +outline, hence the name "pod-shrimps." Several species are known from the +Victorian shales, mudstones, and sandstones; the forms found in Australia +if complete would seldom attain five inches in length, whilst some British +species are known to reach the exceptional length of two feet. The long, +grooved and jointed telson is not uncommon in the sandstones of Melbourne +and Kilmore. Other genera described from Victoria are _Aptychopsis_ and +_Dithyrocaris_. + + +=Lower Cretaceous Crab.--= + +The earliest example of the _DECAPODA_ in the Australian rocks, so far +recorded, is the Lower Cretaceous _Prosopon etheridgei_ (Fig. 120 A) from +Queensland, which has affinities with some Jurassic and Neocomian crabs +found in Europe. Other crustacean remains of less decipherable nature +occur in this same deposit. + +[Illustration: =Fig. 120--FOSSIL CRABS and INSECTS.= + + A--Prosopon etheridgei, H. Woodw. L. Cretaceous. Queensland + + B--Ommatocarcinus corioensis, Cressw. sp. Cainozoic (Jan.) Vic. + + C--Harpactocarcinus tumidus, H. Woodw. Cainozoic (Oamaru). New Zealand + + D--Aeschna flindersensis, H. Woodw. L. Cretaceous. Queensland + + E--Ephemera culleni, Eth. fil. and Olliff. Cainozoic (Deep Leads). New + South Wales + +] + + +=Cainozoic Crabs.--= + +Of the Cainozoic decapod Crustacea there is a Victorian species of a +stalk-eyed crab, _Ommatocarcinus corioensis_ (Fig. 120 B), found in the +marls of Curlewis and Port Campbell, and probably of Janjukian age. +Various portions of similar Crustacea, consisting of claws and fragmentary +carapaces, are found from time to time in the Victorian clays and +limestones of Balcombian and Janjukian ages, but they are insufficient +for identification. A carapace of one of the Oxystomata (with rounded +cephalo-thorax and non-salient frontal region) has occurred in the +Kalimnan marl of the Beaumaris Cliffs, Port Phillip. It is closely allied +to a crab now found in Hobson's Bay and generally along the Victorian +coast. + +Remains of a shore-crab (Fam. Cancridae) are found at three localities, +in the Oamaru Series, in New Zealand; near Brighton, in Nelson and at +Wharekuri in the Waitaki Valley. It has been described under the name of +_Harpactocarcinus tumidus_ (Fig. 120 C), a genus of the Cyclometopa or +"bow crabs." + + +=Pleistocene Lobster.--= + +Numerous remains of a lobster, _Thalassina emerii_ (see _antea_, Fig. 20), +supposed to be of Pleistocene age, occur in nodules found on Queensland +and North Australian (Port Darwin) beaches. + + +=Eurypterids in the Silurian.--= + +The order _EURYPTERIDA_ comprises an extinct group of Crustacea closely +allied to the modern King-crab (_Limulus_). The body was covered with a +thin chitinous skeleton, ornamented with regular scale-like markings. This +group is represented in Victorian rocks by the remains of _Pterygotus_ +("Sea-scorpions"), animals which often attained a length of six feet. +_Pterygotus_ (see Fig. 121 A) had the fore part of the body fused, forming +the cephalo-thorax, which was furnished with anterior, marginal facetted +eyes and central ocelli or smaller simple ones. To the ventral surface +of the body were attached six pairs of appendages. The first pair are +modified antennae with pincer-like terminations, used for prehensile +purposes. Then come four pairs of slender walking feet. The sixth pair of +appendages is in the form of powerful swimming feet or paddles, at the +bases of which are the comb-like jaws. The abdomen consists of thirteen +joints, the last of which, the telson, is spatulate and posteriorly +pointed. Fragments of a tolerably large species of _Pterygotus_ occur in +the Silurian shales of South Yarra, Melbourne, Victoria. It was probably +about 18 inches in length when complete. Of this form, known as _P. +australis_ (Fig. 121 B), portions of the chelate (clawed) appendages, and +parts of the abdominal segments have been found from time to time, but no +complete fossil has yet been discovered. + +[Illustration: =Fig. 121--SILURIAN EURYPTERIDS.= + + A--Pterygotus osiliensis, Schmidt. I. of Oesel. (_After Schmidt_) + + B--Pterygotus australis, McCoy. Part of a body-segment. Silurian + (Melb.) Victoria + +] + + +=Jurassic Insects.--= + +Of the group of the _INSECTA_, the Ipswich Coal measures (Jurassic) +of Queensland have yielded an interesting buprestid beetle +(_Mesostigmodera_), whilst beds of the same age in New South Wales contain +the remains of a probable _Cicada_, associated with leaves of the fern +_Taeniopteris_. + + +=Lower Cretaceous Dragon-fly.--= + +From the Lower Cretaceous of the Flinders River district, Queensland, +there has been obtained a fossil dragon-fly, _Aeschna flindersensis_ (Fig. +120 D). + + +=Cainozoic Insects.--= + +Certain Cainozoic beds of New South Wales, of the age of the Deep-leads +of Victoria, and probably equivalent to the Kalimnan terrestrial series, +contain a species of _Cydnus_, a bug-like insect belonging to the order +Rhynchota; and there are in the same series a Midge (_Chironomus_), +a Day-fly (_Ephemera_, Fig. 120 E) and several beetles (? _Lagria_, +_Palaeolycus_, _Cyphon_ and _Oxytelus_). The occurrence of these insects +of the Deep-leads helps to complete the landscape picture of those +far-off Lower Pliocene times, when the old river systems brought down +large contributions of vegetable waste from higher lands, in the form of +twigs with leaves and fruits; with occasional evidences of the rich and +varied fauna of insect life which was especially promoted in the damp and +vegetative areas of the lower lands. + + +COMMON OR CHARACTERISTIC SPECIES OF THE FOREGOING CHAPTER. + + +TRILOBITES. + + _Ptychoparia howchini_, Eth. fil. Lower Cambrian: South Australia. + + _Dolichometopus tatei_, H. Woodward. Lower Cambrian: South Australia. + + _Olenellus browni_, Eth. fil. Lower Cambrian: Northern Territory. + + _Agnostus australiensis_, Chapm. Upper Cambrian: Victoria. + + _Ptychoparia thielei_, Chapm. Upper Cambrian: Victoria. + + _Dikellocephalus florentinensis_, Eth. fil. Upper Cambrian: Tasmania. + + _Dinesus ida_, Eth. fil. Lower Ordovician: Victoria. + + _Asaphus illarensis_, Eth. fil. Ordovician: Central S. Australia. + + _Ampyx parvulus_, Forbes, var. _jikaensis_, Chapm. Silurian + (Melbournian): Victoria. + + _Illaenus jutsoni_, Chapm. Silurian (Melbournian): Victoria. + + _Proetus euryceps_, McCoy. Silurian: Victoria. + + _Cyphaspis spryi_, Gregory. Silurian (Melbournian): Victoria. + + _Bronteus enormis_, Eth. fil. Silurian (Yeringian): Victoria. + + _Lichas australis_, McCoy. Silurian (Yeringian): Victoria. + + _Odontopleura jenkinsi_, Eth. fil. Silurian: New South Wales. Silurian + (Yeringian): Victoria. + + _Encrinurus punctatus_, Brunnich sp. Silurian: New South Wales. + Silurian (Yeringian): Victoria. + + _Encrinurus (Cromus) murchisoni_, de Koninck. Silurian: New South + Wales. + + _Encrinurus (Cromus) spryi_, Chapm. Silurian (Melbournian): Victoria. + + _Calymene blumenbachii_, Brongn. Silurian (Wangapeka Series): New + Zealand. + + _Homalonotus expansus_, Hector. Silurian (Wangapeka Series): New + Zealand. + + _Homalonotus knightii_, König. Silurian (Wangapeka Series): New + Zealand. + + _Homalonotus harrisoni_, McCoy. Silurian (Melbournian): Victoria. + + _Homalonotus vomer_, Chapm. Silurian: Victoria. + + _Cheirurus insignis_, Beyrich. Silurian: New South Wales. + + _Phacops sweeti_, Eth. fil. and Mitch. Silurian: New South Wales. + Silurian (Yeringian): Victoria. + + _Phacops serratus_, Foerste. Silurian (Yeringian): Victoria. Silurian: + New South Wales. + + _Dalmanites meridianus_, Eth. fil. and Mitch, sp. Silurian: New South + Wales, Victoria and Tasmania. + + _Cheirurus_ sp. Middle Devonian: Victoria. + + _Proetus_ sp. Devonian: Western Australia. + + _Phillipsia seminifera_, Phillips. Carbopermian: New South Wales. + + _Phillipsia grandis_, Eth. fil. Carbopermian: W. Australia and + Queensland. + + _Griffithides eichwaldi_, Waldheim. Carbopermian: New South Wales and + Queensland. + + _Brachymetopus strzelecki_, McCoy. Carbopermian: New South Wales. + + +PHYLLOPODA. + + _Leaia mitchelli_, Eth. fil. Upper Carboniferous: New South Wales. + + _Estheria coghlani_, Cox. Trias: New South Wales. + + _Estheria minuta_, Alberti sp. Trias: New Zealand. + + _Estheria mangaliensis_, Jones. Jurassic: Queensland. + + +OSTRACODA. + + _Leperditia_ sp. Lower Cambrian: S. Australia. + + _Beyrichia kloedeni_, McCoy. Silurian (Yeringian): Victoria. + + _Beyrichia wooriyallockensis_, Chapm. Silurian (Yeringian): Victoria. + + _Beyrichia maccoyiana_, Jones, var. _australis_, Chapm. Silurian: + (Yeringian): Victoria. + + _Bythocypris hollii_, Jones. Silurian (Yeringian): Victoria. + + _Macrocypris flexuosa_, Chapm. Silurian (Yeringian) Victoria. + + _Primitia reticristata_, Jones. Silurian (Yeringian): Victoria. + + _Leperditia shearsbii_, Chapm. Silurian: New South Wales. + + _Primitia cuneus_, Chapm. Middle Devonian: Victoria. + + _Beyrichia varicosa_, T. R. Jones. Carboniferous: Queensland. + + _Primitia dunii_, Chapm. Carbopermian: New South Wales. + + _Jonesina etheridgei_, Chapm. Carbopermian: New South Wales. + + _Entomis jonesi_, de Koninck. Carbopermian: New South Wales. + + _Synaphe mesozoica_, Chapm. sp. Trias: New South Wales. + + _Cythere lobulata_, Chapm. Jurassic: W. Australia. + + _Paradoxorhyncha foveolata_, Chapm. Jurassic: W. Australia. + + _Loxoconcha jurassica_, Chapm. Jurassic: W. Australia. + + _Cytheropteron australiense_, Chapm. Jurassic: W. Australia. + + _Bairdia amygdaloides_, Brady. Cainozoic and living: Victoria. + + _Cythere senticosa_, Baird. Cainozoic. Also living: Victoria. + + _Cythere clavigera_, G. S. Brady. Cainozoic and living: Victoria. + + _Cytherella punctata_, G. S. Brady. Cainozoic and living: Victoria. + + _Cytherella pulchra_, G. S. Brady. Cainozoic and living: Victoria. + + +CIRRIPEDIA. + + _Turrilepas mitchelli_, Eth. fil. Silurian: New South Wales. + + _Turrilepas yeringiae_, Chapm. Silurian (Yeringian): Victoria. + + _Lepas pritchardi_, Hall. Cainozoic (Janjukian): Victoria. + + _(?) Pollicipes aucklandicus_, Hector sp. Cainozoic (Oamaru Series): + New Zealand. + + _Balanus_ sp. Cainozoic (Janjukian and Kalimnan): Victoria. + + +PHYLLOCARIDA. + + _Rhinopterocaris maccoyi_, Eth. fil. sp. Lower Ordovician: Victoria. + + _Hymenocaris hepburnensis_, Chapm. L. Ordovician: Victoria. + + _Caryocaris marri_, Jones and Woodw. L. Ordovician: Victoria and + Tasmania. + + _Caryocaris angusta_, Chapm. L. Ordovician: Victoria. + + _Saccocaris tetragona_, Chapm. L. Ordovician: Victoria. + + _Ceratiocaris_ cf. _murchisoni_, Agassiz sp. Silurian: Victoria. + + _Ceratiocaris pinguis_, Chapm. Silurian (Melbournian): Victoria. + + _Ceratiocaris pritchardi_, Chapm. Silurian: Victoria. + + _Aptychopsis victoriae_, Chapm. Silurian (Melbournian): Victoria. + + _Dithyrocaris praecox_, Chapm. Silurian (Melbournian): Victoria. + + +DECAPODA. + + _Prosopon etheridgei_, H. Woodw. Lower Cretaceous: Queensland. + + _Ommatocarcinus corioensis_, Cresswell sp. Cainozoic (Janjukian): + Victoria. + + _Ebalia_ sp. Cainozoic (Kalimnan): Victoria. + + _Harpactocarcinus tumidus_, H. Woodw. Cainozoic (Oamaru Series): New + Zealand. + + _Thalassina emerii_, Bell. (?) Pleistocene: Queensland and Northern + Territory. + + +EURYPTERIDA. + + _Pterygotus australis_, McCoy. Silurian (Melbournian): Victoria. + + +INSECTA. + + _Mesostigmodera typica_, Etheridge fil. and Olliff. Jurassic: + Queensland. + + _(?) Cicada lowei_, Etheridge fil. and Olliff. Jurassic: New South + Wales. + + _Aeschna flindersensis_, H. Woodward. Lower Cretaceous: Queensland. + + _Chironomus venerabilis_, Eth. fil. and Oll. Cainozoic: New South + Wales. + + _Ephemera culleni_, Eth. fil. and Oll. Cainozoic: New South Wales. + + _Palaeolycus problematicum_, Eth. fil. and Oll. Cainozoic: New South + Wales. + + * * * * * + + +LITERATURE. + + +TRILOBITES. + + McCoy, F. Prod. Pal. Vict., Dec. III. 1876, pp. 13-20, pls. XXII. + and XXIII. (Silurian). Hector, J. Trans. N.Z. Inst., vol. IX. + 1877, p. 602, pl. XXVII. (_Homalonotus_). Woodward, H. Geol. + Mag., Dec. III. vol. I. 1884, pp. 342-344, pl. XI. (Cambrian). + Mitchell, J. Proc. Linn. Soc. New South Wales, vol. II. 1888, + pp. 435-440, pl. XI. (Silurian). Foerste, A. F. Bull. Sci. Lab. + Denison Univ., vol. III. pt. V. 1888, pp. 122-128, pl. XIII. + Etheridge, R. jnr. Proc. Linn. Soc. New South Wales, vol. V. + pp. 501-504, pl. XVIII. (_Bronteus_). Idem, Parl. Papers, Leg. + Assemb. S.A., vol. I. No. 23, 1892; ibid., vol. 2, No. 52, 1893 + (_Asaphus_). Id., Geol. Queensland, 1892, pp. 214-216, pls. + VII. VIII. and XLIV. (Carboniferous). Id., Proc. R. Soc. Vict., + vol. VI. (N.S.), 1894, pp. 189-194, pl. XI. (_Bronteus_). Id., + ibid, vol. VIII. (N.S.), 1896, pp. 56, 57, pl. I. (_Dinesus_). + Id., Rec. Austr. Mus., vol. V. No. 2, 1904, pp. 98-101, pl. X. + (Cambrian). Id., Trans. R. Soc. S. Austr., vol. XXII. 1898, pp. + 1-3, pl. IV. (Cambrian). Etheridge, R. jnr. and Mitchell, J. + Proc. Linn. Soc. New South Wales, vol. VI. 1892, pp. 311-320, pl. + XXV.; ibid., vol. VIII. 1894, pp. 169-178, pls. VI. VII.; ibid., + vol. X. 1896, pp. 486-511, pls. XXXVIII.-XL.; ibid., vol. XXI. + 1897, pp. 694-721, pls. L.-LV. Tate, R. Rep. Horn Exped., 1896, + Part 3, Palaeontology, pp. 111, 112, pl. III. De Koninck, L. G. + Mem. Geol. Surv. New South Wales, Pal. No. 6, 1898, pp. 36-47 pl. + I. (Silurian); pp. 276-281, pl. XXIV. (Carboniferous). Gregory, + J. W. Proc. R. Soc. Vict., vol. XIII. (N.S.) pt. II, 1901, pp. + 179-182, pl. XXII. (_Cyphaspis_). Ibid., vol. XV. (N.S.) pt. + II. 1903, pp. 154-156, pl. XXVI. (_Dinesus_ and _Notasaphus_.) + Chapman, F. Proc. R. Soc. Vict., vol. XXIII. (N.S.), pt. II. + 1910, pp. 314-322, pls. LVIII. and LIX. (Cambrian). Ibid., + vol. XXIV. (N.S.) pt. II. 1912, pp. 293-300, pls. LXI.-LXIII. + (Silurian). + + +PHYLLOPODA. + + Cox, J. C. Proc. Linn. Soc. New South Wales, vol. V., pt. 3, 1881, p. + 276 (_Estheria_). Etheridge, R. jnr. ibid., vol. VII. 1893, pp. + 307-310, text fig. (_Leaia_). Idem, Mem. Geol. Surv. New South + Wales, Pal. No. 1, 1888, pp. 6-8, pl. I. (_Estheria_). + + +OSTRACODA. + + Brady, G. S. in Etheridge, jnr. Geol. Mag., 1876, p. 334 (Cainozoic). + De Koninck, L. G. Mem. Geol. Surv. New South Wales, Pal. No. + 6, 1898, pp. 33, 36 (Silurian); ibid., pp. 275, 276, pl. XXIV. + (Carboniferous). Chapman, F. Proc. R. Soc. Vict., vol. XVI. + (N.S.), pt. II. 1904, pp. 199-204, pl. XXIII. (Jurassic). + Idem, ibid., vol. XXII. (N.S.), pt. I. 1909, pp. 1-5, pl. I. + (_Leperditia_). Idem, Rec. Geol. Surv. New South Wales, vol. + VIII. pt. 4, 1909, pp. 1-3, pl. LIV. (Triassic). Idem, Rec. + Geol. Surv. Vict., vol. III. pt. 2, 1912, p. 221, pl. XXXVI. + (_Primitia_). Idem, Proc. R. Soc. Vict., vol. XV. (N.S.), pt. + II. 1903, pp. 109-113, pl. XVI. (_Beyrichia_). Ibid., vol. XVII. + (N.S.) pt. I. 1904, pp. 299-312, pls. XIII.-XVII. (Silurian). + + +CIRRIPEDIA. + + Etheridge, R. jnr. Geol. Mag., Dec. III. vol. VII. 1890, pp. 337, 338, + pl. XI. (_Turrilepas_). Hall, T.S. Proc. R. Soc. Vict., vol. + XV. (N.S.) pt. I. 1902, pp. 83, 84, pl. XI. (_Lepas_). Benham, + W. B. Geol. Mag., Dec. IV. vol. X. pp. 110-119, pls. IX. X. (_? + Pollicipes_). Chapman, F. Proc. R. Soc. Vict. vol. XXII. (N.S.) + pt. II. 1910, pp. 105-197, pls. XXVIII. XXIX. (_Turrilepas_). + + +PHYLLOCARIDA. + + Etheridge, R. jnr. Rec. Geol. Surv. New South Wales, vol. III. pt. I. + 1894, pp. 5-8, pl. IV. (Ordovician). Chapman, F. Proc. R. Soc. + Vict. vol. XV. (N.S.), pt. II. 1903, pp. 113-117, pl. XVIII. + (Ordovician); ibid., vol. XVII. (N.S.) pt. I. 1904, pp. 312-315, + pl. XVII.; ibid., vol. XXII. (N.S.), pt. II. 1910, pp. 107-110, + pl. XXVIII. (Silurian). Idem, Rec. Geol. Surv. Vict., vol. III. + pt. 2, 1912, pp. 212, 213, pls. XVII. XVIII. (Ordovician). + + +DECAPODA. + + Bell, T. Proc. Geol. Soc. Lond., vol. I. 1845, pp. 93, 94. Text-fig. + (_Thalassina_). Woodward, H. Quart. Journ. Geol. Soc., vol. + XXXII. 1876, pp. 51-53, pl. VII. (_Harpactocarcinus_). Idem., + Proc. Linn. Soc. New South Wales, vol. VII. (2), pt. 2, 1892, pp. + 301-304 pl. IV. (_Prosopon_). + + Hall, T. S. Proc. R. Soc. Vict., vol. XVII. (N.S.) pt. II. 1905, pp. + 356-360, pl. XXIII. (_Ommatocarcinus_). + + +EURYPTERIDA. + + McCoy, F. Geol. Mag. Dec. IV. vol. VI. 1899, pp. 193, 194, text fig. + (_Pterygotus_). + + +INSECTA. + + Woodward, H. Geol. Mag. Dec. III. vol. I. 1884, pp. 337-339, pl. XI. + (_Aeschna_). Etheridge, R. jnr. and Olliff, A. S. Mem. Geol. + Surv. New South Wales, Pal. No. 7, 1890 (Mesozoic and Cainozoic). + + + + +CHAPTER XII. + +FOSSIL FISHES, AMPHIBIANS, REPTILES, BIRDS, AND MAMMALS. + + +=Vertebrates.--= + +The above-named classes of animals are distinguished from those previously +dealt with, by the presence of a vertebral column. The vertebral axis +may be either cartilaginous as in some fishes, or bony as in the greater +number of animals belonging to this sub-kingdom. + + +=Chordata.--= + +_LINKS BETWEEN THE INVERTEBRATES AND FISHES._--The curious little +ascidians or "sea-squirts," belonging to the group Tunicata, are held +by some authorities to be the degenerate descendants of a free-swimming +animal having a complete notochord and nerve-tube, structures which +are now only seen in the tails of their tadpole-like larvae. The fully +developed tunicate is generally sessile and provided with a thick outer +coat (tunic) and muscular inner lining. This outer coat in some forms, as +_Leptoclinum_, is strengthened with tiny calcareous spicules, and these +are sometimes found in the fossil state in Cainozoic clays, as well as +in some of the calcareous deep-sea oozes. The little stellate spicules of +_Leptoclinum_ are abundant in the Balcombian clays of Mornington, Victoria. + +Another primitive form with a notochord is the Lancelet, but this, having +no hard parts, is not found in the fossil state. + + +=Primitive Types of Fishes.--= + +_FISHES._--The remains of fishes are naturally more abundant in the +fossil condition, owing to their aquatic habits, than those of other +vertebrates. The earliest fishes were probably entirely cartilaginous, +and some have left only a mere trace or impression on the rocks in which +they were embedded. These primitive fishes have no lower jaw, and are +without paired limbs. They are sometimes placed in a class by themselves +(_AGNATHA_). The orders of this primitive fish series as represented in +Australasia are the Osteostraci ("bony shells"), of which the remains of +the _Cephalaspis_-like head-shield of _Thyestes_ has been found in the +Silurian of N.E. Gippsland, Victoria (Fig. 122); and the Antiarchi, with +its many-plated cuirass, armoured body-appendages, internal bony tissue, +and coarsely tuberculated exterior, as seen in _Asterolepis australis_, +a fossil occasionally found in the Middle Devonian Limestone of Buchan, +Gippsland. + + +=True Fishes.--Devonian.--= + +Of the true fishes (Pisces), the Elasmobranchii ("slit-gills"), a +sub-class to which the modern sharks belong, are represented in the +Devonian series by the paired spines of a form resembling _Climatius_, +found both in Victoria and New South Wales. Remains of Dipnoi +("double-breather" or lung-fishes) occur in the Devonian of Barker Gorge, +Western Australia, represented by a new species allied to _Coccosteus_ +("berry-bone" fish); and in a bed of the same age at the Murrumbidgee +River, New South Wales by the cranial buckler of _Ganorhynchus süssmilchi_. + +[Illustration: =Fig. 122--Incomplete Head-Shield= of Thyestes magnificus, +Chapm. From the Silurian (Yeringian) of Wombat Creek, N.E. Gippsland. 4/5 +nat. size] + +[Illustration: =Fig. 123= + +=Gyracanthides murrayi=, + +A. S. Woodw. + +L. Carboniferous. Mansfield, Victoria. + +(Restoration). + +About 1/12 nat. size] + +[Illustration: =Fig. 124--TEETH and SCALES of PALAEOZOIC and MESOZOIC +FISHES.= + + A--Strepsodus decipiens, A. S. Woodw. L. Carboniferous. Victoria + + B--Elonichthys sweeti, A. S. Woodw. L. Carboniferous. Victoria + + C--Corax australis, Chapm. L. Cretaceous. Queensland + + D--Belouostomus sweeti, Eth. fil. and Woodw. L. Cretaceous. Q. + +] + + +=Carboniferous Fishes.--= + +The Lower Carboniferous sandstone of Burnt Creek and other localities near +Mansfield, Victoria, contains an abundant fish fauna, associated with +stems of _Lepidodendron_. The slabs of sandstone are often ripple-marked +and show signs of tracks and castings of shore-living animals. These +deposits were probably laid down in shallow water at the shore margin +or in salt lagoons or brackish areas skirting the coast, into which +at intervals the remains of the giant lycopods were drifted. The more +important of these fish remains are Elasmobranchs, as _Gyracanthides +murrayi_ (Fig. 123) and _Acanthodes australis_; the Dipnoan, _Ctenodus +breviceps_; a Rhizodont or fringe-finned ganoid, _Strepsodus decipiens_ +(Fig. 124 A); and a genus related to _Palaeoniscus_, _Elonichthys_ +(_E. sweeti_, Fig. 124 B, and _E. gibbus_). The defence spines of +_Gyracanthides_ are fairly abundant in the sandstones; whilst on some +slabs the large enamelled scales of _Strepsodus_ are equally conspicuous. + +From the sandstones of the same age, Lower Carboniferous, in the Grampians +of Western Victoria, some small but well-preserved spines belonging to the +genus _Physonemus_ have been found associated with a new variety of the +well-known European Carboniferous brachiopod, _Lingula squamiformis_ (var. +_borungensis_). + + +=Carbopermian Fishes.--= + +In the Carbopermian (Gympie Beds) of the Rockhampton District, Queensland, +a tooth of a Cochliodont ("snail tooth") occurs, which has been doubtfully +referred to the genus _Deltodus_ (? _D. australis_). The Cochliodontidae +show dentition remarkably like that of the _Cestracion_ or Port Jackson +Shark. Another tooth having the same family relationship has been +referred to _Tomodus ? convexus_, Agassiz; this is from the Carbopermian +of the Port Stephen district of New South Wales. From the Newcastle Coal +Measures in New South Wales a _Palaeoniscus_-like fish, _Urosthenes +australis_ has been described. + +Carbopermian fish remains are rare in Western Australia. They comprise +a wrinkled tooth of _Edestus_ (_E. davisii_) from the Gascoyne River, +belonging to a fish closely related to the Port Jackson shark; and a +cochliodont, _Poecilodus_ (_P. jonesi_, Ag.) from the Kimberley district. + + +=Triassic Fishes.--= + +Fossil fishes are important and numerous in Australian Triassic beds, +especially in New South Wales. At the base of the Hawkesbury or close +of the Narrabeen series, the railway ballast quarry near Gosford has +yielded an extensive and extremely interesting collection. Near the floor +of the quarry there is a band of sandy shale and laminated sandstone 5 +feet 9 inches in thickness, and this contains the following genera:--A +dipnoan, _Gosfordia_; and the following ganoids or enamelled scale +fishes--_Myriolepis_, _Apateolepis_, _Dictyopyge_, _Belonorhynchus_, +_Semionotus_, _Pristisomus_ (see _antea_, Fig. 18), _Cleithrolepis_ (Fig. +125), _Pholidophorus_ and ? _Peltopleurus_. + + +=Upper Triassic Fishes.--= + +In the middle of the Wianamatta or Upper Trias Series at St. Peter's, +near Sydney, which contains a fauna described as slightly older in aspect +than that of Gosford, having Carbopermian affinities, there occur in +the hard shale or clay stone the genera _Pleuracanthus_ (a Palaeozoic +shark); _Sagenodus_ (a dipnoan related to _Ctenodus_ of the Victorian +Carboniferous); and the following ganoids,--_Palaeoniscus_, _Elonichthys_, +_Myriolepis_, _Elpisopholis_, _Platysomus_ and _Acentrophorus_. From the +soft shales were obtained _Palaeoniscus_, _Semionotus_, _Cleithrolepis_ +and _Pholidophorus_; an assemblage of genera somewhat comparable with the +Gosford fauna. + +[Illustration: =Fig. 125--Cleithrolepis granulatus, Egerton.= + +Triassic (Hawkesbury Series). Gosford, New South Wales. 3/4 nat. size. + + (_After Smith Woodward_.) +] + + +=Lower Mesozoic Fishes.--= + +From the Lower Mesozoic sandstone (?Triassic) of Tasmania, two species of +_Acrolepis_ have been described, viz., _A. hamiltoni_ and _A. tasmanicus_. +The former occurs in the thick bed of sandstone, of nearly 1,000 feet, +at Knocklofty; the latter species in the sandstone with _Vertebraria_ +conformably overlying the Carbopermian at Tinderbox Bay. + +[Illustration: =Fig. 126--REMAINS of JURASSIC and OTHER VERTEBRATES.= + + 1--Ceratodus avus, A. S. Woodw. Left splenial with lower tooth. Cape + Paterson, Victoria. About 1/3 nat. size + + 2--Ceratodus forsteri, Krefft. Left lower tooth. Living. Queensland. + About 1/3 nat. size + + 3--Phalangeal of Carnivorous Dinosaur. Cape Paterson. About 1/3 nat. + size + + 4--Phalangeal of Megalosaurian. Wealden, Sussex, England. 1/4 nat. size + +] + +=Jurassic Fishes.--= + +The Jurassic beds of Victoria contain three genera. _Psilichthys selwyni_, +a doubtful palaeoniscid was described from Carapook, Co. Dundas; whilst +_Leptolepis_, a genus found in the Trias of New South Wales and the Lias +and Oolite of Europe, is represented by _L. crassicauda_ from Casterton, +associated with the typical Jurassic fern, _Taeniopteris_. In the Jurassic +beds of South Gippsland, at Cape Paterson, an interesting splenial tooth +of the mudfish, _Ceratodus_, was found, named _C. avus_ (Fig. 126). Since +then, in a bore-core from Kirrak near the same place a fish scale was +discovered (Fig. 127) which, by its shape, size and structure seems to +differ in no way from the living lung-fish of Queensland (Fig. 128). It +is reasonable to infer that tooth and scale belong to the same species; +and in view of the close relationship of the tooth with that of the living +mudfish, rather than with that of the _Ceratodus_ found fossil in the +Mesozoic of Europe, it may be referred to _Neoceratodus_, in which genus +the living species is now placed. + +[Illustration: =Fig. 127--Scale of Ceratodus (Neoceratodus)= (?)avus, A. +S. Woodw. Jurassic. Kirrak, S. Gippsland, Victoria. About nat. size] + +[Illustration: =Fig. 128--The Queensland Lung-Fish= + +or Barramunda (Neoceratodus forsteri). About 1/12th. nat. size + + (_After Lydekker, in Warne's Natural History_) +] + +[Illustration: =Fig. 129--Leptolepis gregarius=, A. S. Woodw. + +Talbragar Series, Jurassic. Talbragar River, New South Wales 1/2 nat. +size] + +From the Jurassic beds (Talbragar Series) of New South Wales, an +interesting collection of ganoid fishes has been described, comprising +_Coccolepis australis_, _Aphnelepis australis_, _Aetheolepis mirabilis_, +_Archaeomaene tenuis_, _A. robustus_, _Leptolepis talbragarensis_, _L. +lowei_ and _L. gregarina_ (Fig. 129). + + +=Lower Cretaceous Fishes.--= + +Fish remains are fairly abundant in the Lower Cretaceous of Queensland. +They comprise both the sharks and the ganoids. Of the sharks, a specimen, +showing seven conjoined vertebrae has been named _Lamna daviesii_, from +the Richmond Downs, Flinders River district; and a tooth referred to +_Lamna appendiculatus_, Agassiz, from Kamileroy, Leichhardt River, N.W. +Queensland. The typical Cretaceous genus _Corax_ is represented by a +small tooth named _C. australis_ (Fig. 124 C), from the Hamilton River, +Queensland, and which closely approaches the tooth of _Corax affinis_, +Agassiz, from the Upper Cretaceous of Europe. Of the ganoid fishes two +genera, both members of the family _Aspidorhynchidae_, have been found +in Queensland. _Aspidorhynchus_ sp. and _Belonostomus sweeti_ (Fig. 124 +D) have both occurred at Hughenden, Flinders River district. The former +genus has a slender body and produced rostrum; in Europe it is more +characteristic of Jurassic strata. _Belonostomus_ ranges from the Upper +Oolite, Bavaria, to the Upper Cretaceous in other parts of the world. +Remains of a species of _Portheus_, one of the predaceous fishes which +lived in the Cretaceous period, consisting of a portion of the cranium +with the anterior part of the jaws, has been obtained from the Rolling +Downs Formation (Lower Cretaceous) near Hughenden, Queensland. + + +=Cretaceous Fishes, New Zealand.--= + +[Illustration: =Fig. 130--CRETACEOUS and CAINOZOIC FISH-TEETH.= + + A--Notidanus marginalis, Davis. Cainozoic. New Zealand + + B--Callorhynchus hectori, Newton. Cainozoic. New Zealand + + C--Oxyrhina hastalis, Ag. Cainozoic. Victoria + + D--Lamna apiculata, Ag. Cainozoic. Victoria + + E--Carcharodon auriculatus, Blainv. sp. Cainozoic. Victoria + + F--Sargus laticonus, Davis. Cainozoic. New Zealand + +] + +The Cretaceous beds of New Zealand are grouped in ascending order as the +Waipara Greensands, the Amuri Limestone and the Weka Pass Stone. In the +Waipara beds occur the teeth of _Notidanus marginalis_ (Fig. 130 A), and +_N. dentatus_. In the Amuri Limestone _N. dentatus_ is again found, as +well as the genus _Lamna_, represented by _L. compressa_, Ag. (originally +described as _L. marginalis_, Davis), _L. carinata_ and _L. hectori_. Two +forms of "Elephant fish" are represented by their dental plates, namely +_Callorhynchus hectori_ (Fig. 130 B) and _Ischyodus thurmanni_, Pictet and +Campiche (recorded as _I. brevirostris_, Ag.). + + +=Cainozoic Fishes.--= + +Fish remains principally consisting of teeth, are common fossils in the +Cainozoic beds of southern Australia, particularly in Victoria, and also +in New Zealand. + + +=Balcombian Series, Southern Australia.--= + +The Balcombian beds as seen at Mornington and in the Lower Beds at Muddy +Creek, Hamilton, contain the teeth of sharks as _Odontaspis contortidens_, +_Lamna crassidens_, _L. apiculata_, _Oxyrhina hastalis_ (rarely), _O. +minuta_, _Carcharodon megalodon_, and _C. robustus_. + + +=Janjukian.--= + +The Janjukian Series (Miocene), represented at Torquay, Waurn Ponds and +Table Cape, contains an abundant fish fauna, including amongst sharks, +_Cestracion cainozoicus_, _Asteracanthus eocaenicus_, _Galeocerdo davisi_, +_Carcharoides totuserratus_, _Odontaspis contortidens_, _O. incurva_, +_O. cuspidata_, _Lamna crassidens_, _L. apiculata_ (Fig. 130 D), _L. +compressa_, _L. bronni_, _Oxyrhina hastalis_ (occasional) (Fig. 130 C), +_O. desori_, _O. retroflexa_, _O. minuta_, _Carcharodon auriculatus_ (Fig. +130 E), _C. megalodon_ and _C. robustus_. A species of chimaeroid or +Elephant fish is represented by a left mandibular tooth named _Ischyodus +mortoni_, from the Table Cape Beds, Tasmania. + +The Corio Bay series contains teeth of _Acanthias geelongensis_, _Sphyrna +prisca_, _Odontaspis contortidens_, _O. attenuata_, _Oxyrhina minuta_, +_Carcharodon megalodon_, amongst sharks; whilst the spine of a Porcupine +Fish, _Diodon connewarrensis_ has been obtained from the clays of Lake +Connewarre, Victoria. + + +=Kalimnan.--= + +[Illustration: =Fig. 131--CAINOZOIC FISH REMAINS.= + + A--Carcharoides tenuidens, Chapm. Cainozoic (Janj.) Victoria + + B--Odontaspis contortidens. Agassiz. Cainozoic (Kal.) Victoria + + C--Galeocerdo latidens, Agassiz. Cainozoic (Kal.) Victoria + + D--Myliobatis morrabbinensis, Chapm. and Pritch. Cainozoic (Kal.) + Victoria + + E--Labrodon confertidens. Chapm. and Pritch. Cainozoic (Kal.) Vict. + + F--Diodon formosus, Chapm. and Pritch. Cainozoic (Kal.) Vict. + +] + +The Kalimnan Series is also prolific in the remains of fishes, the +principal localities being Beaumaris and Grange Burn, Hamilton. Amongst +the sharks there found are, _Notidanus jenningsi_ (related to the Indian +Grey Shark), _Cestracion cainozoicus_ (related to the Port Jackson +Shark), _Asteracanthus eocaenicus_, _Galeocerdo davisi_, _G. latidens_ +(Fig. 131 C), _G. aduncus_, _Odontaspis contortidens_ (Fig. 131 B), +_O. incurva_, _O. cuspidata_, _O. attenuata_, _Lamna apiculata_, _L. +compressa_, _Oxyrhina hastalis_ (abundant), _O. desori_, _O. retroflexa_, +_O. eocaena_, _O. minuta_, _Carcharodon auriculatus_ and _C. megalodon_. +An extinct species of Sting Ray, _Myliobatis moorabbinensis_ (Fig. 131 D), +is found at Beaumaris, represented by occasional palatal teeth. Mandibular +and palatine teeth of an extinct genus of Elephant Fish, _Edaphodon_ +(_E. sweeti_) are occasionally found at Beaumaris, and at Grange Burn +near Hamilton. Two extinct forms of the Wrasse family, the Labridae, are +found in Victoria; the pharyngeals of _Labrodon confertidens_ (Fig. 131 +E), occurring at Grange Burn, Hamilton, and those of _L. depressus_, at +Beaumaris. The palatal jaws of a Porcupine Fish, _Diodon formosus_ (Fig. +131 F), are frequently met with at the base of the Kalimnan Series, both +at Grange Burn and Beaumaris. + + +=Oamaru Series, New Zealand.--= + +In New Zealand the Oamaru Series, which is comparable in age with the +Victorian Janjukian, contains numerous fish remains, chiefly teeth +of sharks. These are: _Notidanus primigenius_, _N. marginalis_ (also +occurring in the Waipara Series), _Galeocerdo davisi_, _Odontaspis +incurva_, _O. cuspidata_, _O. attenuata_, _Lamna apiculata_, _L. +compressa_, _Oxyrhina retroflexa_, _Carcharodon auriculatus_, _C. +megalodon_ and _C. robustus_. The teeth of a Sting Ray, _Myliobatis +plicatilis_ and of a species of Sea-bream, _Sargus laticonus_, also occur +in this series (Fig. 130 F). + + +=Pleistocene.--= + +A species of fish belonging to the family of the Perches, _Ctenolates +avus_, has been described from freshwater carbonaceous shale of +Pleistocene age from Nimbin on the Richmond River, New South Wales. + + +=Amphibians: Their Structure.--= + +_AMPHIBIANS._--This group includes amongst living forms the Frogs, +Toads, Newts, and Salamanders. The remains of amphibia are rare in +Australasian rocks, and practically limited to the group of the Triassic +Labyrinthodonts. The Amphibia are distinguished from Reptiles by certain +changes which their young undergo after leaving the egg. In this +intermediate stage they breathe by external gills, these being sometimes +retained together with the internal lungs in the adult stage. In the +older forms of this group the vertebra is of the nature of a notochord, +the joints consisting of a thin bony ring with a gelatinous interior. +The Labyrinthodontia have a long, lizard-like body, short pectoral limbs +as compared with the pelvic, and five-toed feet. The skull is completely +roofed over. The teeth are pointed, with a large pulp cavity and wall of +infolded or plicated dentine (hence the name labyrinthodont--maze-tooth). +The vertebrae are hollow on both sides, sometimes imperfectly ossified, +and with a notochordal canal. Ventral aspect with bony thoracic plates. +Cranial bones deeply sculptured, and carrying mucus canals. + + +=Carbopermian Labyrinthodonts.--= + +The genus _Bothriceps_, probably an Archegosaurian, is represented by two +species, _B. australis_ and _B. major_ from New South Wales (Fig. 132). +The latter species occurs in the Oil Shale (Carbopermian) of Airly. + +[Illustration: =Fig. 132--Bothriceps major, A. S. Woodward.= + +Carbopermian. New South Wales. About 1/11th. nat. size + + (_After A. S. Woodward_). +] + + +=Triassic Labyrinthodonts.--= + +From the Hawkesbury Series near Gosford, New South Wales, the +labyrinthodont, _Platyceps wilkinsoni_ has been described. The skeleton +is nearly complete and exposed on the ventral face; the head is 27mm. +long and 32mm. broad. This specimen is associated with the remains of +ganoid fishes, as _Palaeoniscus_ and _Cleithrolepis_, together with the +equisetum-like plant _Phyllotheca_. + +Other, somewhat doubtful remains having similar affinities to the +labyrinthodonts are also recorded from the Wianamatta beds (Upper Trias) +at Bowral, New South Wales, consisting of a maxilla with teeth and 11 +vertebrae with ribs of the left side. Remains of a labyrinthodont, +_Biloela_, supposed to be related to _Mastodonsaurus_, have been +recorded from the Hawkesbury Series of Cockatoo Island, Port Jackson, +New South Wales, by W. J. Stephens, and consisting of a pectoral plate +compared by that author with _M. robustus_ (now transferred to the genus +_Capitosaurus_). + +The only other recorded remains of this group in Australasia are those +noted by W. J. Stephens from the Kaihiku Series (Trias) at Nugget Point, +Otago; and in the Otapiri Series (Upper Trias) of the Wairoa district, New +Zealand. + + +=Reptilia: Their Structure.--= + +_REPTILIA._--The Reptiles are cold-blooded, vertebrated animals, with +a scaly skin or armour. Their respiration is essentially by means of +lungs, and they are terrestrial or aquatic in habit. The skeleton is +completely ossified (bony). Reptiles, although resembling amphibians +externally, are more differentiated in structure and of generally +larger proportions. They exhibit great diversity of form, especially as +regards their extremities. They were even adapted for flying, as in +the Pterosaurs ("Flying Dragons") with their membranous wing attached +to the anterior limb. The Deinosaurs ("Terrible Reptiles") were often +of great size, exceeding the dimensions of any land mammals, and their +limbs were adapted for walking. The marine reptiles, as the Ichthyosauria +("Fish-lizards") and Sauropterygia ("lizard-finned") had the limbs +transformed into paddles. The neural spines in the vertebra of the Turtles +are laterally expanded into a carapace and united with dermal plates. +The vertebrae of Reptilia show great variation of form, being biplanate +(amphiplatyan), biconcave (amphicoelus), hollow in front (procoelus), +or hollow at the back (opisthocoelus). In the case of Reptiles having +both pairs of limbs developed, the cervical, dorsal, sacral and caudal +regions may be separately distinguished. Amongst the Ophidia (Snakes), +Pythonomorpha ("Sea-lizards") and Ichthyosaurs ("Fish-lizards") there is +no differentiated sacral region. The skull of the Reptiles is nearer that +of Birds than Amphibians. The basiocciput (basal bone of the skull at the +back) articulates with the atlas (top joint of the backbone) by means of +a single condyle (protuberance). All reptiles, with the exception of the +Chelonians (Turtles), and a few others, are furnished with teeth: these +are formed chiefly of dentine with a layer of enamel. + + +=Dentition.--= + +Some teeth have solid crowns (pleodont); some grow from persistent pulps +(coelodont); socketed teeth (thecodont) are inserted in alveoli; some are +fused with the supporting bone along the outer rim or top (acrodont); +whilst others are developed laterally along the flange-like inner rim of +the jaw (pleurodont). + + +=Permian and Triassic Reptiles.--= + +The history of Reptilia commences in Permian and Triassic times, when +they were notably represented by the Theromorphs, _Pareiasaurus_ +and _Tritylodon_ in South Africa; the Proterosauria of the European +and American Permian and Trias, represented by the lizard-like +_Palaeohatteria_ and the dorsally frilled _Dimetrodon_, with its +formidable array of neural spines; also the Rhynchosauria, with their +beak-like jaws of the same formations. These two groups constitute the +order Rhynchocephalia, which is represented at the present day by the +Tuatera of New Zealand. + + +=Triassic Reptile, New Zealand.--= + +The earliest Australian reptilian record is that of a vertebra of +_Ichthyosaurus_ from the Kaihiku Series of Mount Potts, New Zealand +(Triassic). This specimen was named _I. australis_ by Hector, but since +that species name was preoccupied by McCoy in 1867 it is suggested here +that the New Zealand species should be distinguished as _I. hectori_. The +New Zealand occurrence of _Ichthyosaurus_ makes the geological history of +the genus very ancient in this part of the world. + + +=Jurassic Reptiles.--= + +At Cape Paterson, Victoria, in the Jurassic coal-bearing sandstone +an extremely interesting discovery was made a few years ago, of the +ungual bone (claw) of a carnivorous Deinosaur, probably related to +_Megalosaurus_ of the European Jurassic and Cretaceous beds (See Fig. 126, +3, 3 A). The presence of an animal like this in Australia points to the +former existence of a concomitant terrestrial animal fauna, upon which the +deinosaur must have preyed. + +[Illustration: =Fig. 133--Ichthyosaurus australis, McCoy.= + + A--Part of head, showing eye protected by sclerotic plates + + B--Left pectoral paddle. L. Cretaceous. Flinders River, Queensland. + 1/8 nat. size + + (_Nat. Mus. Coll._) +] + + +=Lower Cretaceous Reptiles.--= + +The Rolling Downs formation (Lower Cretaceous) of the Thompson and +Flinders Rivers in Queensland has yielded remains of a Tortoise, +_Notochelone costata_ (see _antea_, Fig. 17); and the interesting +Fish-lizard _Ichthyosaurus_. Numerous and well preserved remains of _I. +australis_ McCoy come from the Flinders River (Fig. 133); whilst _I. +marathonensis_ is recorded from Marathon Station, Queensland. The former +species is typically represented by a nearly complete skeleton, and was +considered by McCoy to be one of the largest examples of the genus, +since a perfect specimen would probably reach the length of 25 feet. Its +teeth resemble those of _I. campylodon_, Carter, from the English Chalk. +Of the Sauropterygia two species of _Pliosaurus_ (_P. macrospondylus_ +and _P. sutherlandi_) have been described from the Lower Cretaceous +of the Flinders River; whilst the latter species has also occurred at +Pitchery Creek, Central Queensland and at Marathon. _P. macrospondylus_ +is distinguished from _P. sutherlandi_ by the roughened edges of +the vertebral centra. Another genus of the "lizard-finned" reptiles +(Sauropterygia), viz., _Cimoliosaurus_, occurs in the Upper Cretaceous of +White Cliffs, New South Wales (Fig. 134 B, C.) + +[Illustration: =Fig. 134--FOSSIL REPTILES.= + + A--Taniwhasaurus oweni. Hector. (Lower jaw). Cretaceous. New Zealand + + B--Cimoliosaurus leucoscopelus, Eth. fil. (Teeth). Up. Cretaceous. New + South Wales + + C--Cimoliosaurus leucoscopelus, Eth. fil. (Phalangeal). Up. + Cretaceous. New South Wales + + D--Miolania oweni, A. S. Woodw. Pleistocene. Queensland + +] + + +=Cretaceous Reptiles, New Zealand.--= + +The Waipara Series (Cretaceous) of New Zealand contains a fairly large +number of reptilian species belonging to several genera among which may +be mentioned _Plesiosaurus_, _Polycotylus_, and _Cimoliosaurus_ among the +Sauropterygia; and _Tylosaurus_ and _Taniwhasaurus_ (Fig. 134 A), marine +lizard-like reptiles, belonging to the sub-order Pythonomopha. + + +=Cainozoic and Pleistocene Reptiles.--= + +The later Cainozoic deposits of Queensland contain remains of Crocodiles +referred to _Pallymnarchus pollens_ (from Maryvale Creek) and _Crocodilus +porosus_ (from Chinchilla and Arcola, near Brisbane, Queensland). The +former species has also occurred at Clunes, whilst _Crocodilus porosus_ is +recorded from the Loddon Valley, both in Victoria. Another late Tertiary +reptile is the remarkable Horned Turtle, _Miolania oweni_, which is found +in Queensland in Pleistocene deposits (Fig. 134 D), and in the Pliocene +(Deep Leads) of Gulgong, New South Wales; whilst a second species of the +same genus, _M. platyceps_, is found in coral sand at Lord Howe Island, +400 miles distant from Australia. This genus has a skull with large bony +protuberances, giving it a horned appearance, and the tail is encased in +a bony sheath. A species of _Miolania_ is also described from Patagonia. +The Cave deposits of Wellington Valley, New South Wales, as well as the +fluviatile deposits of Queensland, have, yielded the bones of several +genera of lizards, including the Giant Lizard (_Megalania_), which, in its +length of 20 feet exceeded that of most living crocodiles. + + +=Birds.--= + +_BIRDS (AVES)._--These warm-blooded animals are closely related to +Reptiles in many essential particulars; and are generally considered +to more nearly approach the Deinosaurs than any other group. The +Ratitae ("Raft-breasted" or keel-less birds) and Carinatae (with keeled +breast-bones), a sub-class including most modern birds, were probably +differentiated before the Cainozoic period. + + +=Jurassic Bird.--= + +The oldest recorded bird, the remarkable _Archaeopteryx_, of the +Upper Jurassic of Bavaria in Europe, belonging to the Saururae +(Reptilian-tailed) is, so far, restricted to the beds of that age. + + +=Miocene Bird, New Zealand.--= + +The earliest known birds in Australasia occur in the Miocene rocks (Oamaru +Series), of New Zealand. In this series, in the Marawhenua Greensands, +a Giant Penguin, _Palaeeudyptes antarcticus_ is found at Kakanui near +Oamaru, at Curiosity Shop near Christchurch and at Brighton near Nelson, +New Zealand: this interesting occurrence shows that these restricted +antarctic birds had already become an established type as early as the +Miocene. + + +=Victorian Cainozoic Bird.--= + +The impression of a bird's feather, probably of a Wader, has lately been +described from Western Victoria (see _antea_ Fig. 16 and Fig. 135). This +occurs in ironstone, on the surface of which are also impressions of Gum +(_Eucalyptus_) and Native Honeysuckle (_Banksia_) leaves, of species +closely related to those now growing in the same locality. This ironstone +is probably of Janjukian age, and may therefore be coincident with the New +Zealand occurrence of the _Palaeeudyptes_ in the Oamaru Series. + + +=Pliocene Moa, New Zealand.--= + +In the Wanganui System (Pliocene) the Putiki Beds have yielded bones of a +small Moa (_Dinornis_), probably the oldest example of the group of great +flightless birds which later predominated in New Zealand. + +[Illustration: =Fig. 135--Impression of Bird's Feather in Ironstone.= + +Wannon River, Victoria. (Enlarged).] + + +=Pleistocene Struthious Birds, Australia.--= + +Bones of a struthious or Ostrich-like bird, described by Owen under +the name of _Dromornis australis_, a bird as large as the Moa, have +been recorded from the Pleistocene of Peak Downs and the Paroo River, +Queensland. Indeterminate species of the same genera occur in Phillip Co., +New South Wales, and the Mount Gambier Caves, South Australia; whilst +_Dromaeus patricius_ is known from King's Creek, Darling Downs, Queensland. + +_Genyornis newtoni_ is an extinct bird allied to the Emeus; it has been +found in Pleistocene deposits at Lake Callabonna, South Australia, and +other fragmentary remains have been identified by Dr. Stirling and Mr. +Zietz from Mount Gambier and Queensland. Regarding the build and habits +of _Genyornis_, those authors remark that "Its legs combine a huge femur +nearly as massive, in all but length, as that of _Dinornis maximus_, and +a tibia equalling that of _Pachyornis elephantopus_ with the relatively +slender metatarse of _Dinornis novae-zealandiae_ (_ingens_) and toes which +are insignificant beside those of any of the larger moas."... "In height +it may be confidently stated to have been from 6 feet to 6 feet 6 inches, +that is if the neck should have been of proportions similar to those of +_Pachyornis elephantopus_." Those authors also attribute a slow, sluggish +habit to the bird, and suggest that herbage rather than roots formed its +food. It is very probable that the footprints of birds found in the older +dune rock of Warrnambool, Victoria, associated with the doubtful "human +footprints" may have been made by _Genyornis_ or a related form. + +An extinct Emu, _Dromaeus minor_, has lately been described from the +sub-recent deposits in King Island, Bass Strait. + + +=Pleistocene Carinate Birds, Australia.--= + +Many genera of carinate birds belonging to living Australian types have +been identified by De Vis from the fluviatile deposits on the Darling +Downs, Queensland. These include Falcons (_Taphaetus_ and _Necrastur)_; +a Pelican (_Pelicanus_); an Ibis (_Palaeopelargus_); a Spoonbill +(_Platalea_); Ducks (_Anas_, _Dendrocygna_, _Biziura_ and _Nyroca_); a +Darter (_Plotus_); a Pigeon (_Lithophaps_); a Ground-pigeon (_Progura_); a +Mound-builder (_Chosornis_); a Rail (_Porphyrio_); Moor-hens (_Gallinula_, +_Tribonyx_ and _Fulica_); and a Stork (_Xenorhynchus_). + + +=Pleistocene and Holocene Birds, New Zealand.--= + +In New Zealand numerous remains of birds are found, chiefly in the +Pleistocene strata, associated with Moa bones: such are _Cnemiornis_, the +Flightless Pigeon Goose (Fig. 135); _Harpagornis_, a predatory hawk-like +bird larger than any existing eagle; and _Aptornis_, an extinct Rail. The +sand-dunes, peat bogs, swamps, river alluvium, caves and rock shelters +of New Zealand often contain numerous remains of the gigantic Moa birds +included in the genera _Dinornis_, _Pachyornis_ and _Anomalopteryx_, of +which perhaps the best known are _D. giganteus_, _D. maximus_ (Fig. 136), +_D. robustus_, _P. elephantopus_ (Fig. 137), and _A. antiqua_. Some of the +species have become so recently extinct that remains of their skin and +feathers have been preserved in fissures in the rocks where they were +shielded from the influence of air and moisture. The remains of Moa birds +are very abundant in some of the localities as at Hamilton in Southland, +where, as Hutton estimated, the remains of at least 400 birds were +contained within a radius of 25 feet. + +[Illustration: =Fig. 136--Cnemiornis calcitrans, Owen.= + +Pleistocene. New Zealand. 1/15th. nat. size + + (_After Owen_). +] + +[Illustration:=Fig. 137--Dinornis maximus, Owen. (Great Moa).= + +Pleistocene and Holocene. New Zealand. + +Vertical height, 8 ft. Measured along spine, 10 ft. 8 in. + + (_Nat. Mus. Coll._) +] + +[Illustration: =Fig. 138--Pachyornis elephantopus, Owen sp.= + +Pleistocene. New Zealand. About 1/26th. nat. size. + + (_After Owen_). +] + + +=Mammalia: Early Types.--= + +_MAMMALIA._--The history of those warm-blooded animals, the mammals, +commences in the early part of the Mesozoic period. It was then that the +skull began to assume the characters seen in the modern quadrupeds, and +their well-formed limb-bones, and fusion of the three bones on each side +of the pelvic arch to form the innominate bone, also show relationship to +the later types. The earliest ancestral mammalian forms seem to be related +to the theromorphic reptiles, predominant in the Permian and Trias. The +mammals first to make their appearance were probably related to those of +the Monotreme and Marsupial orders. More nearly related to the former is +the group of mammals of the Mesozoic period, the Multituberculata. + + +=Multituberculata.--= + +This group comprises the Triassic _Tritylodon_ (South Africa and Germany); +the Upper Jurassic _Bolodon_ (England and United States); the Upper +Jurassic to Lower Cainozoic _Plagiaulax_ (England, United States and +France); and the Lower Eocene _Polymastodon_ (New Mexico). The molar teeth +are ridged longitudinally, and carry numerous tubercles, hence the name of +the group, and resemble the deciduous teeth of the Duck-billed Platypus +(_Ornithorhynchus_). + + +=Monotremata.--= + +The Monotremata are represented at the present day in Australia and New +Guinea by the _Echidna_ or Spiny Anteater, and by the _Ornithorhynchus_ or +Duck-billed Platypus of Eastern Australia and Tasmania. These egg-laying +mammals show relationship towards the reptiles both in structure and in +methods of reproduction. + +A Pliocene species of _Ornithorhynchus_ (_O. maximus_) has been recorded +from the Deep-leads of Gulgong, New South Wales, and the same beds have +yielded the remains of _Echidna (Proechidna) robusta_. Remains of another +species, _Echidna, (P.) oweni_, have been described from the Pleistocene +Cave-breccias of the Wellington Valley Caves, New South Wales; and +_Ornithorhynchus agilis_ is found in deposits of similar age in Queensland. + + +=Marsupials.--= + +The Marsupials or pouched mammals belong to the sub-class Metatheria. They +are divided into Diprotodontia and Polyprotodontia, accordingly as they +possess a single pair of incisor teeth in the lower jaw, or many front +teeth, hence the names of the two sub-orders. A later classification of +the Marsupials is that of their division into syndactyla and diadactyla. + +The diadactyla have the second and third toes separate, and are +represented by the family Dasyuridae or Native Cats. These are +polyprotodont. They are the most archaic of the marsupial group. Remains +of _Dasyurus_, both of extinct and still living species are found in +Pleistocene Cave-breccias in Victoria and New South Wales. The Tasmanian +Devil (_Sarcophilus ursinus_) (Fig. 138, 139) and the Tasmanian Wolf +(_Thylacinus cynocephalus_), still living in Tasmania, have left numerous +remains on the mainland, in Victoria and New South Wales. Of the latter +genus an extinct species is _T. major_ from the Pleistocene of Queensland +(Fig. 140). + +[Illustration: =Fig. 139= + +=Skeleton of Sarcophilus ursinus, Harris sp. (Tasmanian devil).= + + (_F. J. Moore, prep._) +] + +[Illustration: =Fig. 140= + +=Skull of Sarcophilus ursinus, Harris sp. (Tasmanian devil).= + +Pleistocene. Queenscliff, Victoria. About 1/2 nat. size + + (_After McCoy_). +] + +The syndactyla have the second and third toes enclosed in a common skin. +The Peramelidae and the Notoryctidae are polyprotodont. The remainder are +all diprotodont. The Peramelidae or Bandicoot family are represented in +Pleistocene Cave-breccias in New South Wales by the genera _Peragale_ and +_Perameles_. + +[Illustration: =Fig. 141--Thylacinus major, Owen.= + +Hind part of mandible, outer side. Pleistocene. Queensland. + +1/2 nat. size] + + +=Pleistocene Diprotodonts.--= + +Pleistocene remains of the diprotodont forms of this syndactylous group +are _Phascolomys_ (the Wombat), perhaps ranging as low as Upper Pliocene +(_P. pliocenus_) (Fig. 141); _Phascolonus (P. gigas)_ (Fig. 142 A)[4], +a large Wombat from Queensland and New South Wales and South Australia; +the Giant Kangaroos, as _Macropus titan_ (Queensland, New South Wales, +Victoria and South Australia), _Procoptodon goliah_ (Queensland, New South +Wales and Victoria), _Sthenurus atlas_ (New South Wales, Queensland, +Victoria and South Australia), _Palorchestes azael_ (Victoria, New +South Wales and Queensland); also the great _Diprotodon_, the largest +known marsupial, as large as, and rather taller than, a rhinoceros, +found in almost every part of Australia, with an allied form referred +to _Nototherium_ occurring also in Tasmania (Figs. 143, 144, 145). +_Nototherium_ (Queensland, South Australia and Victoria), was a smaller +animal than _Diprotodon_, with a shorter and broader skull and similar +dentition. Remains of the extinct "Marsupial Lion," _Thylacoleo carnifex_, +an animal allied to the phalangers, have been found in Cave-deposits in +New South Wales, Queensland, Victoria and Western Australia. Incised bones +of other animals, which are believed to have been gnawed by _Thylacoleo_, +have been found associated with its remains. _Thylacoleo_ possessed a +peculiar dentition, the first pair of incisors in the upper jaw being +very large and trenchant, whilst the canine and two anterior premolars +are small and functionless: the lower jaw has also a pair of large +first incisors, behind which are two small premolars, and an enormous +chisel-edged last premolar biting against a similar tooth in the upper jaw +(Fig. 146). + +[Footnote 4: This genus was described by Owen in 1872 as a sub-genus of +_Phascolomys_ founded on some cheek-teeth; and subsequently, in 1884, +the same author described some incisors under the name of _Sceparnodon +ramsayi_, which are now known to belong to the same animal that bore the +cheek-teeth.] + +[Illustration: =Fig. 142--Mandible of Phascolomys pliocenus, McCoy.= + +(?) Upper Pliocene ("Gold Cement.") Dunolly, Vict. + +About 1/2 nat. size. + + (_After McCoy_). +] + +[Illustration: =Fig. 143--CAINOZOIC TEETH and OTOLITH.= + + A--Phascolonus gigas, Owen. (Molar). Pleistocene. Queensland + + B--Parasqualodon wilkinsoni, McCoy. (Molar). Cainozoic (Janj.) Vict. + + C--Parasqualodon wilkinsoni, McCoy. (Incisor). Cainozoic (Janj.) Vict. + + D--Metasqualodon harwoodi, Sanger sp. (Molar). Cainozoic (Janj.) South + Australia + + E--Kekenodon onamata, Hector. (Molar). Cainozoic (Oamaruian). New + Zealand + + F--Cetotolithes nelsoni, McCoy. (Tympanic bone). Cainozoic (Janj.) + Victoria + +] + +[Illustration: =Fig. 144--Diprotodon australis, Owen.= + +Pleistocene. South Australia. + + (_After Stirling and Zeitz_). +] + +[Illustration: =Fig. 145--Upper Surface of the Right Hind Foot of +Diprotodon australis=. + + A--With the Astragalus (ankle-bone) in position. + + B-- " " " " removed. + +Cir. 1/8 nat. size.] + +[Illustration: =Fig. 146--Diprotodon australis, Owen. (Restored).= + +From a sketch by C. H. Angas.] + +[Illustration: =Fig. 147--Thylacoleo carnifex, Owen.= + +Right lateral aspect of skull and mandible. + + Pleistocene. Australia. 1/5th nat. size. + + c, canine. i, incisors. m, molars. pm, premolars. +] + +[Illustration: =Fig. 148--Wynyardia bassiana, Spencer.= + +Upper Cainozoic (Turritella bed). Table Cape. Tasmania. + +2/7th nat. size. + + (_Casts in Nat. Mus. Coll._) +] + + +=Oldest Known Marsupial.= + +The oldest marsupial found in Australia is probably _Wynyardia bassiana_ +(Fig. 147), whose remains occurred in the _Turritella_-bed at Table +Cape, which is either of Miocene or Lower Pliocene age. This stratum +occurs above the well-known _Crassatellites_-bed (Miocene) of that +locality. So far as can be gathered from its incomplete dentition, +_Wynyardia_ represents an annectant form between the Diprotodonts and the +Polyprotodonts. + + +=Pleistocene Genera, also Living.--= + +Besides the genera above enumerated, many other marsupials of well-known +living species are represented by fossil remains in Cave-deposits and on +"sand-blows" in most of the Australian States. The genera thus represented +in the Pleistocene deposits of Australia are _Bettongia_ (Prehensile +Rat-Kangaroo); _Dasyurus_ (Native Cat); _Hypsiprymnus_ (Rat-Kangaroo); +_Macropus_ (Kangaroo); _Perameles_ (Bandicoot); _Petaurus_ (Flying +Phalanger); _Phalanger_ (Cuscus); _Phascolomys_ (Wombat); _Sarcophilus_ +(Tasmanian Devil); _Thylacinus_ (Tasmanian Wolf). + + +=Cetacea.--= + +The order Cetacea includes Whales, Dolphins and Porpoises. The earliest +known forms belong to the sub-order Archaeoceti, and whilst absent from +Australian deposits, are found in the Eocene of Europe, Northern Africa +and North America. + + +=Odontoceti: Toothed Whales.--= + +Remains of Cetacea are first met with in Australian rocks in the Oligocene +(Balcombian) of Victoria. At Muddy Creek near Hamilton fragments of ribs +and other bones of cetacea, not yet determined, occur in the tenacious +blue clays of the lower part of the Clifton Bank section. In Australia +and New Zealand the oldest determinable remains of this order belong to +the Odontoceti, members of which range from Miocene to Pliocene. Teeth +of the toothed whales like _Squalodon_ of the Miocene of France and +Bavaria have been found in New Zealand (_Kekenodon_); in South Australia +(_Metasqualodon_); and in Victoria (_Parasqualodon_). In Victoria the +teeth of Squalodontidae occur in the Janjukian beds of Cape Otway, +Waurn Ponds and Torquay, represented by molars and anterior teeth of +_Parasqualodon wilkinsoni_ (Fig. 142 B, C). The same species also occurs +at Table Cape, Tasmania, in beds of similar age. Teeth of _Metasqualodon +harwoodi_ (Fig. 142 D) occasionally occur in the white polyzoal rock of +the Mount Gambier district, South Australia. The gigantic toothed whale, +_Kekenodon onamata_ (Fig. 142 E) occurs in the Marawhenua Greensands +(Oamaru Series) at Waitaki Valley, Waihao, Ngapara, Waikouaiti and Milburn +in New Zealand. The molar teeth of this striking species, with their +serrated crowns, measure nearly five inches in length. + + +=Ear-bones of Whales.--= + +The tympanic bones of whales are not uncommon in the Janjukian beds of +Waurn Ponds, near Geelong, Victoria; and they are occasionally found +in the basement bed of the Kalimnan at Beaumaris, Port Phillip. In the +absence of any distinctive generic characters they have been referred +to the quasi-genus _Cetotolithes_ (Fig. 142 F). McCoy has expressed the +opinion that they may perhaps be referable to the ziphioid or beaked +whales, for undoubted remains of that group, as teeth of _Ziphius +geelongensis_, occur in these same beds; as well as portions of their +rostrate crania, in the Kalimnan basement beds at Grange Burn, near +Hamilton. The large curved and flattened teeth of _Ziphius (Dolichodon) +geelongensis_ are occasionally found, more or less fragmentary, in the +polyzoal rock of Waurn Ponds. + +[Illustration: =Fig. 149.--Tooth of Scaldicetus macgeei, Chapm.= + +An Extinct Sperm Whale. + +From the Kalimnan beds of Beaumaris, Port Phillip, Victoria. + +About 3/4 nat. size.] + + +=Kalimnan-Scaldicetus.--= + +From the Kalimnan Series (Lower Pliocene) of Beaumaris, Port Phillip, +there was described a short time since, a remarkably well preserved +specimen of _Scaldicetus_ tooth belonging to a new form, _S. macgeei_ +(Fig. 148). Another species of the genus, with teeth of a slender form, +has been found in the same geological series, at Grange Burn, near +Hamilton. In only one other locality besides Australia does the genus +occur, viz., at Antwerp, Belgium, in Crag deposits of Lower Pliocene age. + + +=Sirenia.--= + +The order Sirenia (Manatees and Dugongs) is represented in the Australian +Pleistocene by _Chronozoön australe_. The remains consist of the parietal +and upper part of the occipital bones of the skull, and were discovered +in the fluviatile deposits on the Darling Downs, Queensland. This fossil +skull, according to De Vis, had a shallower temporal fossa and feebler +masticating muscles, as well as a less highly developed brain than the +existing Dugong. + + +=Carnivora.--= + +The order Carnivora is represented in Australia by the Native Dog or Dingo +(_Canis dingo_). It is by no means a settled question whether the Dingo +can boast of very great antiquity. The evidence of its remains having +been found under volcanic tuff beds in Victoria is not very convincing, +for the original record does not indicate the precise position where +the bones were found. The fact of the remains of the Dingo having been +found in Cave deposits often associated with extinct marsupials, goes a +good way to prove its antiquity. McCoy was strongly inclined to the view +of its Pleistocene age, and points out that it shows cranial characters +intermediate between the Dogs of South America and the Old World. Fossil +remains of the Dingo, associated with Pleistocene mammalian forms have +been recorded from the Wellington, Valley Caves, New South Wales; from +the Mount Macedon Cave, near Gisborne; and in the neighbourhood of +Warrnambool, Western Victoria. + + +=Pinnipedia.--= + +Of the fin-footed Carnivores or Seals and Walruses, the earliest +Australasian record is that of the remains of a small seal in the Okehu +shell-beds near Wanganui, found in association with the bones of a small +Moa-bird (_Dinornis_). + + +=Newer Pliocene Seal.--= + +This seal was referred by Hector to _Arctocephalus cinereus_, a species +synonymous, however, with the widely distributed living Seal, _Otaria +forsteri_, Lesson, of the Southern Ocean. Another and larger species of +eared seal allied to the living Fur Seal, _Otaria forsteri_, occurs in +Victoria. + + +=Pleistocene Seal.--= + +This fossil was named _Arctocephalus williamsi_ by McCoy, and was found +in Pleistocene deposits at Queenscliff, Port Phillip, at 5 feet below the +surface, in marl and sand stone overlain with limestone. Although referred +at the time of description to the Pliocene, it has since been proved +that at this locality there is a considerable thickness of practically +sub-recent material which is more accurately classed with the Pleistocene. +Similar remains of eared seals are not uncommon in the Pleistocene +deposits of the Otway Coast. + + +=Subrecent Human Remains.= + +On turning to the occurrence of "human fossils" in Australia we find the +geological evidence for any great antiquity of man on this continent to +be very scanty and inconclusive. This does not, however, imply that +man's existence in Australia will not eventually be proved to date back +far beyond the period of the "kitchen middens" of modern aspect, such as +are now exposed on the slopes behind the sea-beaches, and on the inland +camping grounds. Almost all the records of Australian human remains that +have been found in other than ordinary burial places, have proved to be +of comparatively recent date. For example, the partially lime-encrusted +body found in the cave in the Mosquito Plains, north of Penola, South +Australia, recorded by Tenison Woods, is that of an aborigine who, in the +early days of settlement, crawled into the cave in a wounded condition. +Other occurrences of human remains in caves, but of fairly recent date +are, a child's skull found in a small cave at Bungonia, Co. Argyle, New +South Wales, recorded by Etheridge; and the non-petrified limb-bones +found in a cave at Wellington, New South Wales, recorded by Krefft, which +were probably washed in from the surface in recent times. As regards the +former, in Western Australia, as observed by Froggatt, the natives at the +present time seek shelter in caves, where these occur, instead of building +mia-mias. + +A more interesting, because probably much older, occurrence of human +remains has been described by Etheridge and Trickett from one of the +Jenolan Caves (Skeleton Cave); and those authors conclude from "The +great lapse of time that must have accrued to enable the changes already +outlined to have taken place since the introduction of the remains into +the Skeleton Cave," that these remains are ancient. + +[Illustration: =Fig. 150--Impressions of Footprints in dune sand-rock.= + +Warrnambool, Victoria. 1/9 nat. size. + + (_F. C. Photo_). (_Warrnambool Museum_). +] + +Curious footprints supposed to resemble impressions of human feet with +accompanying impress as if made by natives seated, have been long known +from the older sand-dune rock of Warrnambool. They were found at Kellas' +Quarry, on the Port Fairy Road in 1890 and at a depth of 54 feet. In +November, 1912, a further discovery of similar footprints were found at +Messrs. Steere Bros.' Quarry, Warrnambool, at a depth of 10 feet, as a +block of stone was being removed for building purposes. These footprints +are even more obscure than those previously found, and it would be unsafe +to affirm their human origin, although they are suggestive of such. Their +antiquity is certainly great, since the lavas and tuffs of the Tower +Hill district are found overlying this old dune-rock. Other footprints +associated with these resemble those of the Dingo and a gigantic bird, +possibly like _Genyornis_. + + +=Probable Origin of Aborigines.--= + +Ethnology appears to throw more light upon the subject than does geology. +Australia has in the past been peopled by two distinct types of man. +(1), the ancestors of the Tasmanians, now alas, extinct, who according +to some authorities came by way of Australia from Papua through the +Malay Peninsula, passing over to Tasmania from the mainland before the +separation caused by the subsidence of the Bass Strait area; and who +were represented by a negroid or woolly-haired type: (2), the present +aboriginals of Australia, showing affinities with the Dravidians of +Southern India, a primitive race from whose original stock the white +Caucasian races of Europe were derived. By intermarriage with a negroid +race like the Melanesian, it is supposed that the black Caucasian gave +rise to the present Australian mixed aboriginal type, with negroid +features, but possessing the long black hair and keener intellect of the +"melanochroi," as the dark Eurasian stock was termed by Huxley. + + +=Aboriginal Implements.--= + +The stone implements fashioned by the Tasmanian aboriginals were roughly +chipped and of primitive type, of such forms as used at the present day by +the Bushmen of South Africa, and representing the eoliths and palaeoliths +of early man in the south of England. The implements of the Australian +aboriginals on the other hand include besides these both flakes and +worked and polished tools, such as were produced by the Neolithic men of +Europe, as contrasted with the typically rough palaeolithic tools of the +Tasmanian, who never grooved his axes for hafting as did the Australian +aboriginal. According to some authorities the Tasmanians represent +palaeolithic or even eolithic man in the character of their implements; +whilst the Australian resembles the Middle or Mousterian stage of early +man in certain of their ethnological characters and in the forms of their +implements, although a marked exception is seen in their manufacture of +polished adzes, of the neolithic period and in the use of bone implements +such as were used in Europe in Upper Palaeolithic times. So far no human +remains or handiwork in the form of chipped implements have been found +in other than superficial deposits, either in Tasmania or Australia. The +incised bone-fragment found near Ballarat, in a bed of silt beneath a +sheet of basalt which flowed from Mount Buninyong, is believed by some +to be evidence of man's handiwork in the early Pleistocene, though by +others thought to have been cut by the teeth of the "marsupial lion" +(_Thylacoleo_). A stone axe of basalt, grooved for the purpose of +mounting in a handle, was found in gravel at Ballarat at a depth of 22 +inches from the surface. This, however, is no proof of man's antiquity, +for superficial deposits of much greater depth are easily accumulated +within a short period. Another implement was found at Maryborough in +Queensland in gravels at a depth of 4 feet from the surface, but not +below the basalt of the main lead. In this case it is believed that the +implement may have fallen into a natural hollow or wombat-burrow. A bone +pointer, such as used by native medicine men, was some years ago found +buried in the Miocene marls of Waurn Ponds near Geelong. Its presence in +so old a rock is easily explained from the fact that in the aboriginal +ceremonies the pointer was buried after the incantations. Seeing the +difficulties in the way of discovering reliable occurrences of man's +handiwork in isolated examples amongst the older superficial deposits of +silt and gravels, the ancient sand-dunes of Victoria, which date back +at least to Upper Pliocene, should afford favourable conditions for the +preservation of any really ancient kitchen middens, did such exist. +Moreover, these deposits would have been less liable to disturbance when +once they were covered, than the inland deposits, for the former are now +consolidated into a tolerably hard stone. + + +=Antiquity of Man in Australia.--= + +A strong argument in favour of a considerable antiquity for man in +Australia is the fact that the dialects are many, and marriage and tribal +customs more complex and intricate than would be found in a comparatively +recent primitive race. In any case, it is quite possible, if not probable, +that man was in southern Australia before the termination of the last +phase of volcanic activity, since the tuff beds of Koroit, for example, +are quite modern and were laid down on a modern sea-beach strewn with +shells identical in species and condition with those now found thrown +up in the vicinity at high tide. This view is quite compatible with the +occurrence of dingo remains (assuming this animal was introduced by man) +in cave deposits in Australia, associated with extinct forms of marsupials. + + * * * * * + + +COMMON OR CHARACTERISTIC FOSSILS OF THE FOREGOING CHAPTER. + + +FISHES. + + _Thyestes magnificus_, Chapman. Silurian: Victoria. + + _Asterolepis australis_, McCoy. Middle Devonian: Victoria. + + _Ganorhynchus süssmilchi_, Etheridge fil. Devonian: New South Wales. + + _Gyracanthides murrayi_, A. S. Woodward. Lower Carboniferous: Victoria. + + _Acanthodes australis_, A. S. Woodward. Lower Carboniferous: Victoria. + + _Ctenodus breviceps_, A. S. Woodward. Lower Carboniferous: Victoria. + + _Strepsodus decipiens_, A. S. Woodward. Lower Carboniferous: Victoria. + + _Elonichthys sweeti_, A. S. Woodward. Lower Carboniferous: Victoria. + + _Physonemus micracanthus_, Chapman. Lower Carboniferous: Victoria. + + _(?) Deltodus australis_, Eth. fil. Carbopermian: Queensland. + + _Tomodus (?) convexus_, Agassiz. Carbopermian: New South Wales. + + _Edestus davisii_, H. Woodward. Carbopermian: W. Australia. + + _Peocilodus jonesi_, Agassiz. Carbopermian: W. Australia. + + _Gosfordia truncata_, A. S. Woodw. Triassic: New South Wales. + + _Myriolepis clarkei_, Egerton. Triassic: New South Wales. + + _Apateolepis australis_, A. S. Woodw. Triassic: New South Wales. + + _Dictyopyge robusta_, A. S. Woodw. Triassic: New South Wales. + + _Belonorhynchus gigas_, A. S. Woodw. Triassic: New South Wales. + + _Semionotus australis_, A. S. Woodw. Triassic: New South Wales. + + _Pristisomus latus_, A. S. Woodw. Triassic: New South Wales. + + _Cleithrolepis granulatus_, Egerton. Triassic: New South Wales. + + _Pholidophorus gregarius_, A. S. Woodw. Triassic: New South Wales. + + _Pleuracanthus parvidens_, A. S. Woodw. Upper Trias: New South Wales. + + _Sagenodus laticeps_, A. S. Woodw. Upper Trias: New South Wales. + + _Palaeoniscus crassus_, A. S. Woodw. Upper Trias: New South Wales. + + _Elonichthys armatus_, A. S. Woodw. Upper Trias: New South Wales. + + _Elpisopholis dunstani_, A. S. Woodw. Upper Trias: New South Wales. + + _Pholidophorus australis_, A. S. Woodw. Upper Trias: New South Wales. + + _Psilichthys selwyni_, Hall. Jurassic: Victoria. + + _Leptolepis crassicauda_, Hall. Jurassic: Victoria. + + _Ceratodus avus_, A. S. Woodw. Jurassic: Victoria. + + _Coccolepis australis_, A. S. Woodw. Jurassic: New South Wales. + + _Aphnelepis australis_, A. S. Woodw. Jurassic: New South Wales. + + _Aetheolepis mirabilis_, A. S. Woodw. Jurassic: New South Wales. + + _Archaeomaene tenuis_, A. S. Woodw. Jurassic: New South Wales. + + _Leptolepis talbragarensis_, A. S. Woodw. Jurassic: New South Wales. + + _Lamna daviesii_, Eth. fil. Lower Cretaceous: Queensland. + + _Lamna appendiculatus_, Agassiz. Lower Cretaceous: Queensland. + + _Corax australis_, Chapm. Lower Cretaceous: Queensland. + + _Aspidorhynchus_ sp. Lower Cretaceous: Queensland. + + _Belonostomus sweeti_, Eth. fil. and A. S. Woodw. Lower Cretaceous: + Queensland. + + _Portheus australis_, A. S. Woodw. Lower Cretaceous: Queensland. + + _Cladocyclus sweeti_, A. S. Woodw. Lower Cretaceous: Queensland. + + _Notidanus marginalis_, Davis. Cretaceous: New Zealand. + + _Lamna compressa_, Agassiz. Cretaceous: New Zealand. + + _Callorhynchus hectori_, Newton. Cretaceous: New Zealand. + + _Ischyodus thurmanni_, Pictet and Campiche. Cretaceous: New Zealand. + + _Odontaspis contortidens_, Agassiz. Cainozoic (Bal. and Janj.): + Victoria. + + _Lamna apiculata_, Ag. sp. Cainozoic (Bal. and Janj.): Victoria. Also + Cainozoic (Oamaru Series): New Zealand. + + _Carcharodon megalodon_, Agassiz. Cainozoic (Bal. Janj. and Kal.): + Victoria. Also Cainozoic (Oamaru Series): New Zealand. + + _Cestracion cainozoicus_, Chapm. and Pritch. Cainozoic (Janj. and + Kal.): Victoria. + + _Asteracanthus eocaenicus_, Tate sp. Cainozoic (Janj. and Kal.): + Victoria. + + _Galeocerdo davisi_, Chapm. and Pritch. Cainozoic (Janj.): Victoria. + Also Cretaceous (Waipara Series) and Cainozoic (Oamaru Series): + New Zealand. + + _Carcharoides totuserratus_, Ameghino. Cainozoic (Janj.): Victoria. + + _Odontaspis incurva_, Davis sp. Cainozoic (Janj. and Kal.): Victoria. + Also Cainozoic (Oamaru Series): New Zealand. + + _Oxyrhina retroflexa_, Agassiz. Cainozoic (Janj.): Victoria. Also + Cainozoic (Oamaru Series): New Zealand. + + _Carcharodon auriculatus_, Blainville sp. Cainozoic (Janj. and Kal.): + Victoria. + + _Acanthias geelongensis_, Chapm. and Pritch. Cainozoic (Janj.): + Victoria. + + _Ischyodus mortoni_, Chapm. and Pritch. Cainozoic (Janj.): Tasmania. + + _Notidanus jenningsi_, Chapm. and Pritch. Cainozoic (Kal.): Victoria. + + _Galeocerdo aduncus_, Agassiz. Cainozoic (Kal.): Victoria. + + _Oxyrhina hastalis_, Agassiz. Cainozoic (rare in Balc. and Janj., + abundant in Kal.): Victoria. + + _Myliobatis moorabbinensis_, Chapm. and Pritch. Cainozoic (Kal.): + Victoria. + + _Edaphodon sweeti_, Chapm. and Pritch. Cainozoic (Kal.): Victoria. + + _Labrodon confertidens_, Chap. and Pritch. Cainozoic (Kal.): Victoria. + + _Diodon formosus_, Chapm. and Pritch. Cainozoic (Kal.): Victoria. + + _Notidanus marginalis_, Davis. Cretaceous (Waipara Series); and + Cainozoic (Oamaru Series): New Zealand. + + _Myliobatis plicatilis_, Davis. Cainozoic (Oamaru Series): New Zealand. + + _Sargus laticonus_, Davis. Cainozoic (Oamaru Series): New Zealand. + + _Ctenolates avus_, A. S. Woodw. Pleistocene: New South Wales. + + _Neoceratodus forsteri_, Krefft, sp. Pleistocene: New South Wales. + + +AMPHIBIA. + + _Bothriceps australis_, Huxley. Carbopermian: New South Wales. + + _Bothriceps major_, A. S. Woodw. Carbopermian: New South Wales. + + _Platyceps wilkinsoni_, Stephens. Triassic: New South Wales. + + +REPTILIA. + + _Ichthyosaurus hectori_, Ch. (nom. mut.). Triassic: New Zealand. + + _(?) Megalosaurus_ sp. Jurassic: Victoria. + + _Notochelone costata_, Owen sp. Lower Cretaceous: Queensland. + + _Ichthyosaurus australis_, McCoy. Lower Cretaceous: Queensland. + + _Ichthyosaurus marathonensis_, Eth. fil. Lower Cretaceous: Queensland. + + _Cimoliosaurus leucoscopelus_, Eth. fil. Upper Cretaceous: New South + Wales. + + _Plesiosaurus australis_, Owen. Cretaceous: New Zealand. + + _Polycotylus tenuis_, Hector. Cretaceous: New Zealand. + + _Cimoliosaurus haastii_, Hector sp. Cretaceous: New Zealand. + + _Tylosaurus haumuriensis_, Hector sp. Cretaceous: New Zealand. + + _Taniwhasaurus oweni_, Hector. Cretaceous: New Zealand. + + _Pallymnarchus pollens_, De Vis. Pleistocene: Queensland and Victoria. + + + _Crocodilus porosus_, Schneider. Pleistocene: Queensland and Victoria. + + _Miolania oweni_, A. S. Woodw. Pliocene (Deep-leads): New South Wales. + Pleistocene: Queensland. + + _Miolania platyceps_, Owen. Pleistocene: Lord Howe Island. + + _Megalania prisca_, Owen. Pleistocene: Queensland. + + +BIRDS. + + _Palaeeudyptes antarcticus_, Huxley. Cainozoic (Oamaru Series): New + Zealand. + + _Dinornis_ sp. Cainozoic (Petane Series): New Zealand. + + _Pelecanus proavis_, De Vis. Pleistocene: Queensland. + + _Platalea subtenuis_, De Vis. Pleistocene: Queensland. + + _Anas elapsa_, De Vis. Pleistocene: Queensland. + + _Gallinula strenuipes_, De Vis. Pleistocene: Queensland. + + _Fulica prior_, De Vis. Pleistocene: Queensland. + + _Dromornis australis_, Owen. Pleistocene: Queensland and New South + Wales. + + _Dromaeus patricius_, De Vis. Pleistocene. Queensland. + + _Dromaeus minor_, Spencer. Pleistocene: King Island. + + _Genyornis newtoni_, Stirling and Zietz. Pleistocene: S. Australia. + + _Cnemiornis calcitrans_, Owen. Pleistocene: New Zealand. + + _Harpagornis moorei_, von Haast. Pleistocene: New Zealand. + + _Aptornis otidiformis_, Owen sp. Pleistocene: New Zealand. + + _Dinornis giganteus_, Owen. Pleistocene and Holocene: N. Id., New + Zealand. + + _Pachyornis elephantopus_, Owen sp. Pleistocene and Holocene: S. Id., + New Zealand. + + _Anomalopteryx antiqua_, Hutton. Pleistocene: S. Id., New Zealand. + + +MAMMALIA. + + _Ornithorhynchus maximus_, Dun. Cainozoic (Kalimnan or L. Pliocene): + New South Wales. + + _Echidna (Proechidna) robusta_, Dun. Cainozoic (Kalimnan): New South + Wales. + + _Ornithorhynchus agilis_, De Vis. Pleistocene: New South Wales. + + _Echidna (Proechidna) oweni_, Krefft. Pleistocene: New South Wales. + + _Wynyardia bassiana_, Spencer. Cainozoic (Kalimnan): Tasmania. + + _Dasyurus maculatus_, Kerr sp. Pleistocene: Victoria and New South + Wales. Living: Queensland, New South Wales, Victoria and Tasmania. + + _Phascolomys pliocenus_, McCoy. Cainozoic (Werrikooian): Victoria. + + _Sarcophilus ursinus_, Harris sp. Pleistocene: Victoria and New South + Wales. Living: Tasmania. + + _Thylacinus cynocephalus_, Harris sp. Pleistocene: Victoria and New + South Wales. Living: Tasmania. + + _Thylacinus spelaeus_, Owen. Pleistocene: Queensland and New South + Wales. + + _Thylacinus major_, Owen. Pleistocene: Queensland. + + _Peragale lagotis_, Reid sp. Pleistocene: New South Wales. Living: S. + Australia and W. Australia. + + _Perameles gunni_, Gray. Pleistocene: Victoria. Living: Queensland and + Victoria. + + _Phascolomys parvus_, Owen. Pleistocene: Queensland. + + _Phascolonus gigas_, Owen. Pleistocene: Queensland, New South Wales + and S. Australia. + + _Macropus titan_, Owen. Pleistocene: Queensland, Victoria, New South + Wales and S. Australia. + + _Macropus anak_, Owen. Pleistocene: Queensland, S. Australia and New + South Wales. + + _Procoptodon goliah_, Owen sp. Pleistocene: Queensland, New South + Wales and Victoria. + + _Sthenurus atlas_, Owen sp. Pleistocene: Queensland, New South Wales, + Victoria, and South Australia. + + _Sthenurus occidentalis_, Glauert. Pleistocene: W. Australia. + + _Palorchestes azael_, Owen. Pleistocene: Queensland, New South Wales + and Victoria. + + _Diprotodon australis_, Owen. Pleistocene: Queensland, New South + Wales, Victoria and S. Australia. + + _Nototherium mitchelli_, Owen. Pleistocene: Queensland, S. Australia + and Victoria. + + _Thylacoleo carnifex_, Owen. Pleistocene: Queensland, New South Wales, + Victoria and W. Australia. + + _Parasqualodon wilkinsoni_, McCoy sp. Cainozoic (Janjukian): Victoria + and Tasmania. + + _Metasqualodon harwoodi_, Sanger sp. Cainozoic (Janjukian): S. + Australia. + + _Kekenodon onamata_, Hector. Cainozoic (Oamaru Series): New Zealand. + + _Cetotolithes nelsoni_, McCoy. Cainozoic (Janjukian): Victoria. + + _Ziphius (Dolichodon) geelongensis_, McCoy. Cainozoic (Janjukian): + Victoria. + + _Scaldicetus macgeei_, Chapm. Cainozoic (Kalimnan): Victoria. + + _Chronozoön australis_, De Vis. Pleistocene: Queensland. + + _Canis dingo_, Blumenbach. Late Pleistocene or Holocene: Victoria. + + _Otaria forsteri_, Lesson. Pliocene (Petane Series): N. Id., New + Zealand. + + _Arctocephalus williamsi_, McCoy. Pleistocene: Victoria. + + + * * * * * + + +LITERATURE. + + +FISHES. + + Silurian.--Chapman, F. Proc. R. Soc. Vict., vol. XVIII. (N.S.), pt. + II. 1906, pp. 93-100, pls. VII. and VIII. (_Thyestes_). + + Devonian.--McCoy, F. Prod. Pal. Vict., Dec. IV. 1876, pp. 19, 20, + pl. XXXV. figs. 7, 7_a_, 7_b_ (_Asterolepis_). Etheridge, + R. jnr. Rec. Austr. Mus., vol. VI. pp. 129-132, pl. XXVIII. + (_Ganorhynchus_). + + Carboniferous and Carbopermian.--Woodward, H. Geol. Mag., Dec. III. + vol. III. 1886, pp. 1-7, pl. I. (_Edestus_). Etheridge, R. + jnr. Geol. and Pal. Queensland, 1892, p. 296, pl. XXXIX. fig. + 1 (_Deltodus_). De Koninck, L. G. Mem. Geol. Surv. New South + Wales, Pal. No. 6, 1898, p. 281, pl. XXIV., fig. 11 (_Tomodus_). + Woodward, A. S. Mem. Nat. Mus. Melbourne, No. 1. 1906 (Mansfield + Series). + + Triassic.--Johnston, R. M. and Morton, A. Proc. R. Soc. Tasmania + (1889), 1890, pp. 102-104; ibid. (1890), 1891, pp. 152-154 + (_Acrolepis_). Woodward, A. S. Mem. Geol. Surv. New South Wales, + Pal. No. 4, 1890 (Gosford). Ibid. No. 10, 1908 (St. Peters). + + Jurassic.--Woodward, A. S. Mem. Geol. Surv. New South Wales, Pal. No. + 9, 1895. Id., Ann. Mag. Nat. Hist., Ser. VII. Vol. XVIII. 1906, + pp. 1-3, pl. I. (_Ceratodus_). Hall, T. S. Proc. R. Soc. Vict. + vol. XII. (N.S.) pt. II. 1900, pp. 147-151, pl. XIV. Chapman, F. + Rec. Geol. Surv. Vict. vol. III. pt. 2, 1912, pp. 234-235, pl. + XXXIX. (_Ceratodus_). + + Cretaceous.--Etheridge, R. jnr. Proc. Linn. Soc. New South Wales, + vol. III. ser. 2, 1889, pp. 156-161, pl. IV. Idem, Geol. and + Pal. Queensland, 1892, pp. 503-504. Davis, J. W. Trans. R. Dubl. + Soc. vol. IV. ser. 2. 1888, pp. 1-48, pls. I.-VII. (Cretaceous + and Cainozoic of New Zealand). Etheridge, R. jnr. and Woodward, + A. S. Trans. R. Soc. Vict., vol. II. pt. II. 1892, pp. 1-7, + pl. I. (_Belonostomus_). Woodward, A. S. Ann. Mag. Nat. Hist., + ser. 6, vol. XIX. 1894, pp. 444-447, pl. X. (_Portheus_ and + _Cladocyclus_). Chapman, F. Proc. R. Soc. Vict., vol. XXI. + (N.S.), pt. II. 1909, pp. 452, 453 (_Corax_). + + Cainozoic.--McCoy, F. Prod. Pal. Vict., Dec. II. 1875, pp. 8-10, pl. + XI. (_Carcharodon_). Chapman, F. and Pritchard, G. B. Proc. R. + Soc. Vict., vol. XVII. (N.S.), pt. I. 1904, pp. 267-297, pls. + V.-VIII. Idem, ibid, vol. XX. (N.S.), pt. I. 1907, pp. 59-75, + pls. V.-VIII. See also Davis, J. W. (_Cretaceous_). + + Pleistocene.--Etheridge, R. jnr. Geol. and Pal. Queensland, 1892, p. + 646 (_Neoceratodus_). Woodward, A. S. Rec. Geol. Surv. New South + Wales, vol. VII. pt. 2, 1902, pp. 88-91, pl. XXIV. (_Ctenolates_). + + +AMPHIBIA. + + Huxley, T. H. Quart. Journ. Geol. Soc., vol. XV. 1859, pp. 647-649, + pl. XXII. figs. 1, 2 (_Bothriceps_). Stephens, W. J. Proc. Linn. + Soc. New South Wales, ser. 2. vol. I. 1886, pp. 931-940. Ibid., + 1887, pp. 1175-1182, pl. XXII. Ibid., vol. II. 1887, pp. 156-158. + Woodward, A. S. Rec. Geol. Surv. New South Wales, vol. VIII. pt. + 4, 1909, pp. 317-319, pl. LI. (_Bothriceps_). + + +REPTILIA. + + Jurassic and Cretaceous.--Hector, J. Trans. N.Z. Inst., vol. VI. 1874, + pp. 333-358. + + Cretaceous.--McCoy, F. Proc. R. Soc. Vic., vol. VIII. pt. I. 1868, + p. 42 (_Plesiosaurus_). Ibid., vol. IX. pt. II. 1869, p. 77 + (_Ichthyosaurus_). Owen, R. Geol. Mag., Dec. I. vol. VII. + 1870, pp. 49-53, pl. III. (_Plesiosaurus_). Id., Quart. Journ. + Geol. Soc. vol. XXXVIII. 1882, pp. 178-183 (_"Notochelys" = + Notochelone_). Etheridge, R. jnr. Proc. Linn. Soc. New South + Wales, ser. 2, vol. III. 1889, pp. 405-413, pls. VII. and VIII. + (_Ichthyosaurus_). Id., Geol. and Pal. Queensland, 1892, pp. + 505-510. Hutton, F. W. Trans. N.Z. Inst. vol. XXVI. 1894, pp. + 354-358, 1 pl. (_Cimoliosaurus_). + + Pleistocene.--Etheridge, R. jnr. Rec. Geol. Surv. New South Wales, + vol. I. pt. 3, 1889, pp. 149-152 (_Miolania_). Id., Geol. and + Pal. Queensland, 1892, pp. 647-653. + + +AVES. + + Miocene.--Huxley, T. H. Quart. Journ. Geol. Soc. vol. XV. 1859, pp. + 670-677. Also Hector, J. Trans. N.Z. Inst. vol. IV. 1872, pp. + 341-346, 1 pl. (_Palaeeudyptes_). Chapman, F. Proc. R. Soc. Vict. + (N.S.) pt. I. 1910, pp. 21-26, pls. IV. and V. + + Pleistocene and Holocene.--Von Haast, J. Trans. N.Z. Inst., vol. IV., + 1872, pp. 192-196; and vol. VI. 1874, pp. 62-75 (_Harpagornis_). + Owen, R. Memoirs on the Extinct Wingless Birds of New Zealand, + London, 1879, 2 vols. De Vis, C. W. Proc. R. Soc. Queensland, + vol. VI. pt. I. 1889, pp. 6-8. Id., Proc. Linn. Soc. New South + Wales, vol. III. ser. 2, 1888, pp. 1277-1292, pls. XXXIII.-XXXVI. + (Carinatae). Etheridge, R. jnr. Rec. Geol. Surv. New South Wales, + vol. I. pt. 2, 1889, pp. 126-136, pls. XI.-XIII. (_Dromornis_). + Id., Geol. and Pal. Queensland, 1892, pp. 653-663. Hutton, F. + W. Trans. N.Z. Inst., vol. XXIV. 1892, pp. 93-172 (Moas). Id., + ibid., vol. XXV. 1893, pp. 14-16, 1 pl. (_Anomalopteryx_). Id., + ibid., vol. XXIX. 1897, pp. 441-557, figs. (Moas). Id., ibid., + vol. XXXVIII. 1906, pp. 66 and 67 (_Emeus crassus_). Hamilton, A. + Ibid, vol. XXVI. 1894, pp. 227-257 (Bibliography of Moas). Ibid, + vol. XXX. 1898, pp. 445 and 446 (_Euryapteryx_). Stirling, E. C. + and Zietz, A. H. C. Mem. R. Soc. S. Austr., vol. I. pt. II. 1900, + pp. 41-80, pls. XIX.-XXIV. (_Genyornis_). Spencer, W. B. Vict. + Nat. vol. XXIII. 1906, pp. 139 and 140; also Spencer, W. B. and + Kershaw, J. A. Mem. Nat. Mus. Melbourne No. 3, 1910, pp. 5-35, + pls. I.-VII. (_Dromaeus minor_). + + +MAMMALS. + + Huxley, T. H. Quart. Journ. Geol. Soc., vol. XV. 1859, pp. 676-677 + (_Phocaenopsis_). McCoy, F. Prod. Pal. Vict., Dec. I. 1874, + pp. 21, 22, pls. III.-V. (_Phascolomys_). Ibid, Dec. II. + 1875, pp. 7-8, pl. XI. and Dec. VI. 1879, pp. 20 and 21, pl. + LV. (_Squalodon_). Ibid, Dec. III. 1876, pp. 7-12, pl. XXI. + (_Thylacoleo_). Ibid, Dec. IV. 1876, pp. 7-11, pl. XXXI.-XXXIII. + (_Diprotodon_). Ibid, Dec. V. 1877, pp. 7-9, pl. XLI. and + XLII. (_Arctocephalus_). Ibid, Dec. VI. 1879, pp. 5-7, pl. LI. + (_Macropus_): pp. 9-11, pl. LI.-LIII. (_Procoptodon_): pp. 13-17, + pl. LIV. (_Cetotolithes_); pp. 19 and 20, pl. LV. (_Physetodon_). + Ibid, Dec. VII. 1882, pp. 7-10, pl. LX. (_Canis dingo_): pp. + 11-13, pl. LXXII. and LXII. (_Sarcophilus_): pp. 23-26, pl. + LIX. (_Ziphius_). Owen, R. Extinct Mammals of Australia, London + 1877, 2 vols. Hector, J. Trans. N.Z. Inst., vol. XIII. 1881, pp. + 434-436, 1 pl. (_Kekenodon_). Lydekker, R. Cat. Foss. Mammalia, + Brit. Mus. part V. 1887. Id., Handbook to the Marsupialia, and + Monotremata. Allen's Nat. Library, 1894, pt. III. pp. 249-286. + De Vis, C. W. Proc. Linn. Soc. New South Wales, vol. VIII. pt. + 3, 1883, p. 395 (Sirenian). Id., ibid, vol. X. 1895, pp. 75-133, + pls. XIV.-XVIII. (Macropodidae). Id., Proc. R. Soc. Vict., vol. + XII. (N.S.), pt. I, 1899, pp. 107-11 (Marsupials). Etheridge, R. + jnr. Geol. and Pal. Queensland, 1892, pp. 663-683 (Pleistocene + Mammals). Dun, W. S. Rec. Geol. Surv. New South Wales, vol. III. + pt. 4, 1893, pp. 120-124, pl. XVI. (_Palorchestes_). Ibid., vol. + IV. pt. 3, 1895, pp. 118-126, pls. XI. and XII. (Monotremes). + Stirling, E. C. and Zietz, A. H. C. Mem. Roy. Soc. S. Australia, + vol. I. pt. I. 1899 (Descr. of _Diprotodon_, Manus and Pes.). + Spencer, W. B. Proc. Zool. Soc. 1900, pp. 776-794, pls. XLIX. + and L. (_Wynyardia_). Hall, T. S. Proc. R. Soc. Vict. vol. + XXIII. (N.S.), pt. II. 1911, pp. 257-265, pl. XXXVI. (Rev. of + Squalodontidae). Spencer, W. B. and Walcott, R. H. Proc. R. + Soc. Vict., vol. XXIV. (N.S.), pt. I. 1912, pp. 92-123, pls. + XXXVI.-XXIX. (_Thylacoleo_). Chapman, F. Rec. Geol. Surv. Vict., + vol. III. pt. 2, 1912, pp. 236-238, pl. XL. (_Scaldicetus_). + Woods, J. E. T. Geol. Observations in S. Australia, 1862, pp. 329 + and 330 (Human Remains): also Krefft, G. Australian Vertebrata, + Recent and Fossil, 1867, p. 91; Etheridge, R. jnr. Rec. Geol. + Surv. New South Wales, vol. III. pt. 4, 1893, pp. 128-132; + Etheridge, R. jnr. and Trickett, O. Ibid., vol. VII. pt. 4, 1904, + pp. 325-328. + + + + +APPENDIX.--ON THE COLLECTION AND PRESERVATION OF FOSSILS. + + +The tools and other paraphernalia necessary for fossil collecting +are fortunately within the reach of all. The principal of these is a +geological hammer, preferably with a pick at one end of the head and the +opposite end square-faced. The pick end is useful for digging out fossils +from soft clays, or for extracting a block of fossils entire. The square +end is employed for breaking up the slabs or masses containing fossils. To +get good results, much will of course depend upon one's skill in striking +the right face of a block. If bedding planes are present on the lump from +which we wish to extract our fossils, it will be well to strike at right +angles to these layers in order to split them asunder, thus exposing a +shell-layer corresponding to the original surface of the ancient sea-bed +upon which the organisms accumulated. In some cases the splitting of +fossiliferous rocks may be best carried out with the pick end, provided it +be not too sharply curved. The hammer should be faced with steel, for many +fossiliferous rocks, especially compact limestones, are apt to severely +try the temper of an ill-made tool. + +A chisel, of chilled steel, should accompany the hammer, since this is +often of the greatest use in working out large fossils, more particularly +those that are buried in a cliff or quarry face. The process of extracting +difficult specimens should never be hurried, for one often gets +surprisingly good results with a little extra care. + +A strong pocket knife may be used in trimming specimens and partially +cleaning shells that can be safely manipulated on the spot, but the final +cleaning should be left until the return home. The knife is also useful +for cleaning slates and shales, since the chisel-edge is frequently a +trifle too thick for this kind of work. + +For the more delicate fossils, means for careful packing should be +provided; chip-boxes and cotton-wool being indispensable for the smaller +specimens. A ready method of packing the fossils obtained from the +friable, sandy tertiary deposits is to store them in tins, the contents of +which can be firmly secured from rattling by filling up with sand. This +sand, however, should be taken from the same bed in which the fossils +occur, so as to get no admixture of the smaller shells from another +formation or deposit; for although we may not wish to examine the finer +material ourselves, it will yield in many cases a rich harvest to our +microscopical friends, such residues containing microzoa, as shells +of foraminifera, polyzoa and carapaces of the ostracoda. The residues +referred to may be obtained from many of our marls and rubbly limestones +by the simple process of washing in water, and repeatedly pouring off the +finest clayey mud, until only a sandy deposit remains, which can then be +dried and sorted over by the aid of a lens or low power microscope. + + +=Hints on Fossil Collecting.--= + +As regards the places most suitable for collecting fossils, the Cainozoic +beds are perhaps, the most accessible to a beginner, especially in +Victoria. For instance, the cliff exposures at Beaumaris, Port Phillip, +will afford a plentiful supply of the little heart-shaped sea-urchin, +_Lovenia_, and an occasional _Trigonia_ and _Limopsis_, as well as +many other fossils of the great group of the shell-fish or mollusca. +The richest bed containing the sharks' teeth at the above locality is +almost perpetually covered with a bed of shingle, but can be reached by +digging at the cliff-base. Isolated specimens, however, although rather +the worse for wear, may often be picked up amongst the shingle, having +been washed up from the foreshore by the tide. An enticing band of large +bivalve shells (_Dosinea_), can be seen halfway up the cliffs, near the +baths at this locality, but are somewhat disappointing, for when obtained +they crumble to pieces in the hand, since their shells are composed of +the changeable form of carbonate of lime called aragonite, which has +decomposed in place in the bed, after the shells were covered up by the +deposit. + +Good collections of shells of the Balcombian series may be easily made at +Balcombe's Bay and Grice's Creek, Port Phillip. They can there be dug out +of the grey-blue clay with a knife, and afterwards cleaned at leisure by +means of a soft tooth brush dipped in water. In the cement stone at the +same place there are numerous shells of pteropods or "sea-butterflies" +(_Vaginella_), and specimens of the stone may be obtained, showing myriads +of the porcelain-like shells, and also their internal casts in the hard +greenish coloured matrix. + +The ferruginous or ironstone beds seen in the Flemington Railway cutting, +Melbourne, is an old marine shell-bank, resting on basalt. The shells have +all been dissolved away, and only their casts and moulds remain. These +impressions are, however, so faithfully moulded that the ornamentation +of each shell can often be reproduced on a squeeze taken with a piece of +modelling wax or plasticine. Such fossil remains are easily collected by +carefully breaking up the blocks of ironstone with a hammer. + +Quarries in the older limestones and mudstones in Victoria, New South +Wales and other States, are often good hunting grounds for fossils. The +quarry at Cave Hill, Lilydale, for example, will be found very profitable, +for the limestone is full of corals and molluscan shells; whilst the +friable or rubbly portion is worth breaking down for the smaller fossils. +The bed-rock (Silurian) of Melbourne is in places very fossiliferous; +the sandstones of Moonee Ponds Creek generally affording a fair number +of brachiopods, and occasionally corals. The mudstones of South Yarra, +Studley Park, Yan Yean, and other places on the same geological horizon, +contain a rich fauna, to be obtained only by the assiduous collector +who will search over and break up a large number of blocks. Practice +in this work makes a good collector; although of course one must know +something about the objects looked for, since many apparently obscure +fossil remains of great interest might easily be passed over for lack +of knowledge as to what should be expected to occur at each particular +locality. + +Many other good collecting grounds might here be alluded to, but we have +purposely cited only a few near Melbourne, since a selection from other +parts of Australasia may easily be made from the localities mentioned in +connection with the various groups of fossils dealt with in the systematic +portion of this work. + + +=Preservation of Fossils.--= + +Many of the Cainozoic fossils from the shelly sands and clays are +extremely delicate, owing in some cases to their being imperfectly +preserved, seeing that they frequently contain in their shell-structure +layers of the unstable form of carbonate of lime called aragonite. Fossils +containing aragonite are:--Calcareous Sponges; Corals; Bivalved shells, +except Oysters, Pectens, and the outer layer of _Spondylus_, _Pinna_, and +_Mytilus_; Gasteropods (with a few exceptions); and Cephalopods. In some +of these, however, a transformation of the aragonite into calcite enables +the fossil to be permanently preserved. The delicate fossils referred to +should be dipped in weak glue or gelatine and left to dry; after which +their final cleaning can be done with the aid of a little warm water and a +soft brush. + +Certain of the clays and mudstones, both of Cainozoic and Jurassic ages +which show remains of plants, such as leaves and fern fronds, are often +best treated with a thin surface layer of paper varnish, before they +lose the natural moisture of the rock; for when they become perfectly dry +the thin carbonaceous film representing the original leaf-substance peels +off, and the fossil is consequently destroyed. A method of treatment for +Cainozoic leaves, by dipping them in warm vaseline and brushing off the +superfluous material, has been described by Mr. H. Deane. + + +=Storing Fossils for Reference.--= + +Fossils specimens are generally best displayed in cardboard trays; or if +thin wooden paper-covered tablets are used, say of about 3-16in. thickness +and cut to proportionate sizes, the fossils should be held in place by +pins for easy removal, unless more than one example can be shown together, +exhibiting all aspects, when they can be secured to the tablet by a touch +of seccotine. The smaller shells may be displayed in glass topped boxes, +which in turn may be stuck down to tablets or placed in trays. + + + + +INDEX. + + + Aboriginal implements, 303 + Aborigines, probable origin of, 302 + _Acanthias_, 270 + _Acanthodes_, 261 + _Acanthosphaera_, 103 + _Acanthothyris_, 166, 167 + _Acentrophorus_, 263 + _Acrolepis_, 263 + _Actaeon_, 197 + _Actinoceras_, 205, 207 + _Actinocrinus_, 136 + _Actinodesma_, 178, 179 + _Actinopteria_, 178, 179 + _Actinostroma_, 121, 122 + _Adeona_, 158 + _Aechmina_, 237 + _Aeschna_, 250 + _Aetheolepis_, 267 + _Agathiceras_, 207 + AGNATHA, 258 + _Agnostus_, 227 + _Allodesma_, 176 + _Ambonychia_, 177 + _Ammodiscus_, 96, 97 + _Ammonites_, 204, 209, 210 + AMMONOIDEA, 205 + _Amoeba_, 36, 95 + AMPHIBIA, structure of, 272 + _Amphistegina_, 100 + _Amplexus_, 117 + _Ampyx_, 229 + _Amusium_, 185 + _Anas_, 283 + _Anchura_, 197 + _Ancilla_, 198, 199, 202 + _Ancyloceras_, 209, 210 + ANGIOSPERMEAE, characters of, 40 + ANNELIDA, 152 + _Anomalina_, 98 + _Anomalopteryx_, 283 + _Antedon_, 138 + ANTHOZOA, 64, 113 + Antiquity of man in Australia, 304 + _Aparchites_, 237 + _Apateolepis_, 262 + _Aphnelepis_, 267 + _Apocynophyllum_, 91 + _Aptornis_, 283 + _Aptychopsis_, 246 + _Arabellites_, 153 + _Arachnoides_, 146 + _Araucarioxylon_, 68 + _Araucarites_, 89 + _Arca_, 184, 186, 188 + _Archaeocidaris_, 144 + _Archaeocyathina_, 113 + ARCHAEOCYATHINAE, 112 + _Archaeomaene_, 267 + _Archaeopteryx_, 280 + _Arctocephalus_, 299 + _Arenicolites_, 153 + Argillaceous rocks, 69 + _Argilloecia_, 237 + _?Argiope_, 166 + _Argonauta_, 205 + ARTHROPODA, structure and subdivisions of, 38, 220 + _Asaphus_, 227, 228 + _Aspidorhynchus_, 267 + _Astarte_, 182 + _Asteracanthus_, 269, 271 + ASTEROIDEA, 139 + _Asterolepis_, 258 + _Astralium_, 198, 200 + _Astropecten_, 141 + _Athyris_, 161, 162, 165 + _Atrypa_, 158, 160, 162 + _Aturia_, 210 + _Atys_, 204 + _Aucella_, 183 + _Aulopora_, 116 + Australian fossiliferous strata, 45-48. + AVES, 280 + Aviculopecten, 179, 180 + _Axopora_, 119 + + + Bactronella, 112 + _Baculites_, 210 + _Baiera_, 89, 164 + _Bairdia_, 240 + _Balanophyllia_, 118 + _Balanus_, 243 + Balcombian bivalves, 186 + " gasteropods, 199 + Bandicoot, 289, 295 + _Bankivia_, 201 + _Banksia_, 91, 281 + _Barbatia_, 184, 185 + Barnacles, 240 + _Barnea_, 187 + _Bathytoma_, 201 + _Bela_, 201 + _Belemnites_, 205, 209, 210 + BELEMNOIDEA, 205 + _Bellerophon_, 193, 194, 195, 196 + _Belonorhynchus_, 262 + _Belonostomus_, 267 + _Bettongia_, 295 + _Beyrichia_, 235, 236, 237 + _Biloela_, 274 + _Bipora_, 158 + Birds, fossil, 53, 280 + _Biziura_, 283 + BLASTOIDEA, distribution and characters of, 61, 138 + Blue-green Algae, 76, 82 + Bog iron-ore, 80 + _Bolodon_, 286 + _Bombax_, 91 + Bone-beds, 78 + Bone-breccias, 79 + _Bothriceps_, 273 + _Botryocrinus_, 136 + BRACHIOPODA, structure of, 57, 158 + Brachiopod limestone, 74 + _Brachymetopus_, 232 + _Brachyphyllum_, 89 + Bracken fern, 91 + _Brissopsis_, 148 + Brittle-stars, characters of, 61, 141 + _Bronteus_, 229, 230 + _Bryograptus_, 124, 126, 227 + BRYOPHYTA, characters of, 39 + _Buccinum_, 191 + _Buchozia_, 199 + _Bulimina_, 97, 98 + _Bulinus_, 69, 191 + _Bulla_, 204 + _Bullinella_, 198, 199 + _Bythocypris_, 236 + _Bythotrephis_, 82 + + + Cainozoic Balanidae, 243 + " bird, Victoria, 281 + " bivalves, 184 + " brachiopods, 166 + " brittle-stars, 143 + " chitons, 190 + " corals, 118 + " crabs, 247 + " echinoids, irregular, 146 + " echinoids, regular, 145 + " fishes, 269 + " Foraminifera, 99 + " gasteropods, 198 + " gasteropods, New Zealand, 202 + " Holothuroidea, 148 + " insects, 250 + " Lepadidae, 243 + " Ostracoda, 239 + " and Pleistocene reptiles, 279 + " plants, 89 + " Polyzoa, 158 + Cainozoic Radiolaria, 104 + " scaphopods, 189 + " sponges, 110 + " starfishes, 141 + " strata, 45, 46 + Calcareous rocks, 72 + " sponges, 112 + _Callograptus_, 122 + _Callorhynchus_, 269 + _Calymene_, 229, 230, 231 + CALYPTOBLASTEA, 122 + _Calyptraea_, 198, 200, 201 + _Camarotoechia_, 160, 161, 162 + Cambrian bivalves, 177 + " brachiopods, 159 + " crinoids, 134 + " Foraminifera, 96 + " gasteropods, 192 + " Ostracoda, 235 + " plants, 82 + " Radiolaria, 102 + " sponges, 107 + _Cameroceras_, 207 + _Campanularia_, 122 + _Campophyllum_, 115, 117 + _Cancellaria_, 198, 199, 202 + _Canis_, 298 + Cannel coal, 76 + _Capitosaurus_, 274 + _Capulus_, 194 + Carbonaceous rocks, 76 + Carboniferous brachiopods, 162 + " crinoids, 136 + " fishes, 259 + " Foraminifera, 96 + " gasteropods, 196 + " Ostracoda, 237 + " plants, 85 + Carbopermian bivalves, 179 + " blastoids, 139 + " brachiopods, 163 + " cephalopods, 207 + " corals, 116 + " crinoids, 137 + " fishes, 261 + " Foraminifera, 97 + " gasteropods, 196 + " labyrinthodonts, 273 + " Ostracoda, 237 + " palaeechinoids, 144 + " Phyllopoda, 233 + " plants, 86 + " sponges, 110 + " starfishes, 141 + " trilobites, 232 + _Carcharodon_, 269, 270, 271 + _Carcharoides_, 269 + _Cardiola_, 177, 178 + _Cardita_, 184, 187 + _Cardium_, 176, 184, 186, 187 + CARNIVORA, 298 + _Carposphaera_, 102 + _Carpospongia_, 109 + _Caryocaris_, 244, 246 + _Cassidulus_, 148 + _Catenicella_, 158 + _Cellaria_, 158 + _Cellepora_, 158 + _Cenellipsis_, 102 + _Cenosphaera_, 102, 103 + CEPHALOPODA, characters of, 204 + _Ceratiocaris_, 246 + _Ceratodus_, 265, 267 + _Ceratotrochus_, 118 + _Cerithiopsis_, 200 + _Cerithium_, 198, 200 + _Cestracion_, 261, 269, 271 + CETACEA, 295 + _Cetotolithes_, 296 + _Chaenomya_, 181 + CHAETOPODA, 152 + _Chama_, 185 + Changes of climate in the past, 31 + CHEILOSTOMATA, 155, 157 + _Cheirurus_, 229, 231 + _Chelodes_, 190 + Cherts, 71 + _Chione_, 185, 187, 188 + _Chiridota_, 148 + _Chironomus_, 250 + _Chiton_, 190 + _Chonetes_, 160, 161, 162 + CHORDATA, 257 + _Chosornis_, 283 + _Chronozoön_, 298 + _Cicada_, 250 + _Cidaris_, 145 + _Cimoliosaurus_, 279 + _Cinnamomum_, 91 + _Cinulia_, 197 + CIRRIPEDIA, habits and structure of, 240 + _Cladochonus_, 117 + _Cladophlebis_, 89, 164, 182 + CLADOPHORA, 122 + Classification of animals, 35 + _Clathrodictyon_, 121 + _Clausilia_, 191 + _Clavigera_, 165 + Clays, 69 + _Cleiothyris_, 164 + _Cleithrolepis_, 262, 263, 274 + _Climacograptus_, 127 + _Climatius_, 258 + _Clonograptus_, 123, 124, 126 + _Clypeaster_, 146 + _Cnemiornis_, 283 + Coals, 76 + _Coccolepis_, 267 + _Cocconema_, 92 + _Coccosteus_, 259 + COELENTERATA, characters of, 37 + _Coleolus_, 193 + Collecting fossils, 317 + _Colubraria_, 199 + _Columbarium_, 198, 201, 202 + _Columbella_, 198 + _Conchothyra_, 184 + _Conocardium_, 177, 178 + Conodonts, 153 + _Conosmilia_, 118 + _Conularia_, 193, 194, 196 + _Conus_, 198, 199, 202, 204 + _Coprosmaephyllum_, 90 + Coral limestone, 73 + Corals, 64, 113 + _Corax_, 267 + _Corbicula_, 182 + _Corbula_, 177, 185, 187, 188 + _Cordaites_, 85 + _Cornulites_, 154 + _Coscinocyathus_, 113 + _Coxiella_, 69 + _Crassatellites_, 176, 184 + _Crenella_, 176 + _Crepicephalus_, 227 + _Crepidula_, 198 + Cretaceous (Lower and Upper) cephalopods, 209 + " cephalopods, New Zealand, 210 + " Cheilostomata, 157 + " crinoids, 137 + " echinoids (irregular), 146 + " (Lower) fishes, 267 + " fishes, New Zealand, 268 + " Foraminifera, 98 + " gasteropods, 197 + " plants, 89 + " Radiolaria, 103 + " (Lower) reptiles, 277 + " reptiles, New Zealand, 279 + " scaphopods, 189 + " sponges, 110 + Crinoidal limestone, 74 + CRINOIDEA, occurrence and structure of, 61, 133 + _Crioceras_, 209 + _Crisia_, 158 + _Cristellaria_, 98 + _Crocodilus_, 279 + _Cromus_, 229 + Crustacea, an archaic group, 221 + " development of, 221 + " fossil, 54 + _Cryptodon_, 186 + _Cryptograptus_, 127 + _Cryptoplax_, 190 + _Cryptostomata_, 155, 156 + _Ctenodonta_, 177, 178 + _Ctenodus_, 261, 263 + _Ctenolates_, 272 + _Ctenostreon_, 182 + _Cucullaea_, 182, 184, 185 + _Cultellus_, 188 + _Cuna_, 184, 186, 187 + _Cupressinoxylon_, 78, 89 + _Cupressus_, 91 + _Cuscus_, 295 + Cuttle-fishes, 205 + CYANOPHYCEAE, 82 + _Cyathocrinus_, 137 + _Cyathophyllum_, 113, 115, 117 + _Cyclas_, 69 + _Cycloceras_, 206 + _Cyclolituites_, 207 + _Cyclometopa_, 248 + _Cyclonema_, 194 + CYCLOSTOMATA, 155 + _Cydnus_, 250 + _Cymbella_, 92 + _Cyphaspis_, 229 + _Cyphon_, 250 + _Cypraea_, 191, 198, 199, 200, 202 + _Cypricardinia_, 178 + Cyprid limestone, 75 + _Cyrenopsis_, 184 + _Cyrtoceras_, 204, 207 + _Cyrtograptus_, 128 + _Cyrtina_, 162, 164 + _Cyrtolites_, 193 + Cystideans, 61 + _Cystiphyllum_, 116 + _Cythere_, 239, 240 + _Cytherella_, 240 + _?Cytheridea_, 238 + _Cytheropteron_, 239 + + + _Dadoxylon_, 68 + _Dalmanites_, 224, 225, 229, 231 + _Daonella_, 182 + Darter, 283 + _?Darwinula_, 238 + _Dasyurus_, 287, 295 + DECAPODA, 246 + Deep Leads, fruits of, 91 + " insects from, 250 + _Deltodus_, 261 + _Deltopecten_, 180 + _Dendrocrinus_, 134, 135 + _Dendrocygna_, 283 + _Dendrograptus_, 122 + _Dendrophyllia_, 119 + _Dennantia_, 198 + _Dentalium_, 189 + Dentition of Reptiles, 275 + _Deontopora_, 120 + _Desmoceras_, 209 + Devonian bivalves, 178 + " brachiopods, 161 + " cephalopods, 207 + " corals, 115 + " crinoids, 136 + " fishes, 258 + " gasteropods, 195 + " Ostracoda, 237 + " plants, 85 + " Radiolaria, 102 + " scaphopods, 189 + " stromatoporoids, 121 + " trilobites, 231 + DIADACTYLA, 287 + Diatomite, 72 + Diatoms, 92 + _Dicellograptus_, 126, 127 + _Dichograptus_, 126 + _Dicranograptus_, 126, 127 + _Dictyonema_, and allies, 122 + _Dictyopyge_, 262 + _Didymograptus_, 124, 126 + _?Didymosorus_, 89 + _Dielasma_, 164, 165 + _Dikellocephalus_, 227 + _Dimetrodon_, 276 + _Dimya_, 184, 185, 186 + _Dinesus_, 227 + Dingo, 298, 305 + _Dinornis_, 281, 282, 283, 299 + _Diodon_, 270, 271 + _Dione_, 188 + _Diphyphyllum_, 113 + _Diplograptus_, 124, 126, 127, 128 + _Diprotodon_, 51, 290, 293 + _Diprotodon_-breccias, 203 + DIPROTODONTIA, 287 + _Discina_, 166 + _Discorbina_, 98 + _Dissocheilus_, 199 + _Dithyrocaris_, 246 + _Ditrupa_, 154 + _Ditrupa_ limestone, 74 + _Dolichodon_, 296 + _Dolichometopus_, 226 + _Dolium_, 201 + _Donax_, 175, 187 + _Dorsetensia_, 209 + _Dosinea_, 185, 188 + _Drillia_, 198, 202 + _Dromaeus_, 282, 283 + _Dromornis_, 282 + Duck, 283 + _Duncaniaster_, 147 + + + Ear-bones of whales, 296 + Early observers, 24 + _Eburnopsis_, 199, 200 + _Echidna_, 286, 287 + _Echinocyamus_, 146 + ECHINODERMATA, characters of, 37, 59 + " divisions of, 133 + ECHINOIDEA, 143 + _Echinolampas_, 147, 148 + _Echinoneus_, 147 + _Echinus_, 145 + _Ecionema_, 112 + _Edaphodon_, 271 + _Edestus_, 262 + _Edmondia_, 177, 180, 182 + _Eglisia_, 202 + Elephant-fish, 269, 271 + Elephant-tusk shells, 188 + Elevated sea-beds, 27 + _Elonichthys_, 261, 263 + _Elpisopholis_, 263 + _Emarginula_, 198 + Emu, 283 + _Encrinurus_, 229 + _Endoceras_, 205 + _Endothyra_, 96, 98 + _Entalophora_, 158 + _Entomis_, 238 + _Ephemera_, 250 + _Equisetites_, 40 + Errant worms, 153 + _Erycina_, 187 + _Erymnoceras_, 209 + _Estheria_, 233 + _Eucalyptus_, 90, 91, 281 + _Eulima_, 198 + _Eunema_, 193 + _Eunicites_, 153 + _Euomphalus_, 194, 195, 196 + _Eupatagus_, 147 + _Euphemus_, 196 + _Eurydesma_, 181 + EURYPTERIDA, 248 + _Euthria_, 198 + _Eutrochus_, 200 + Evolution of life-forms, 33 + + + _Fagus (Notofagus)_, 91 + Falcon, 283 + _Fasciolaria_, 198, 199 + _Favosites_, 73, 114, 115, 116 + Feather-star, 138 + _Fenestella_, 156, 157 + _Fibularia_, 146 + Fishes, fossil, 53 + " primitive types, 258 + " true, 258 + Fish-lizards, 275, 276, 277, 278 + _Fissilunula_, 183, 184 + _Fissurellidea_, 198 + _Fistulipora_, 155, 156 + _Flabellina_, 98 + _Flabellum_, 118, 119 + Flightless pigeon goose, 283 + Flints, 71 + Flying phalanger, 295 + Foraminifera, characters of, 36, 95 + " fossil, 65 + Foraminiferal limestone, 73 + Fossil faunas, differences in, 43 + Fossiliferous strata, Australia, 45-48 + " strata, New Zealand 49 + Fossil, origin of name, 23 + Fossils an index to age of strata, 26, 32 + " nature of, 21 + " petrifaction of, 23 + " preservation of, 23 + " structure preserved in, 24 + Fossil wood, 24, 66, 68 + _Frondicularia_, 97, 98 + Fruits of the deep leads, 91 + _Fulica_, 283 + _Fusus_, 198, 201 + + + _Galeocerdo_, 269, 271 + _Gallinula_, 283 + _Gangamopteris_, 86 + _Ganorhynchus_, 259 + _Gari_, 185 + GASTEROPODA, characters of, 190 + _Gastrioceras_, 207 + _Geinitzina_, 98 + _Genyornis_, 282, 302 + Geological epochs, 45-49 + Geology, scope of, 21 + Giant kangaroo, 289 + " lizard, 280 + " penguin, 280 + _Gibbula_, 198 + _Ginkgo_, 89, 91 + _Girvanella_, 76, 82, 86 + Glauconite casts of foraminifera, 96 + _Glossograptus_, 126, 127 + _Glossopteris_, 86 + _Glycimeris_, 184, 187 + _Glyphioceras_, 207 + _Gomphonema_, 93 + Gondwana-Land, 87 + _Goniatites_, 207, 208 + _Goniograptus_, 124, 126 + _Gosfordia_, 262 + _Gosseletina_, 196 + _Grammysia_, 177 + _Granatocrinus_, 139 + _Graphularia_, 118, 119 + Graptolites, Bendigo series, 124 + " Lancefield series, 124 + " nature of, 63, 123 + " Tasmania, 128 + GRAPTOLITOIDEA, 123 + _Gregoriura_, 142 + _Griffithides_, 232 + _Gromia_, 95 + Ground pigeon, 283 + _Gryphaea_, 182 + _Grypotherium_, 53 + Guide fossils, 43 + GYMNOSPERMEAE, characters of, 40 + _Gyracanthides_, 261 + _Gyroceras_, 207 + _Gyrodoma_, 194 + + + _Halimeda_ limestone, 75 + _Haliotis_, 198, 200 + _Haliserites_, 83 + _Halysites_, 114 + _Hamites_, 210 + _Hapalocrinus_, 136 + _Haploceras_, 209 + _Haplophragmium_, 97, 98 + _Harpa_, 198, 199, 201 + _Harpactocarcinus_, 248 + _Harpagornis_, 283 + _Hawk_, 283 + _Helicocrinus_, 136 + _Helicotoma_, 195 + _Heliolites_, 115, 116 + _Heliopora_, 115 + _Heliosphaera_, 103 + _Helix_, 203 + _Hemiaster_, 148 + _Hemipatagus_, 148 + _Heterocrinus_, 135 + HETEROPODA, 190 + _Heteropora_, 158 + Hexactinellid sponge, 107, 110 + Hinge-structure, in bivalves, 175 + _Holaster_, 147 + HOLOTHUROIDEA, 148 + _Homalonotus_, 229, 231 + _Hornera_, 158 + _Huenella_, 159 + Human remains, sub-recent, 299 + _Hyalostelia_, 108, 110 + _Hybocrinus_, 135 + _Hydractinia_, 119, 120 + HYDROZOA, 63, 119 + _Hymenocaris_, 244 + _Hyperammina_, 97 + _Hyolithes_, 192, 193, 194 + _Hypothyris_, 164 + _Hypsiprymnus_, 295 + + + Ibis, 283 + _Ichthyosaurus_, 276, 277, 278 + _Idiostroma_, 121 + _Idmonea_, 158 + _Illaenus_, 229 + Indusial limestone, 75 + _Inoceramus_, 183, 184 + Insects, 53, 250 + Ironstone, 80 + Irregular echinoids, 146 + _Ischnochiton_, 190 + _Ischyodus_, 269, 270 + _Isochilina_, 237 + _Isocrinus_, 137, 138 + + + Janjukian bivalves, 186 + " gasteropods, 200 + _Jonesina_, 237 + Jurassic bird, 280 + " bivalves, 182 + " brachiopods, 165 + " cephalopods, 208 + " fishes, 264 + " Foraminifera, 98 + " gasteropods, 196 + " insects, 250 + " Ostracoda, 238 + " Phyllopoda, 233 + " plants, 89 + " reptiles, 276 + " scaphopods, 189 + + + Kalimnan bivalves, 187 + " gasteropods, 201 + Kangaroo, 295 + _Keeneia_, 196 + _Kekenodon_, 295, 296 + Kerosene shale, 77 + _Kionoceras_, 206 + _Kloedenia_, 237 + + + _Labrodon_, 271 + LABYRINTHODONTIA, 272 + _Lagena_, 98 + _?Lagria_, 250 + _Lamna_, 267, 269, 271 + Lamp-shells, 57, 158 + _Lasiocladia_, 110 + _Lasiograptus_, 126, 127 + _Latirus_, 198, 201 + _Laurus_, 91 + _Leaia_, 233 + Leda, 182, 184, 185, 187, 188 + Leonardo da Vinci, 25 + _Lepas_, 243 + _Leperditella_, 234 + _Leperditia_, 233, 234, 235, 237, 238 + _Lepidocyclina_, 99, 100 + " limestone, 73 + _Lepidodendron_, 40, 85, 261 + " beds, 162 + _Lepralia_, 157, 158 + _Leptaena_, 162, 164 + _Leptoclinum_, 257, 258 + _Leptodesma_, 179 + _Leptodomus_, 177 + _Leptograptus_, 124 + _Leptolepis_, 264, 265, 267 + _Lepton_, 187 + _Lichas_, 229 + _Lichenopora_, 158 + _Lieberkuehnia_, 95 + _Lima_, 184, 185, 186 + _Limatula_, 185 + Limestones formed by organisms, 72 + _Limnaea_, 69 + _Limopsis_, 184, 185, 187 + _Limulus_, 248 + _Lingula_, 160, 162, 166, 261 + _Linthia_, 147, 148 + _Liopyrga_, 201 + _Liotia_, 198, 200 + Lithistid sponges, 109, 110 + Lithological evidence, value of, 44 + _Lithophaps_, 283 + _Lithothamnion_, 75 + _Lituites_, 207 + _Lituola_, 97 + _Loganograptus_, 126 + _Lophophyllum_, 117 + _Lorica_, 190 + _Lotorium_, 198, 200, 202 + _Lovenia_, 147 + Lower Cambrian trilobites, 226 + " Cretaceous bivalves, 183 + " " brachiopods, 166 + " " cephalopods, 209 + " " crab, 246 + " " dragon-fly, 250 + " " fishes, 267 + " " reptiles, 277 + " Mesozoic fishes, 263 + " Ordovician graptolites, New Zealand, 126 + " Ordovician graptolites, Victoria, 124 + _Loxoconcha_, 239 + _Loxonema_, 193, 194, 195, 196 + _Lucina_, 185, 187 + Lung-fish, 265 + _Lunucammina_, 98 + _Lunulicardium_, 178 + _Lunulites_, 158 + _Lyriopecten_, 179 + + + _Maccoyella_, 183, 184 + _Macrocephalites_, 209 + _Macrocheilus_, 196 + _Macrocypris_, 236, 240 + _Macropora_, 158 + _Macropus_, 289, 295 + _Macrotaeniopteris_, 88 + _Mactra_, 177, 185, 188 + Madrepore limestone, 73 + _Magasella_, 166, 168 + _Magellania_, 166, 167, 168 + _Magnolia_, 91 + Maiden-hair tree, 89 + Mail-shells, 189 + MAMMALIA, early types, 285 + Mammals, fossil, 51 + Manatees and dugongs, 298 + _Marginella_, 198, 199 + _Marginulina_, 98 + Marsupial lion, 293 + Marsupial, oldest known Australian, 294 + Marsupials, 287 + " Pleistocene and living, 295 + _Martiniopsis_, 164 + _Mastodonsaurus_, 274 + Material for fossil collecting, 315 + _Megalania_, 280 + _Megalosaurus_, 277 + _Melania_, 203 + _Melosira_, 92 + _Membranipora_, 157, 158 + _Meretrix_, 177, 185, 187 + _Mesoblastus_, 139 + _Mesostigmodera_, 250 + Mesozoic strata, 46 + _Metablastus_, 139 + _Metasqualodon_, 295, 296 + METAZOA, 95 + _Micraster_, 146 + _Microdiscus_, 227 + _Mikrogromia_, 95 + _Millepora_, 119 + _Milleporids_, 119 + _Miliolina_, 96, 100, 101 + Miocene bird, New Zealand, 280 + " leaf-beds, 90 + Miolania, 279 + Mitra, 198, 199, 204 + Moa-birds, 281-285, 299 + _Modiola_, 183, 186 + _Modiolaria_, 186 + _Modiolopsis_, 177 + MOLLUSCA, characters of, 38, 56, 174 + MOLLUSCOIDEA, characters of, 38, 57, 154 + Monactinellid sponges, 109, 110 + _Monogenerina_, 97 + _Monograptus_, 124, 128 + _Monostychia_, 146 + _Monotis_, 182 + MONOTREMATA, 286 + _Monticulipora_, 155 + Monticuliporoids, 117 + _Montlivaltia_, 118 + Moor-hen, 283 + _Mopsea_, 119 + _Morio_, 198, 200 + Mound-builders, 283 + _Mourlonia_, 196 + Mud-fish, 265, 267 + Muds, 69 + Mudstone, 70 + MULTITUBERCULATA, 286 + _Murchisonia_, 104, 195, 196 + _Murex_, 198, 199, 200 + _Myodora_, 185, 187 + _Myriolepis_, 262, 263 + _Mytilarca_, 177 + _Mytilus_, 182, 183, 187, 188 + + + Naming of animals, 34 + _Nassa_, 191, 198, 204 + _Natica_, 191, 197, 198, 200, 201 + Native cat, 287, 295 + " dog, 298 + " honeysuckle, 91, 92 + NAUTILOIDEA, 204 + _Nautilus_, 204, 207, 209, 210 + _Navicula_, 92 + _Nebalia_, 244 + _Necrastur_, 283 + _Neoceratodus_, 267 + Newer Pliocene seal, 299 + _Newtoniella_, 198 + New Zealand fossiliferous strata, 49 + _Niso_, 194, 198 + _Nodosaria_, 98, 100 + _Nonionina_, 96 + _Normanites_, 209 + _Notasaphus_, 227 + _Notidanus_, 268, 269, 270, 271 + _Notochelone_, 53, 277 + _Notophyllia_, 118 + _Nototherium_, 293 + _Nubecularia_, 97, 98 + _Nucleospira_, 160 + _Nucula_, 175, 177, 178, 183, 184, 185 + _Nuculites_, 177, 178 + Nullipore limestone, 75 + _Nummulites_, 65, 99 + Nummulitic limestone, 73 + _Nyroca_, 283 + + + OCTOPODA, 205 + _Octopus_, 205 + _Odontaspis_, 269, 270, 271 + ODONTOCETI, 295 + _Odontopleura_, 229, 231 + _Odostomia_, 198, 200 + _Oenonites_, 153 + _Olenellus_, 226, 227 + _Oliva_, 204 + _Ommatocarcinus_, 247 + _Omphalotrochus_, 194 + Oolitic ironstone, 81 + _Ophileta_, 192, 193 + OPHIUROIDEA, 141 + _Orbiculoidea_, 160 + _Orbitoides_, 99 + Ordovician bivalve, 177 + " brachiopods, 159 + " cephalopods, 205 + " corals, 113 + " crinoids, 135 + " gasteropods, 193 + " Phyllocarida, 244 + " Radiolaria, 102 + " sponges, 108 + " trilobites, 227 + _Ornithorhynchus_, 286, 287 + _Orthis_, 159, 160, 161, 162 + " limestone, 74 + _Orthoceras_, 204, 205, 206, 207, 208 + _Orthonota_, 177 + _Orthothetes_, 162 + OSTRACODA, features of carapace, 234 + " habits of, 234 + " structure of, 233 + _Ostrea_, 182, 184, 187 + _Otaria_, 299 + _Oxyrhina_, 269, 270, 271 + OXYSTOMATA, 247 + _Oxytelus_, 250 + + + _Pachydomus_, 181 + _Pachyornis_, 282, 283 + _Pachypora_, 73, 116 + _Palaeaster_, 140, 141 + _Palaeeudyptes_, 280, 281 + _Palaeohatteria_, 276 + _Palaeolycus_, 250 + _Palaeoneilo_, 177, 178 + _Palaeoniscus_, 261, 263, 274 + _Palaeopelargus_, 283 + Palaeozoic chitons, 189 + " Cladophora, 122 + " Cryptostomata, 156 + " errant worms, 153 + " strata, 47 + " Trepostomata, 155 + _Palissya_, 89, 164 + _Pallymnarchus_, 279 + _Palorchestes_, 290 + _Panda_, 203 + _Panenka_, 177 + _Paracyainus_, 118 + _Paracyclas_, 177, 179 + _Paradoxechinus_, 145 + _Paradoxorhyncha_, 239 + _Parasqualodon_, 295, 296 + _Pareiasaurus_, 276 + _Patella_, 190, 191 + _Pecten_, 175, 182, 183, 184, 185, 186, 187, 188 + PELECYPODA, characters of, 174 + " hinge structure of, 175 + Pelican, 283 + _Pelicanus_, 283 + _Pelosina_, 97 + _?Peltopleurus_, 262 + _Pentacrinus_, 137, 138 + _Pentagonaster_, 141 + _Pentamerus_, 160, 162 + _Penteune_, 91 + _Peragale_, 289 + _Perameles_, 289, 295 + _Perisphinctes_, 209 + Permian and Triassic reptiles, 276 + _Perna_, 187 + _Peronella_, 148 + _Persoonia_, 90 + _Petaurus_, 295 + _Petraia_, 113 + _Phacops_, 229, 230, 231 + Phalanger, 295 + _Phanerotrema_, 194 + _Phascolomys_, 289, 295 + _Phascolonus_, 289 + _Phialocrinus_, 137 + _Phillipsia_, 232 + _Phoenicopsis_, 88 + _Pholas_, 177 + _Pholidophorus_, 262, 263 + _Phos_, 198 + _Phragmoceras_, 207 + _Phryganea_, 75 + PHYLACTOLAEMATA, 155 + PHYLLOCARIDA, structure of, 243 + _Phyllocladus_, 90 + _Phyllograptus_, 123, 126 + PHYLLOPODA, 233 + _Phyllotheca_, 274 + _Physa_, 191 + _Physonemus_, 261 + Pigeon, 283 + _Pinna_, 186 + PINNIPEDIA, 299 + _Pisania_, 202 + _?Pisocrinus_, 136 + _Placopsilina_, 97 + _Placotrochus_, 118 + _Placunanomia_, 184, 187 + _Plagiarca_, 184 + _Plagiaulax_, 286 + _Planorbis_, 191 + Plants, fossil, 66 + Plant series, characters of, 39 + _Platalea_, 283 + _Platyceps_, 273 + _Platyceras_, 192, 194, 195, 196 + _Platycoila_, 91 + _Platycrinus_, 137 + _Platyschisma_, 196 + _Platysomus_, 263 + _Plaxiphora_, 190 + _Plectroninia_, 112 + _Pleioclinis_, 91 + Pleistocene birds, New Zealand, 283 + " bivalves, 188 + " carinate birds, 283 + " diprotodonts, 289 + " fish, 272 + " Foraminifera, 101 + " gasteropods, 202 + " lobster, 248 + " plants, 91 + " seal, 299 + _Plerophyllum_, 117 + _Plesiastraea_, 119 + _Plesiolampas_, 148 + _Plesiosaurus_, 279 + _Pleuracanthus_, 263 + _Pleurodictyum_, 114 + _Pleuromya_, 183 + _?Pleurostomella_, 98 + _Pleurotoma_, 198, 199, 202 + _Pleurotomaria_, 194, 196, 197, 200, 202 + _Plicatula_, 186 + Pliocene moa, New Zealand, 281 + _Pliosaurus_, 278 + _Plotus_, 283 + _Podocarpus_, 90 + _Poecilodus_, 262 + _?Pollicipes_, 243 + POLYCHAETA, 152, 154 + _Polycotylus_, 279 + _Polymastodon_, 286 + _Polymorphina_, 98, 100 + POLYPLACOPHORA, 189 + _Polypora_, 157 + POLYPROTODONTIA, 287 + _Polystomella_, 101 + POLYZOA, characters of, 59, 155 + " subdivisons of, 155 + Polyzoal limestone, 74 + _Porcellia_, 196 + Porcupine fish, 270, 271 + _Porina_, 158 + _Porphyrio_, 283 + _Portheus_, 268 + _Poteriocrinus_, 137 + Prehensile Rat-kangaroo, 295 + Preservation of fossils, 319 + _Primitia_, 236, 237 + _Pristisomus_, 262 + _Procoptodon_, 290 + _Productus_, 162, 163, 164 + _Proechidna_, 287 + _Proetus_, 229, 232 + _Progura_, 283 + _Prosopon_, 246 + _Protaster_, 142 + _Protocardium_, 185 + _Protopharetra_, 113 + _Protoretepora_, 157 + _Protospongia_, 107,108 + PROTOZOA, characters of, 36, 65, 95 + _Psammechinus_, 145 + _Pseudamaura_, 197 + _Psilichthys_, 264 + PTERIDOPHYTA, characters of, 40 + PTERIDOSPERMEAE, characters of, 40 + _Pterinea_, 178, 179 + _Pteris_ (_Pteridium_), 91 + PTEROPODA, 190, 192, 193, 194 + _Pterygotus_, 248, 249 + _Ptilograptus_, 122 + _Ptychoparia_, 226, 227 + _Pugnellus_, 184 + _Pulvinulina_, 98 + Purbeck marble, 74 + _Purisiphonia_, 110 + _Purpura_, 191 + + + RADIOLARIA, characters of, 36, 66 + " habitat of, 101 + " structure of, 101 + " subdivisions, 102 + Rail, 283 + Raised beaches as distinct from middens, 29 + _Ranella_, 204 + Range-in-time of fossils, 50 + _Raphistoma_, 193, 195 + Rat-kangaroo, 295 + _Receptaculites_, 109 + Regular echinoids, 144 + _Reinschia_, 78 + Reptiles, fossil, 53 + " dentition of, 275 + " structure of, 274 + _Reteocrinus_, 135 + _Retepora_, 158 + _Reticularia_, 164 + _Retiolites_, 124, 128 + _Rhacopteris_, 86 + _Rhinopterocaris_, 244, 246 + _Rhipidomella_, 162 + _Rhizophyllum_, 113 + _Rhodocrinus_, 135 + _Rhombopora_, 156 + _Rhynchonella_, 158, 165, 166 + RHYNCHOTA, 250 + _Rhynchotrema_, 160 + _Ringicula_, 202 + _Risella_, 191 + _Rissoa_, 198 + _Rissoina_, 197 + _Rostellaria_, 198 + _Rotalia_, 96, 101 + Rugose corals, 113 + + + _Saccammina_, 96 + _Saccocaris_, 244 + _Sagenodus_, 263 + _Salterella_, 192 + Sandstones, 71 + _Sanidophyllum_, 115 + _Sarcophilus_, 287, 295 + _Sargus_, 272 + _Scala_, 191, 198, 199, 200, 202 + _Scalaetrochus_, 194 + _Scaldicetus_, 297 + _Scaphella_, 202 + _Scaphites_, 209 + SCAPHOPODA, 188 + _Scenella_, 193 + _Sceparnodon_, 289 + _Schizaster_, 148 + _Schizodus_, 175 + _Schizophoria_, 162 + _Schloenbachia_, 209 + _Scutellina_, 146 + Sea-beds far from the present coast, 29 + Sea-bream, 272 + " -cucumbers, 148 + " -firs, 119, 122 + " -mats, 154, 155 + " -pen, 119 + " -urchins, 59, 143 + " characters of, 144 + Sedentary worms, 154 + _Seguenzia_, 199 + _Selenaria_, 158 + _Semele_, 185 + _Semicassis_, 198 + _Seminula_, 164 + _Semionotus_, 262, 263 + SEPIOIDEA, 205 + _Serpula_, 154 + Serpulite limestone, 74 + _Sertularia_, 119, 122 + Shales, 69 + Sharks, 267, 269, 270, 271 + Shell-limestone, 74 + _Shumardia_, 227 + _Sigsbeia_, 143 + Siliceous rocks, 71 + Silicified wood, 24 + _Siliquaria_, 198 + Silurian bivalves, 177 + " brachiopods, 160 + " brittle-stars, 142 + " cephalopods, 206 + " cirripedes, 241 + " conodonts, 153 + " corals, 113 + " crinoids, 135 + " Foraminifera, 96 + " gasteropods, 193 + " graptolites, Victoria, 128 + " Hexacoralla, 114 + " Octocoralla, 115 + " Ostracoda, 235 + " palaeechinoids, 144 + " Phyllocarida, 246 + " plants, 82 + " Radiolaria, 102 + " sponges, 109 + " starfishes, 140 + " stromatoporoids, 121 + " trilobites, 228 + _Siphonalia_, 198 + _Siphonia_, 110 + _Siphonotreta_, 160 + SIRENIA, 298 + _Sistrum_, 202 + Slate, 70 + Smith, William, 26 + Smittia, 158 + _Solarium_, 198 + _Solenocurtus_, 187 + _Soletellina_, 188 + Sphaerosiderite, 80 + _Sphenopteris_, 85, 89 + _Sphenotrochus_, 118, 119 + _Sphenotus_, 177, 179 + _Sphyrna_, 270 + _Spirifer_, 160, 161, 162, 163, 164 + _Spiriferina_, 165 + " -beds, 208 + _Spirillina_, 96 + _Spirorbis_, 154 + _Spirula_, 205 + _Spirulirostra_, 205, 210 + _Spisula_, 188 + _Spondylostrobus_, 91 + _Spondylus_, 175, 184, 185 + SPONGES, characteristics of, 64, 107 + _Spongilla_, 72 + _Spongodiscus_, 103 + _Spongophyllum_, 116 + Spoonbill, 283 + Spore coal, 76 + _Squalodon_, 295 + _Stacheia_, 97 + Star-corals, 119 + Starfishes, characters of, 61, 139 + _Staurolonche_, 103 + _Stauroneis_, 92 + Steno, 25 + _Stenopora_, 117 + _Stenotheca_, 192 + _Stephanella_, 109 + _Stephanograptus_, 126 + _Stephanotrochus_, 118 + _Sthenurus_, 290 + Sting-ray, 271 + _Stomatopora_, 158 + Storing fossils, 320 + Stork, 283 + Strata, superposition of, 41 + " vertically arranged, 44 + Stratigraphical series, general thickness, 44 + Stratigraphy, 27 + _Strepsodus_, 261 + _Streptelasma_, 113 + _Stricklandinia_, 160 + _Stromatopora_, 120, 121 + _Stromatoporella_, 121, 122 + STROMATOPOROIDS, 63, 120 + _Strombus_, 184, 204 + _Strophalosia_, 163 + _Stropheodonta_, 160, 161 + _Strophonella_, 160 + _Struthiolaria_, 202 + _Studeria_, 148 + _Sturtzura_, 143 + _Stutchburia_, 180 + STYLASTERIDS, 119 + _Subemarginula_, 198 + Submerged forests, 30 + _Sunetta_, 187 + Superposition of strata, 41 + _Synaphe_, 238 + SYNDACTYLA, 288 + _Synedra_, 92 + _Syringopora_, 114 + _Syringothyris_, 164 + + + _Tabellaria_, 92 + _Taeniopteris_, 88, 89, 164, 250, 265 + _Taniwhasaurus_, 279 + _Taphaetus_, 283 + Tasmanian devil, 287, 295 + " wolf, 287, 295 + Tasmanite, 77 + _Taxocrinus_, 135 + _Tellina_, 185, 187, 188 + _Temnechinus_, 146 + _Tentaculites_, 193, 194, 195 + _Terebra_, 198, 199, 202, 204 + _Terebratella_, 166, 168 + _Terebratula_, 166 + _Terebratulina_, 166, 167 + Tertiary ironstone, 81 + _Tessarodoma_, 158 + TETRACORALLA, 113 + Tetractinellid sponge, 110, 112 + _Tetragraptus_, 124, 126 + _Textularia_, 98, 100 + _Thalassina_, 248 + THALLOPHYTA, characters of, 39 + _Thalotia_, 200 + _Thamnastraea_, 118 + _Thinnfeldia_, 88, 89, 182 + _Thurammina_, 97 + _Thyestes_, 258 + _Thylacinus_, 287, 295 + _Thylacoleo_, 293, 303 + Time-range of fossils, 50 + _Tomodus_, 262 + Toothed whales, 295 + Torbanite, 77 + _Torlessia_, 154 + _Trachyderma_, 153, 154 + _Trachypora_, 117 + _Trematonotus_, 194 + _Trematotrochus_, 118, 119 + TREPOSTOMATA, 155 + _Tretocalia_, 112 + Triassic bivalves, 181 + " brachiopods, 164 + " cephalopods, 208 + " crinoids, 137 + " fishes, 262 + " Foraminifera, 98 + " labyrinthodonts, 273 + " Ostracoda, 238 + " Phyllopoda, 233 + " plants, 88 + " reptiles, New Zealand, 276 + _Tribonyx_, 283 + _Tribrachiocrinus_, 137 + _Trichograptus_, 124 + _Tricoelocrinus_, 139 + _Trigonia_, 175, 182, 183, 184, 187 + _Trigonograptus_, 126 + TRILOBITES, habits of, 222 + " structure of, 223 + _Tritylodon_, 276, 286 + _Trivia_, 198, 199 + _Trochoceras_, 205 + _Trochonema_, 195 + _Trochus_, 191, 194, 195 + _Trophon_, 202 + _Truncatulina_, 98, 100 + _Tryplasma_, 113 + Tuatera, 276 + _Tudicla_, 201 + TUNICATA, 257 + _Turbo_, 197, 200 + _Turrilepas_, 241, 243 + _Turritella_, 191, 198, 200, 201, 202 + _Turritella_ -limestone, 74 + _Tylosaurus_, 279 + _Tylospira_, 198, 202 + _Typhis_, 198 + + + _Uncinulus_, 162 + _Unio_, 181, 182 + _Unionella_, 181 + Upper Cambrian trilobites, 227 + " Cretaceous bivalves, 184 + " Cretaceous brachiopod, 166 + " Cretaceous cephalopod, 166 + " Triassic fishes, 262 + " Ordovician graptolites, New South Wales, 127 + " Ordovician graptolites, Victoria, 126 + _Urasterella_, 140 + _Urosthenes_, 262 + + + _Vaginella_, 198, 199 + _Vaginulina_, 98 + _Valvulina_, 97, 98 + _Venus_, 177, 185, 187, 188 + VERMES, characters of, 37 + _Vertebraria_, 264 + VERTEBRATA, characters of, 38, 257 + _Verticordia_, 186 + _Vetotuba_, 194 + _Voluta_, 198, 201, 202 + _Volutilithes_, 198, 201, 202 + _Volvox_, 78 + _Volvulella_, 201 + + + Warrnambool footprints, 301 + Werrikooian bivalves, 187 + " gasteropods, 202 + Whales, 295 + White coal, 77 + _Wilsonia_, 160 + Wombat, 289, 295 + Worms, fossil, 59, 152 + Worm-tracks, 154 + Wrasse family, 271 + _Wynyardia_, 294 + + + Xenophanes, 24 + _Xenorhynchus_, 283 + _Xestoleberis_, 237 + _Xiphosphaera_, 103 + + + _Yvania_, 196 + + + _Zaphrentis_, 117 + _Ziphius_, 296 + + + + +INDEX TO AUSTRALASIAN LOCALITIES. + + +Appended letters indicate the State or Country:-- + +N.S.W., New South Wales; N.T., Northern Territory; N.Z., New Zealand; +Q., Queensland; S.A., South Australia; T., Tasmania; V., Victoria; W.A., +Western Australia. + + Adelaide, S.A., 102 + Aire Coast, V., 138 + Airly, N.S.W., 273 + Alice Springs, S.A., 193 + Altona Bay, V., 112 + Arcola, Q., 279 + Arcoona, S.A., 91 + Ardrossan, S.A., 82, 107 + + + Bacchus Marsh, V., 88, 90 + Balcombe's Bay, V., 190, 239, 317 + Bald Hill, V., 88 + Barker Gorge, W.A., 196, 232, 259 + Barraba, N.S.W., 93, 102 + Batesford, V., 73, 100, 138, 141 + Baton River, N.Z., 195, 207 + Bay of Islands, N.Z., 93 + Beaumaris, V., 119, 243, 248, 270, 271, 296, 297, 317 + Bendigo, V., 108, 109, 124, 246 + Berwick, V., 68 + Bindi, V., 109, 121, 161, 195 + Bingera, N.S.W., 102 + Boggy Creek, V., 112 + Bowen River, Q., 117, 137, 164 + Bowning, N.S.W., 144, 153, 207, 231, 241 + Bowral, N.S.W., 274 + Brighton, N.Z., 146, 248, 280 + Broadhurst's Creek, V., 231 + Broken River, N.Z., 146, 167 + Broken River, Q., 136 + Broome, W.A., 183 + Brunswick, V., 136 + Buchan, V., 79, 109, 115, 136, 161, 195, 203, 207, 231, 237, 258 + Bulla, V., 122 + Bungonia, N.S.W., 300 + Burdekin, Q., 115, 116 + Burnt Creek, V., 259 + Burragorang, N.S.W., 180 + + + Camperdown, V., 74 + Canobolas district, N.S.W., 114 + Canowindra, N.S.W., 162 + Canterbury, N.Z., 154 + Cape Liptrap, V., 71 + Cape Otway, V., 119, 296 + Cape Palliser, N.Z., 203 + Cape Paterson, V., 265, 276 + Carapook, V., 264 + Caroline Creek, T., 227 + Casterton, V., 265 + Castlemaine, V., 126, 246 + Cavan, N.S.W., 109 + Cessnock, N.S.W., 237 + Chatham Ids., 138 + Chillagoe, Q., 115 + Chinchilla, Q., 279 + Clarence Town, N.S.W., 139, 162 + Cliftonwood, N.S.W., 237 + Clunes, V., 279 + Cockatoo Id., N.S.W., 274 + Collie, W.A., 98 + Collingwood, V., 206 + Coole Barghurk Creek, V., 193 + Cooma, N.S.W., 93, 102 + Copeland, N.S.W., 85 + Corio Bay, V., 270 + Corner Creek, Q., 237 + Croydon, Q., 89, 166 + Curiosity Shop, N.Z., 138, 280 + Curlewis, V., 112, 247 + Curramulka, S.A., 108, 177, 192, 235 + Currowang, N.S.W., 127 + + + Dalton, N.S.W., 90, 91 + Dargo High Plains, V., 91 + Darling Downs, Q., 53, 110, 282, 283, 298 + Darling River, N.S.W., 154, 157 + Darriwill, V., 126 + Delegate River, N.S.W., 114 + Derrengullen Creek, N.S.W., 190 + Diggers' Rest, V., 126 + Dolodrook River, V., 193, 227 + Dromana, V., 246 + Dundas Co., V., 264 + + + East Maitland, N.S.W., 154 + Elizabeth River, S.A., 91 + + + Fanning River, Q., 207 + Farley, N.S.W., 180, 237 + Fernbrook, N.S.W., 109 + Fifield, N.S.W., 237 + Flemington, V., 136, 142, 143, 206, 318 + Flinders, V., 65, 112 + Flinders River, Q., 183, 250, 267, 277, 278 + Florentine Valley, T., 159, 227 + Fraser's Creek, V., 231 + + + Gascoyne River, W.A., 117, 136, 137, 232, 262 + Geelong, V., 100, 119, 120, 243 + Geilston, T., 203 + Gellibrand River, V., 199 + Geraldton, W.A., 98, 197, 238 + Gippsland Lakes, V., 168, 243 + Gisborne, V., 299 + Glenelg River, V., 168 + Glenwilliam, N.S.W., 139 + Goodradigbee River, N.S.W., 109 + Goonoo, N.S.W., 85 + Gordon River, T., 115 + Gosford, N.S.W., 53, 262, 263, 273 + Grampians, V., 261 + Grange Burn, Hamilton, V., 143, 270, 271, 296, 297 + Greenough River, W.A., 165, 182, 209 + Grey River, N.Z., 78 + Grice's Creek, V., 317 + Grose Vale, N.S.W., 238 + Gulgong, N.S.W., 279, 286 + Gunning, N.S.W., 91 + + + Haddon, V., 68 + Hallett's Cove, S.A., 119 + Hall's Sound, Papua, 201 + Hamilton, N.Z., 285 + Hamilton, V., 190, 243, 270, 271, 295, 296, 297 + Hamilton River, Q., 267 + Hatton's Corner, N.S.W., 114, 231 + Heathcote, V., 160, 177, 227 + Hobart, T., 68, 203 + Hokonui Hills, N.Z., 164, 165 + Hughenden, Q., 267, 268 + + + Iguana Creek, V., 85 + Irwin River, W.A., 97, 98, 137, 207 + Island of Timor, 163 + + + Jenolan Caves, N.S.W., 102, 121, 300 + + + Kakanui, N.Z., 280 + Kamileroy, Q., 267 + Keilor, V., 128 + Kent's Group, T., 203 + Kilmore, V., 144, 206, 231, 246 + Kilmore Creek, V., 231 + Kimberley, W.A., 136, 137, 192, 207, 262 + King Island, T., 53, 104, 283 + King's Creek, Q., 282 + Kirrak, V., 265 + Knocklofty, T., 264 + Knowsley, V., 227 + Koroit, V., 305 + Kowhai River, N.Z., 189 + + + Lake Callabonna, S.A., 51, 282 + Lake Connewarre, V., 270 + Lake Eyre, S.A., 166, 183, 189, 197 + Lake Frome, S.A., 91 + Lancefield, V., 93, 108, 122, 124, 246 + Laurie's Creek, S.A., 193, 205, 228 + Lawson, N.S.W., 127 + Leichhardt River, Q., 267 + Leigh's Creek, S.A., 193 + Lennard River, W.A., 208 + Lilydale, V., 73, 82, 96, 114, 121, 190, 229, 231, 236, 243, 318 + Limeburners Point, V., 79 + Limestone Creek, Glenelg River, V., 202 + Limestone Creek, Yass, N.S.W.; 136, 231 + Loddon Valley, V., 279 + Lord Howe Id., 279 + Loyola, V., 109, 121, 229, 231 + Lyndhurst, N.S.W., 227 + + + Maddingley, V., 90 + Mallee, V., 71, 101, 119, 138, 141 + Mandurama, N.S.W., 102, 127, 227 + Manly, N.S.W., 88 + Mansfield, V., 53, 122, 154, 231, 259 + Marathon Station, Q., 277 + Maria Id., T., 180 + Maryborough, Q., 146, 184, 304 + Maryvale Creek, Q., 279 + McMahon's Creek, V., 207 + Melbourne, V., 82, 136, 140, 153, 178, 246 + Mersey River, T., 77, 97, 193 + Milburn, N.Z., 296 + Mitchell Downs, Q., 137 + Mitta Mitta River, V., 114 + Molong, N.S.W., 114 + Moonee Ponds Creek, V., 229, 318 + Moorabool River, V., 112, 120, 202 + Mornington, V., 65, 70, 90, 112, 118, 258, 269 + Mosquito Plains, S.A., 300 + Mount Angas, Q., 166 + " Buninyong, V., 303 + " Gambier, S.A., 71, 91, 119, 120, 138, 147, 282, 296 + " Lambie, N.S.W., 85 + " Macedon Cave, 298 + " Potts, N.Z., 276 + " Victoria, N.S.W., 88 + " Wellington, V., 126, 134, 159, 193 + " Wyatt, Q., 109 + Muddy Creek, Hamilton, V., 141, 147, 243, 269, 295 + Mudgee, N.S.W., 109 + Muree, Raymond Terrace, N.S.W., 238 + Murray River Cliffs, S.A., 58, 210 + Murrumbidgee River, N.S.W., 114, 189, 259 + + + Napier Range, W.A., 232 + Narrengullen Creek, N.S.W., 237 + Nelson, N.Z., 78, 126, 164, 165, 182, 233, 248 + Newcastle, N.S.W., 233 + Ngapara, N.Z., 296 + Nimbin, Richmond River, N.S.W., 272 + Norseman district, W.A., 110 + Nugget Point, Otago, N.Z., 274 + Nungatta, N.S.W., 85 + Nyrang Creek, N.S.W., 162 + + + Oakey Creek, N.S.W., 178 + Oamaru, N.Z., 110, 280 + Orakei Bay, N.Z., 158 + Otway Coast, V., 90 + + + Pakaraka, N.Z., 93 + Papua, 100, 146, 148, 184, 187, 188, 201, 203, 209, 210 + Paroo River, Q., 282 + Peak Downs, Q., 282 + Penola, S.A., 300 + Petermann Creek, S.A., 193 + Phillip Co., N.S.W., 282 + Pine Creek, Q., 93 + Pitfield Plains, V., 90 + Pitchery Creek, Q., 278 + Pokolbin, N.S.W., 97, 180 + Port Campbell, V., 247 + Port Darwin, N.T., 103, 248 + Port Stephen, N.S.W., 262 + Preservation Inlet, N.Z., 126 + + + Ravensfield, N.S.W., 180 + Reid Gap, Q., 207 + Richmond Downs, Q., 267 + Richmond River, N.S.W., 93 + Rock Flat Creek, N.S.W., 206 + Rockhampton, Q., 110, 139, 144, 153, 164, 196, 261 + Rough Range, W.A., 116, 122 + + + Sale, V., 112 + San Remo, V., 122 + Sebastopol, V., 93 + Seville, V., 229, 231 + Shakespeare Cliff, N.Z., 146 + Southland, N.Z., 285 + South Yarra, V., 128, 136, 143, 206, 229, 249, 318 + Spring Creek, Torquay, V., 141 + St. Peter's, Sydney, N.S.W., 262 + Stanwell, Q., 137 + Stockyard Creek, N.S.W., 127 + Stroud, N.S.W., 86 + Studley Park, V., 128, 318 + Sunbury, V., 126 + + + Table Cape, T., 74, 190, 269, 270, 294, 296 + Talbot, V., 93 + Talbragar, 267 + Tallong, N.S.W., 127 + Tamworth, N.S.W., 85, 103, 115 + Taranaki, N.Z., 203 + Tempe Downs, S.A., 193, 205, 228 + Thompson River, Q., 277 + Thomson River, V., 229 + Tinderbox Bay, T., 264 + Tingaringi, N.S.W., 127 + Toongabbie, V., 74, 135 + Torquay, V., 74, 141, 148, 243, 269, 296 + Tyer's River, V., 82, 144 + + + Upper Finke Basin, S.A., 159 + Upper Yarra, V., 206, 207, 231, 236 + + + Vegetable Creek, N.S.W., 91 + + + Waihao, N.Z., 296 + Waikari River, N.Z., 141 + Waikouaiti, N.Z., 296 + Wairoa, N.Z., 274 + Wairoa Gorge, N.Z., 137, 162 + Waitaki Valley, N.Z., 296 + Walhalla, V., 114, 121, 128 + Wandong, V., 229, 231 + Wanganui, N.Z., 299 + Wannon River district, V., 53, 90 + Waratah Bay, V., 114, 121, 229 + Warburton, V., 207 + Warrnambool, V., 282, 299, 301, 302 + Waurn Ponds, V., 90, 119, 141, 243, 269, 296 + Wellington Valley, N.S.W., 287, 298, 300 + Well's Creek, N.Z., 165 + West Melbourne Swamp, V., 51 + Westport, N.Z., 78 + Wharekuri, N.Z., 248 + White Cliffs, N.S.W., 138, 179, 183, 184, 195, 279 + Whittlesea, V., 206 + Wilberforce, N.Z., 189 + Wilcannia, N.S.W., 138 + Wirrialpa, S.A., 159 + Wollumbilla, Q., 98, 137, 154, 157, 166, 183, 189 + Wombat Creek, V., 109, 126 + Woori Yallock Creek, V., 231 + Wormbete Creek, V., 74 + Wynyard, T., 246 + + + Yan Yean, V., 318 + Yass, N.S.W., 65, 109, 114, 121, 153, 161, 179, 190, 207, 231, 237, 241 + Yering, V., 142 + Yorke Peninsula, S.A., 226 + Yule Id., Papua, 146, 187, 201 + + + Zeehan, T., 154 + + + * * * * * + + +[Illustration: AUSTRALIA _Shewing chief fossiliferous localities._] + + + * * * * * + +CORRIGENDA. + + Page 65, for head-line "_Protozoa_" read "_How Fossils are Found_." + + Page 147, for head-line "_Characteristic Fossils_" read + "_Sea-urchins_." + + Page 273, for head-line "_Reptiles_" read "_Amphibians_." + + [Transcriber Note: These changes were not utilized here as they only + apply to the titles at the top of the printed pages.] + + * * * * * + +ERRATUM--Page 47. + + _In 1st column_--_for_ "Mesozoic or Secondary (continued)." + + _Read_ "Palaeozoic or Primary" and omit divisional line. + + [Transcriber Note: These changes were applied to the text.] + + + * * * * * + + +Transcriber Note + + +Images were moved so paragraphs were not split. Minor typographical +errors were corrected. Hyphenation was standardized to the most +prevalent form utilized. As the æ ligature was only used five times and +"ae" was used more than 50 times, the ligature was converted to "ae". + + + + + + + + + +End of Project Gutenberg's Australasian Fossils, by Frederick Chapman + +*** END OF THE PROJECT GUTENBERG EBOOK 59074 *** |
